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<document id="38193EE11AF5B2840514F07DAD0F72BB" ID-CLB-Dataset="29084" ID-DOI="10.11646/zootaxa.4455.3.8" ID-GBIF-Dataset="44f06df2-9e8d-40d1-92fa-fe2af75207f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457509" ID-ZooBank="0D9ACB6D-8CD9-4895-B4D3-377A3E482212" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539263086379" checkinUser="plazi" docAuthor="Gill, Anthony C. &amp; Psomadakis, Peter N." docDate="2018" docId="650487BAFFFF092DFF0AC94BFA68FBCD" docLanguage="en" docName="zootaxa.4455.3.8.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudanthias emma Gill &amp; Psomadakis 2018, new species" docType="treatment" docVersion="7" lastPageNumber="529" masterDocId="993DFFC2FFFE0929FF9DCB54FFF5FFDD" masterDocTitle="Pseudanthias emma new species, with notes on a collection of anthiadine serranid fishes from off Myanmar (eastern Indian Ocean)" masterLastPageNumber="536" masterPageNumber="525" pageNumber="526" updateTime="1698576116994" updateUser="ExternalLinkService">
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<mods:title id="EA1F1831679ACA2418E4AFCCDEC75C13">Pseudanthias emma new species, with notes on a collection of anthiadine serranid fishes from off Myanmar (eastern Indian Ocean)</mods:title>
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<mods:namePart id="F4B9D4E45665F52BF74C628E54A4F271">Gill, Anthony C.</mods:namePart>
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<mods:namePart id="3C03D52B7C71A6F4D82D99EC3C69FD11">Psomadakis, Peter N.</mods:namePart>
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<mods:title id="E2B1685D283A423AE8AFBE789D7897A5">Zootaxa</mods:title>
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<mods:date id="8E789694A514DB1CC04789B16BD394C3">2018</mods:date>
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<mods:number id="12F1545DFA2FA895EED85BC27CFEB7BD">2018-08-03</mods:number>
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<treatment id="650487BAFFFF092DFF0AC94BFA68FBCD" ID-DOI="http://doi.org/10.5281/zenodo.5977657" ID-GBIF-Taxon="148691454" ID-Zenodo-Dep="5977657" LSID="urn:lsid:plazi:treatment:650487BAFFFF092DFF0AC94BFA68FBCD" httpUri="http://treatment.plazi.org/id/650487BAFFFF092DFF0AC94BFA68FBCD" lastPageId="4" lastPageNumber="529" pageId="1" pageNumber="526">
<subSubSection id="A5B76527FFFF0928FF0AC94BFE88FD86" pageId="1" pageNumber="526" type="nomenclature">
<paragraph id="ED1236ACFFFF0928FF0AC94BFDD4FDE7" blockId="1.[151,545,543,639]" box="[151,545,543,570]" pageId="1" pageNumber="526">
<heading id="B65A81C0FFFF0928FF0AC94BFDD4FDE7" bold="true" box="[151,545,543,570]" fontSize="11" level="1" pageId="1" pageNumber="526" reason="1">
<emphasis id="DFD9EABEFFFF0928FF0AC94BFDD4FDE7" bold="true" box="[151,545,543,570]" pageId="1" pageNumber="526">
<taxonomicName id="2AAD4D2FFFFF0928FF0AC94BFE7FFDE4" authority="Gill &amp; Psomadakis, 2018" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[151,394,543,569]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="526" phylum="Chordata" rank="species" species="emma" status="sp. nov.">
<emphasis id="DFD9EABEFFFF0928FF0AC94BFE7FFDE4" bold="true" box="[151,394,543,569]" italics="true" pageId="1" pageNumber="526">Pseudanthias emma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4EA57C5FFFF0928FE0FC974FDD4FDE7" box="[402,545,544,570]" pageId="1" pageNumber="526" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="ED1236ACFFFF0928FF0AC916FE88FD86" blockId="1.[151,545,543,639]" box="[151,381,578,603]" pageId="1" pageNumber="526">
<figureCitation id="75962A29FFFF0928FF0AC916FF03FD86" box="[151,246,578,603]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[151,1436,1123,1873]" captionTargetId="figure@4.[151,1436,1123,1873]" captionTargetPageId="4" captionText="FIGURE 1. Pseudanthias emma sp. nov., SAIAB 203722, 107 mm SL, male holotype, off Tanintharyi coast, Myanmar, Andaman Sea, Indian Ocean. Note that filamentous tips of caudal fin were cropped during photography. Photo by Oddgeir Alvheim." httpUri="https://zenodo.org/record/1457511/files/figure.png" pageId="1" pageNumber="526">Figure 1</figureCitation>
;
<tableCitation id="A02F0317FFFF0928FE99C916FE88FD86" box="[260,381,578,603]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="2.[151,239,500,523]" captionStartId-1="3.[151,239,152,175]" captionText-0="TABLE 1. Morphometric values for Pseudanthias emma sp. nov. expressed as percentage SL, with comparable data for P. pillai (after Heemstra &amp; Akhilesh 2012). * indicates where data for P. pillai is lacking or not comparable." captionText-1="TABLE 2. Comparison of coloration features of male Pseudanthias emma sp. nov. and P. pillai" pageId="1" pageNumber="526">Tables 1–2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="A5B76527FFFF0928FF0AC933FEB3FDA2" box="[151,326,614,639]" pageId="1" pageNumber="526" type="description">
<paragraph id="ED1236ACFFFF0928FF0AC933FEB3FDA2" blockId="1.[151,545,543,639]" box="[151,326,614,639]" pageId="1" pageNumber="526">Emma’s basslet</paragraph>
</subSubSection>
<subSubSection id="A5B76527FFFF0928FF0AC9FAFED1FCD2" pageId="1" pageNumber="526" type="materials_examined">
<paragraph id="ED1236ACFFFF0928FF0AC9FAFED1FCD2" blockId="1.[151,1437,686,2005]" pageId="1" pageNumber="526">
<materialsCitation id="5DC53CF1FFFF0928FF0AC9FAFED1FCD2" ID-GBIF-Occurrence="1917075941" collectingDate="2015-05-25" collectionCode="SAIAB" collectorName="V Dr. Fridtjof Nansen &amp; P. N. Psomadakis." country="Myanmar" elevation="105" latitude="11.666" location="Andaman Sea" longLatPrecision="13" longitude="97.26933" municipality="Indian Ocean" pageId="1" pageNumber="526" specimenCode="SAIAB 203722, 107" specimenCount="1" stateProvince="Tanintharyi" typeStatus="holotype">
<emphasis id="DFD9EABEFFFF0928FF0AC9FAFEFEFD1A" bold="true" box="[151,267,686,711]" pageId="1" pageNumber="526">
<typeStatus id="3216880EFFFF0928FF0AC9FAFEF3FD1A" box="[151,262,686,711]" pageId="1" pageNumber="526" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="BD0B9ED7FFFF0928FE8EC9FBFE30FD1A" box="[275,453,687,711]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="1" pageNumber="526">
SAIAB
<quantity id="2A559B49FFFF0928FEF3C9FBFE30FD1A" box="[366,453,687,711]" metricMagnitude="5" metricUnit="m" metricValue="2.03722107" pageId="1" pageNumber="526" unit="mm" value="2.03722107E8">203722</quantity>
</specimenCode>
,
<specimenCode id="BD0B9ED7FFFF0928FE4CC9FBFE0EFD1A" box="[465,507,687,711]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="1" pageNumber="526">107</specimenCode>
mm SL, off
<collectingRegion id="2F69F84EFFFF0928FD13C9FAFCFAFD1A" box="[654,783,686,711]" country="Myanmar" name="Taninthayi" pageId="1" pageNumber="526">Tanintharyi</collectingRegion>
coast,
<collectingCountry id="95BA763CFFFF0928FCC2C9FBFC3BFD1A" box="[863,974,687,711]" name="Myanmar" pageId="1" pageNumber="526">Myanmar</collectingCountry>
,
<location id="E8726077FFFF0928FC44C9FAFB8CFD1A" LSID="urn:lsid:plazi:treatment:650487BAFFFF092DFF0AC94BFA68FBCD:E8726077FFFF0928FC44C9FAFB8CFD1A" box="[985,1145,686,711]" country="Myanmar" latitude="11.666" longLatPrecision="13" longitude="97.26933" municipality="Indian Ocean" name="Andaman Sea" pageId="1" pageNumber="526" stateProvince="Tanintharyi">Andaman Sea</location>
,
<collectingMunicipality id="0D76ACD6FFFF0928FB1BC9FAFAE9FD1A" box="[1158,1308,686,711]" pageId="1" pageNumber="526">Indian Ocean</collectingMunicipality>
(
<geoCoordinate id="8899506BFFFF0928FAB0C9FAFF5BFD37" direction="north" orientation="latitude" pageId="1" pageNumber="526" precision="9" value="11.666">11°39.96’ N</geoCoordinate>
,
<geoCoordinate id="8899506BFFFF0928FF22C986FEB6FD36" box="[191,323,722,747]" direction="east" orientation="longitude" pageId="1" pageNumber="526" precision="9" value="97.26933">97°16.16’E</geoCoordinate>
),
<elevation id="6680D19FFFFF0928FEC0C986FE58FD37" box="[349,429,722,747]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="526" unit="m" value="105.0">
<quantity id="2A559B49FFFF0928FEC0C986FE58FD37" box="[349,429,722,747]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="526" unit="m" value="105.0">105 m</quantity>
</elevation>
, R/
<collectorName id="4058537AFFFF0928FE47C987FD16FD37" box="[474,739,723,747]" pageId="1" pageNumber="526">
V
<emphasis id="DFD9EABEFFFF0928FE66C987FD16FD37" box="[507,739,723,747]" italics="true" pageId="1" pageNumber="526">Dr. Fridtjof Nansen</emphasis>
</collectorName>
, stn 149, bottom trawl,
<collectingDate id="8957E984FFFF0928FB93C986FB46FD36" box="[1038,1203,722,747]" pageId="1" pageNumber="526" value="2015-05-25">
<date id="9913106CFFFF0928FB93C986FB46FD36" box="[1038,1203,722,747]" pageId="1" pageNumber="526" value="2015-05-25">25 May 2015</date>
</collectingDate>
, collected by
<collectorName id="4058537AFFFF0928FAFDC987FED1FCD2" pageId="1" pageNumber="526">P. N. Psomadakis.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A5B76527FFFF0928FF5AC84EFE4DFC1E" pageId="1" pageNumber="526" type="diagnosis">
<paragraph id="ED1236ACFFFF0928FF5AC84EFE4DFC1E" blockId="1.[151,1437,686,2005]" pageId="1" pageNumber="526">
<emphasis id="DFD9EABEFFFF0928FF5AC84EFEB4FCEE" bold="true" box="[199,321,794,819]" pageId="1" pageNumber="526">Diagnosis.</emphasis>
The following combination of characters distinguishes
<taxonomicName id="2AAD4D2FFFFF0928FC2FC84FFB67FCEF" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[946,1170,795,819]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="526" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFF0928FC2FC84FFB67FCEF" box="[946,1170,795,819]" italics="true" pageId="1" pageNumber="526">Pseudanthias emma</emphasis>
</taxonomicName>
from congeners: dorsal rays X,16; pectoral rays 18; lateral-line scales 42; third dorsal-fin spine longest, prolonged; no papillae on posterior margin of orbit; soft part of dorsal with low scaly sheath; subopercle and interopercle indistinctly serrated; caudal fin weakly concave centrally, with elongate filaments extending from second to uppermost and second to lowermost branched rays.
</paragraph>
</subSubSection>
<subSubSection id="A5B76527FFFF092DFF5AC89AFF1AFC30" lastPageId="4" lastPageNumber="529" pageId="1" pageNumber="526" type="description">
<paragraph id="ED1236ACFFFF0928FF5AC89AFB1FFADA" blockId="1.[151,1437,686,2005]" pageId="1" pageNumber="526">
<emphasis id="DFD9EABEFFFF0928FF5AC89AFEACFC3A" bold="true" box="[199,345,974,999]" pageId="1" pageNumber="526">Description.</emphasis>
Dorsal rays X,16, all segmented rays branched; third dorsal-fin spine longest, with fleshy pennant extending from tip; anal rays III,7, all segmented rays branched; pectoral rays 18/18, all rays branched except upper 2/2, no serrations on rays; pelvic rays 1,5; caudal fin weakly concave centrally, with elongate filaments extending from second to uppermost and second to lowermost branched rays; principal caudal rays 9 + 8; branched caudal-fin rays 7 + 6; upper procurrent caudal-fin rays 7; lower procurrent caudal rays 7; lateral-line scales 42/42; scales above lateral-line to origin of dorsal fin (counted upwards and forwards from lateral line) 6/6; scales above lateral-line to base of fifth dorsal spine (counted upwards and forwards from lateral line) 4/3; scales above origin of anal fin origin to lateral line (counted upwards and forwards from anal origin) 18/18; circumpeduncular scales 26; gill rakers 11 + 22; branchiostegal rays 7; pseudobranch filaments 20.
</paragraph>
<paragraph id="ED1236ACFFFF0928FF5ACE46FCE2FA03" blockId="1.[151,1437,686,2005]" pageId="1" pageNumber="526">Vertebrae 10 + 16; predorsal formula 0/0+0/2/1+1; main shaft (proximal component) of first dorsal pterygiophore roughly perpendicular to long axis of body; no trisegmental pterygiophores associated with dorsal and anal fins; ribs present on vertebrae 3 through 10; epineurals present on vertebrae 1 through 13; paired parapophyses present on first caudal vertebra (see Baldwin 1990: fig. 21B); parhypural and hypurals autogenous; well-developed hypurapophysis on parhypural; epurals 3; single uroneural (posterior uroneural absent); ventral tip of cleithrum with well-developed posteroventral process.</paragraph>
<paragraph id="ED1236ACFFFF0928FF5ACEBEFD18F9DE" blockId="1.[151,1437,686,2005]" box="[199,749,1514,1539]" pageId="1" pageNumber="526">
Morphometric values are summarised in
<tableCitation id="A02F0317FFFF0928FD0FCEBEFD13F9DE" box="[658,742,1514,1539]" captionStart="TABLE 1" captionStartId="2.[151,239,500,523]" captionText="TABLE 1. Morphometric values for Pseudanthias emma sp. nov. expressed as percentage SL, with comparable data for P. pillai (after Heemstra &amp; Akhilesh 2012). * indicates where data for P. pillai is lacking or not comparable." pageId="1" pageNumber="526">Table 1</tableCitation>
.
</paragraph>
<paragraph id="ED1236ACFFFF0928FF5ACD5AFAD7F8FE" blockId="1.[151,1437,686,2005]" pageId="1" pageNumber="526">Mouth large, slightly oblique, posterior margin of maxilla reaching to vertical through posterior edge of pupil; mouth terminal, lower jaw projecting slightly; premaxilla with an enlarged recurved canine anterolaterally, a band of small conical teeth about five rows wide at symphysis reducing to two rows on sides of jaw, with the outer row teeth larger and slightly curved, and inner pair anteriorly nearest symphysis enlarged and caniniform, directed inwards; dentary with a band of small conical teeth about six rows wide, reducing to single row posteriorly on sides of jaw, an enlarged, curved canine tooth on sides of jaw, and an anteriorly pointing canine tooth anterolaterally; vomer with a triangular patch of small conical teeth, five rows wide in midline; palatine with a narrow band of small conical teeth, three rows wide at widest point; ectopterygoid, mesopterygoid and tongue edentate.</paragraph>
<paragraph id="ED1236ACFFFF092BFF5ACC7AFE21FF09" blockId="1.[151,1437,686,2005]" lastBlockId="2.[151,1437,151,460]" lastPageId="2" lastPageNumber="527" pageId="1" pageNumber="526">Opercle with 3 flat spines, middle spine longest, upper spine mostly concealed by scales; posterior margin of preopercle with 35/37 fine serrations, these gradually increasing in size until angle of preopercle; no serrations on ventral part of preopercle; interopercle with 1/2 indistinct serrations; subopercle with 2/2 indistinct serrations; posttemporal with about 5 indistinct serrations; no apparent serrations on supracleithrum. Anterior nostril positioned at middle of snout, tubular with elongate flap on posterior rim, flap reaching posterior nostril when laid back; posterior nostril at mid-upper, anterior border of orbit. No papillae on posterior rim of orbit. Snout not hypertrophied at symphysis.</paragraph>
<paragraph id="ED1236ACFFFC092BFF5ACB8AFE6FFE11" blockId="2.[151,1437,151,460]" pageId="2" pageNumber="527">
Scales ctenoid with peripheral cteni (
<bibRefCitation id="893C4B5DFFFC092BFDE8CB8AFCE0FF2A" box="[629,789,222,247]" pageId="2" pageNumber="527" refId="ref5478" refString="Roberts, C. D. (1993) Comparative morphology of spined scales and their phylogenetic significance in the Teleostei. Bulletin of Marine Science, 52, 60 - 113." type="journal article">Roberts 1993</bibRefCitation>
); lateral line broadly arched over pectoral fin following body contour to caudal-fin base; head fully scaled except for lips, and areas in front of and immediately below nostrils; no auxiliary scales on body, a few auxiliary scales on cheeks and operculum; low scaly sheath on soft dorsal and anal fins, with indistinct small scales present between segmented rays; caudal fin with scaly basal sheath, with small scales extending over almost all of fin, except for fin tips and posterior part of membranes of middle rays; pectoral fins with basal, wedge-shaped sheath of small scales; pelvic fins with small scales extending on to basal half of fins.
</paragraph>
<caption id="B9D26624FFFC092BFF0ACAA0FAE0FDF6" pageId="2" pageNumber="527">
<paragraph id="ED1236ACFFFC092BFF0ACAA0FAE0FDF6" blockId="2.[151,1436,499,555]" pageId="2" pageNumber="527">
<emphasis id="DFD9EABEFFFC092BFF0ACAA0FEF6FDD6" bold="true" box="[151,259,500,523]" pageId="2" pageNumber="527">TABLE 1</emphasis>
. Morphometric values for
<taxonomicName id="2AAD4D2FFFFC092BFDBECAA7FD0CFDD6" authority="Gill &amp; Psomadakis, 2018" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[547,761,499,523]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="527" phylum="Chordata" rank="species" species="emma" status="sp. nov.">
<emphasis id="DFD9EABEFFFC092BFDBECAA7FD0CFDD6" box="[547,761,499,523]" italics="true" pageId="2" pageNumber="527">Pseudanthias emma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4EA57C5FFFC092BFC9DCAA0FCA3FDD6" box="[768,854,500,523]" pageId="2" pageNumber="527" rank="species">
<emphasis id="DFD9EABEFFFC092BFC9DCAA0FCA3FDD6" bold="true" box="[768,854,500,523]" pageId="2" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
expressed as percentage SL, with comparable data for
<taxonomicName id="2AAD4D2FFFFC092BFF0AC940FF1CFDF6" box="[151,233,531,555]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="527" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFC092BFF0AC940FF1CFDF6" box="[151,233,531,555]" italics="true" pageId="2" pageNumber="527">P. pillai</emphasis>
</taxonomicName>
(after Heemstra &amp; Akhilesh 2012). * indicates where data for
<taxonomicName id="2AAD4D2FFFFC092BFC18C940FC22FDF6" box="[901,983,531,555]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="527" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFC092BFC18C940FC22FDF6" box="[901,983,531,555]" italics="true" pageId="2" pageNumber="527">P. pillai</emphasis>
</taxonomicName>
is lacking or not comparable.
</paragraph>
</caption>
<paragraph id="ED1236ACFFFC092BFD22C914FC1EF83B" pageId="2" pageNumber="527">
<table id="9FADC40CFFFCF6D6FF02C96BFAE7F83B" box="[159,1298,575,2022]" gridcols="3" gridrows="34" pageId="2" pageNumber="527">
<tr id="539D34EEFFFCF6D6FF02C96BFAE7FD4F" box="[159,1298,575,658]" gridrow="0" pageId="2" pageNumber="527" rowspan-0="1">
<th id="104C5D92FFFCF6D6FD22C96BFCA8FD4F" box="[703,861,575,658]" gridcol="1" gridrow="0" pageId="2" pageNumber="527">
<taxonomicName id="2AAD4D2FFFFC092BFD22C914FCE1FD88" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[703,788,576,597]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="527" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFC092BFD22C914FCE1FD88" box="[703,788,576,597]" italics="true" pageId="2" pageNumber="527">P. emma</emphasis>
</taxonomicName>
SAIAB 203722 Holotype
</th>
<th id="104C5D92FFFCF6D6FC36C96BFAE7FD4F" box="[939,1298,575,658]" gridcol="2" gridrow="0" pageId="2" pageNumber="527">
<taxonomicName id="2AAD4D2FFFFC092BFC36C914FC0CFD88" box="[939,1017,575,597]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="527" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFC092BFC36C914FC0CFD88" box="[939,1017,575,597]" italics="true" pageId="2" pageNumber="527">P. pillai</emphasis>
</taxonomicName>
Male holotype and female paratype
</th>
</tr>
<tr id="539D34EEFFFCF6D6FF02C9F1FAE7FD66" box="[159,1298,677,699]" gridrow="1" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C9F1FDC7FD66" box="[159,562,677,699]" gridcol="0" gridrow="1" pageId="2" pageNumber="527">SL (mm)</th>
<td id="104C5D92FFFCF6D6FD22C9F1FCA8FD66" box="[703,861,677,699]" gridcol="1" gridrow="1" pageId="2" pageNumber="527">107</td>
<td id="104C5D92FFFCF6D6FC36C9F1FAE7FD66" box="[939,1298,677,699]" gridcol="2" gridrow="1" pageId="2" pageNumber="527">100–119</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C99BFAE7FD38" box="[159,1298,719,741]" gridrow="2" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C99BFDC7FD38" box="[159,562,719,741]" gridcol="0" gridrow="2" pageId="2" pageNumber="527">Greatest body depth</th>
<td id="104C5D92FFFCF6D6FD22C99BFCA8FD38" box="[703,861,719,741]" gridcol="1" gridrow="2" pageId="2" pageNumber="527">37.9</td>
<td id="104C5D92FFFCF6D6FC36C99BFAE7FD38" box="[939,1298,719,741]" gridcol="2" gridrow="2" pageId="2" pageNumber="527">35–39</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C9ACFAE7FCD3" box="[159,1298,760,782]" gridrow="3" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C9ACFDC7FCD3" box="[159,562,760,782]" gridcol="0" gridrow="3" pageId="2" pageNumber="527">Body depth at anal-fin origin</th>
<td id="104C5D92FFFCF6D6FD22C9ACFCA8FCD3" box="[703,861,760,782]" gridcol="1" gridrow="3" pageId="2" pageNumber="527">34.0</td>
<td id="104C5D92FFFCF6D6FC36C9ACFAE7FCD3" box="[939,1298,760,782]" gridcol="2" gridrow="3" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C876FAE7FCE5" box="[159,1298,802,824]" gridrow="4" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C876FDC7FCE5" box="[159,562,802,824]" gridcol="0" gridrow="4" pageId="2" pageNumber="527">Body width</th>
<td id="104C5D92FFFCF6D6FD22C876FCA8FCE5" box="[703,861,802,824]" gridcol="1" gridrow="4" pageId="2" pageNumber="527">17.9</td>
<td id="104C5D92FFFCF6D6FC36C876FAE7FCE5" box="[939,1298,802,824]" gridcol="2" gridrow="4" pageId="2" pageNumber="527">16–16.8</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C81FFAE7FCBC" box="[159,1298,843,865]" gridrow="5" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C81FFDC7FCBC" box="[159,562,843,865]" gridcol="0" gridrow="5" pageId="2" pageNumber="527">Head length</th>
<td id="104C5D92FFFCF6D6FD22C81FFCA8FCBC" box="[703,861,843,865]" gridcol="1" gridrow="5" pageId="2" pageNumber="527">34.3</td>
<td id="104C5D92FFFCF6D6FC36C81FFAE7FCBC" box="[939,1298,843,865]" gridcol="2" gridrow="5" pageId="2" pageNumber="527">37–38</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C820FAE7FC57" box="[159,1298,884,906]" gridrow="6" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C820FDC7FC57" box="[159,562,884,906]" gridcol="0" gridrow="6" pageId="2" pageNumber="527">Snout length</th>
<td id="104C5D92FFFCF6D6FD22C820FCA8FC57" box="[703,861,884,906]" gridcol="1" gridrow="6" pageId="2" pageNumber="527">7.6</td>
<td id="104C5D92FFFCF6D6FC36C820FAE7FC57" box="[939,1298,884,906]" gridcol="2" gridrow="6" pageId="2" pageNumber="527">7.3–9.1</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C8C9FAE7FC6E" box="[159,1298,925,947]" gridrow="7" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C8C9FDC7FC6E" box="[159,562,925,947]" gridcol="0" gridrow="7" pageId="2" pageNumber="527">Orbit diameter</th>
<td id="104C5D92FFFCF6D6FD22C8C9FCA8FC6E" box="[703,861,925,947]" gridcol="1" gridrow="7" pageId="2" pageNumber="527">9.6</td>
<td id="104C5D92FFFCF6D6FC36C8C9FAE7FC6E" box="[939,1298,925,947]" gridcol="2" gridrow="7" pageId="2" pageNumber="527">8.7–9.1</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C893FAE7FC00" box="[159,1298,967,989]" gridrow="8" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C893FDC7FC00" box="[159,562,967,989]" gridcol="0" gridrow="8" pageId="2" pageNumber="527">Bony interorbital width</th>
<td id="104C5D92FFFCF6D6FD22C893FCA8FC00" box="[703,861,967,989]" gridcol="1" gridrow="8" pageId="2" pageNumber="527">9.3</td>
<td id="104C5D92FFFCF6D6FC36C893FAE7FC00" box="[939,1298,967,989]" gridcol="2" gridrow="8" pageId="2" pageNumber="527">8.1–8.2</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02C8A4FAE7FBDB" box="[159,1298,1008,1030]" gridrow="9" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02C8A4FDC7FBDB" box="[159,562,1008,1030]" gridcol="0" gridrow="9" pageId="2" pageNumber="527">Upper jaw length</th>
<td id="104C5D92FFFCF6D6FD22C8A4FCA8FBDB" box="[703,861,1008,1030]" gridcol="1" gridrow="9" pageId="2" pageNumber="527">14.5</td>
<td id="104C5D92FFFCF6D6FC36C8A4FAE7FBDB" box="[939,1298,1008,1030]" gridcol="2" gridrow="9" pageId="2" pageNumber="527">15</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CF4EFAE7FBED" box="[159,1298,1050,1072]" gridrow="10" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CF4EFDC7FBED" box="[159,562,1050,1072]" gridcol="0" gridrow="10" pageId="2" pageNumber="527">Maxilla width</th>
<td id="104C5D92FFFCF6D6FD22CF4EFCA8FBED" box="[703,861,1050,1072]" gridcol="1" gridrow="10" pageId="2" pageNumber="527">6.3</td>
<td id="104C5D92FFFCF6D6FC36CF4EFAE7FBED" box="[939,1298,1050,1072]" gridcol="2" gridrow="10" pageId="2" pageNumber="527">5.9–6.3</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CF17FAE7FB84" box="[159,1298,1091,1113]" gridrow="11" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CF17FDC7FB84" box="[159,562,1091,1113]" gridcol="0" gridrow="11" pageId="2" pageNumber="527">Caudal peduncle length</th>
<td id="104C5D92FFFCF6D6FD22CF17FCA8FB84" box="[703,861,1091,1113]" gridcol="1" gridrow="11" pageId="2" pageNumber="527">20.5</td>
<td id="104C5D92FFFCF6D6FC36CF17FAE7FB84" box="[939,1298,1091,1113]" gridcol="2" gridrow="11" pageId="2" pageNumber="527">19–22</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CF38FAE7FB5F" box="[159,1298,1132,1154]" gridrow="12" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CF38FDC7FB5F" box="[159,562,1132,1154]" gridcol="0" gridrow="12" pageId="2" pageNumber="527">Caudal peduncle depth</th>
<td id="104C5D92FFFCF6D6FD22CF38FCA8FB5F" box="[703,861,1132,1154]" gridcol="1" gridrow="12" pageId="2" pageNumber="527">16.4</td>
<td id="104C5D92FFFCF6D6FC36CF38FAE7FB5F" box="[939,1298,1132,1154]" gridcol="2" gridrow="12" pageId="2" pageNumber="527">12–13</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CFC1FAE7FB76" box="[159,1298,1173,1195]" gridrow="13" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CFC1FDC7FB76" box="[159,562,1173,1195]" gridcol="0" gridrow="13" pageId="2" pageNumber="527">Predorsal length</th>
<td id="104C5D92FFFCF6D6FD22CFC1FCA8FB76" box="[703,861,1173,1195]" gridcol="1" gridrow="13" pageId="2" pageNumber="527">33.1</td>
<td id="104C5D92FFFCF6D6FC36CFC1FAE7FB76" box="[939,1298,1173,1195]" gridcol="2" gridrow="13" pageId="2" pageNumber="527">33</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CFEBFAE7FB08" box="[159,1298,1215,1237]" gridrow="14" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CFEBFDC7FB08" box="[159,562,1215,1237]" gridcol="0" gridrow="14" pageId="2" pageNumber="527">Preanal length</th>
<td id="104C5D92FFFCF6D6FD22CFEBFCA8FB08" box="[703,861,1215,1237]" gridcol="1" gridrow="14" pageId="2" pageNumber="527">65.6</td>
<td id="104C5D92FFFCF6D6FC36CFEBFAE7FB08" box="[939,1298,1215,1237]" gridcol="2" gridrow="14" pageId="2" pageNumber="527">64–65</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CFBCFAE7FB23" box="[159,1298,1256,1278]" gridrow="15" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CFBCFDC7FB23" box="[159,562,1256,1278]" gridcol="0" gridrow="15" pageId="2" pageNumber="527">Prepelvic length</th>
<td id="104C5D92FFFCF6D6FD22CFBCFCA8FB23" box="[703,861,1256,1278]" gridcol="1" gridrow="15" pageId="2" pageNumber="527">37.0</td>
<td id="104C5D92FFFCF6D6FC36CFBCFAE7FB23" box="[939,1298,1256,1278]" gridcol="2" gridrow="15" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CE46FAE7FAF5" box="[159,1298,1298,1320]" gridrow="16" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CE46FDC7FAF5" box="[159,562,1298,1320]" gridcol="0" gridrow="16" pageId="2" pageNumber="527">Dorsal fin base length</th>
<td id="104C5D92FFFCF6D6FD22CE46FCA8FAF5" box="[703,861,1298,1320]" gridcol="1" gridrow="16" pageId="2" pageNumber="527">61.0</td>
<td id="104C5D92FFFCF6D6FC36CE46FAE7FAF5" box="[939,1298,1298,1320]" gridcol="2" gridrow="16" pageId="2" pageNumber="527">62–63</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CE6FFAE7FA8C" box="[159,1298,1339,1361]" gridrow="17" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CE6FFDC7FA8C" box="[159,562,1339,1361]" gridcol="0" gridrow="17" pageId="2" pageNumber="527">First dorsal spine</th>
<td id="104C5D92FFFCF6D6FD22CE6FFCA8FA8C" box="[703,861,1339,1361]" gridcol="1" gridrow="17" pageId="2" pageNumber="527">broken</td>
<td id="104C5D92FFFCF6D6FC36CE6FFAE7FA8C" box="[939,1298,1339,1361]" gridcol="2" gridrow="17" pageId="2" pageNumber="527">5.6–6.6</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CE30FAE7FAA7" box="[159,1298,1380,1402]" gridrow="18" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CE30FDC7FAA7" box="[159,562,1380,1402]" gridcol="0" gridrow="18" pageId="2" pageNumber="527">Third dorsal spine</th>
<td id="104C5D92FFFCF6D6FD22CE30FCA8FAA7" box="[703,861,1380,1402]" gridcol="1" gridrow="18" pageId="2" pageNumber="527">23.9</td>
<td id="104C5D92FFFCF6D6FC36CE30FAE7FAA7" box="[939,1298,1380,1402]" gridcol="2" gridrow="18" pageId="2" pageNumber="527">16–18</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CED9FAE7FA7E" box="[159,1298,1421,1443]" gridrow="19" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CED9FDC7FA7E" box="[159,562,1421,1443]" gridcol="0" gridrow="19" pageId="2" pageNumber="527">Fourth dorsal spine</th>
<td id="104C5D92FFFCF6D6FD22CED9FCA8FA7E" box="[703,861,1421,1443]" gridcol="1" gridrow="19" pageId="2" pageNumber="527">14.8</td>
<td id="104C5D92FFFCF6D6FC36CED9FAE7FA7E" box="[939,1298,1421,1443]" gridcol="2" gridrow="19" pageId="2" pageNumber="527">14–15</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CEE3FAE7FA10" box="[159,1298,1463,1485]" gridrow="20" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CEE3FDC7FA10" box="[159,562,1463,1485]" gridcol="0" gridrow="20" pageId="2" pageNumber="527">Tenth dorsal spine</th>
<td id="104C5D92FFFCF6D6FD22CEE3FCA8FA10" box="[703,861,1463,1485]" gridcol="1" gridrow="20" pageId="2" pageNumber="527">11.8</td>
<td id="104C5D92FFFCF6D6FC36CEE3FAE7FA10" box="[939,1298,1463,1485]" gridcol="2" gridrow="20" pageId="2" pageNumber="527">13–14</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CEB4FAE7FA2B" box="[159,1298,1504,1526]" gridrow="21" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CEB4FDC7FA2B" box="[159,562,1504,1526]" gridcol="0" gridrow="21" pageId="2" pageNumber="527">First segmented dorsal ray</th>
<td id="104C5D92FFFCF6D6FD22CEB4FCA8FA2B" box="[703,861,1504,1526]" gridcol="1" gridrow="21" pageId="2" pageNumber="527">13.9</td>
<td id="104C5D92FFFCF6D6FC36CEB4FAE7FA2B" box="[939,1298,1504,1526]" gridcol="2" gridrow="21" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CD5EFAE7F9FD" box="[159,1298,1546,1568]" gridrow="22" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CD5EFDC7F9FD" box="[159,562,1546,1568]" gridcol="0" gridrow="22" pageId="2" pageNumber="527">Longest segmented dorsal ray (number)</th>
<td id="104C5D92FFFCF6D6FD22CD5EFCA8F9FD" box="[703,861,1546,1568]" gridcol="1" gridrow="22" pageId="2" pageNumber="527">18.2 (13)</td>
<td id="104C5D92FFFCF6D6FC36CD5EFAE7F9FD" box="[939,1298,1546,1568]" gridcol="2" gridrow="22" pageId="2" pageNumber="527">24–25 (*)</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CD67FAE7F994" box="[159,1298,1587,1609]" gridrow="23" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CD67FDC7F994" box="[159,562,1587,1609]" gridcol="0" gridrow="23" pageId="2" pageNumber="527">Anal fin base length</th>
<td id="104C5D92FFFCF6D6FD22CD67FCA8F994" box="[703,861,1587,1609]" gridcol="1" gridrow="23" pageId="2" pageNumber="527">17.0</td>
<td id="104C5D92FFFCF6D6FC36CD67FAE7F994" box="[939,1298,1587,1609]" gridcol="2" gridrow="23" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CD08FAE7F9AF" box="[159,1298,1628,1650]" gridrow="24" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CD08FDC7F9AF" box="[159,562,1628,1650]" gridcol="0" gridrow="24" pageId="2" pageNumber="527">First anal spine</th>
<td id="104C5D92FFFCF6D6FD22CD08FCA8F9AF" box="[703,861,1628,1650]" gridcol="1" gridrow="24" pageId="2" pageNumber="527">7.2</td>
<td id="104C5D92FFFCF6D6FC36CD08FAE7F9AF" box="[939,1298,1628,1650]" gridcol="2" gridrow="24" pageId="2" pageNumber="527">8.2–10</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CDD1FAE7F946" box="[159,1298,1669,1691]" gridrow="25" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CDD1FDC7F946" box="[159,562,1669,1691]" gridcol="0" gridrow="25" pageId="2" pageNumber="527">Second anal spine</th>
<td id="104C5D92FFFCF6D6FD22CDD1FCA8F946" box="[703,861,1669,1691]" gridcol="1" gridrow="25" pageId="2" pageNumber="527">16.9</td>
<td id="104C5D92FFFCF6D6FC36CDD1FAE7F946" box="[939,1298,1669,1691]" gridcol="2" gridrow="25" pageId="2" pageNumber="527">15–17</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CDFBFAE7F918" box="[159,1298,1711,1733]" gridrow="26" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CDFBFDC7F918" box="[159,562,1711,1733]" gridcol="0" gridrow="26" pageId="2" pageNumber="527">Third anal spine</th>
<td id="104C5D92FFFCF6D6FD22CDFBFCA8F918" box="[703,861,1711,1733]" gridcol="1" gridrow="26" pageId="2" pageNumber="527">15.2</td>
<td id="104C5D92FFFCF6D6FC36CDFBFAE7F918" box="[939,1298,1711,1733]" gridcol="2" gridrow="26" pageId="2" pageNumber="527">15</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CD8CFAE7F933" box="[159,1298,1752,1774]" gridrow="27" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CD8CFDC7F933" box="[159,562,1752,1774]" gridcol="0" gridrow="27" pageId="2" pageNumber="527">First segmented anal ray</th>
<td id="104C5D92FFFCF6D6FD22CD8CFCA8F933" box="[703,861,1752,1774]" gridcol="1" gridrow="27" pageId="2" pageNumber="527">21.1</td>
<td id="104C5D92FFFCF6D6FC36CD8CFAE7F933" box="[939,1298,1752,1774]" gridcol="2" gridrow="27" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CC56FAE7F8C5" box="[159,1298,1794,1816]" gridrow="28" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CC56FDC7F8C5" box="[159,562,1794,1816]" gridcol="0" gridrow="28" pageId="2" pageNumber="527">Longest segmented anal ray (number)</th>
<td id="104C5D92FFFCF6D6FD22CC56FCA8F8C5" box="[703,861,1794,1816]" gridcol="1" gridrow="28" pageId="2" pageNumber="527">25.9 (3)</td>
<td id="104C5D92FFFCF6D6FC36CC56FAE7F8C5" box="[939,1298,1794,1816]" gridcol="2" gridrow="28" pageId="2" pageNumber="527">*</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CC7FFAE7F89C" box="[159,1298,1835,1857]" gridrow="29" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CC7FFDC7F89C" box="[159,562,1835,1857]" gridcol="0" gridrow="29" pageId="2" pageNumber="527">Caudal fin length</th>
<td id="104C5D92FFFCF6D6FD22CC7FFCA8F89C" box="[703,861,1835,1857]" gridcol="1" gridrow="29" pageId="2" pageNumber="527">45.1</td>
<td id="104C5D92FFFCF6D6FC36CC7FFAE7F89C" box="[939,1298,1835,1857]" gridcol="2" gridrow="29" pageId="2" pageNumber="527">27–29</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CC00FAE7F8B7" box="[159,1298,1876,1898]" gridrow="30" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CC00FDC7F8B7" box="[159,562,1876,1898]" gridcol="0" gridrow="30" pageId="2" pageNumber="527">Caudal concavity</th>
<td id="104C5D92FFFCF6D6FD22CC00FCA8F8B7" box="[703,861,1876,1898]" gridcol="1" gridrow="30" pageId="2" pageNumber="527">17.8</td>
<td id="104C5D92FFFCF6D6FC36CC00FAE7F8B7" box="[939,1298,1876,1898]" gridcol="2" gridrow="30" pageId="2" pageNumber="527">2</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CC29FAE7F84E" box="[159,1298,1917,1939]" gridrow="31" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CC29FDC7F84E" box="[159,562,1917,1939]" gridcol="0" gridrow="31" pageId="2" pageNumber="527">Pectoral fin length</th>
<td id="104C5D92FFFCF6D6FD22CC29FCA8F84E" box="[703,861,1917,1939]" gridcol="1" gridrow="31" pageId="2" pageNumber="527">28.3</td>
<td id="104C5D92FFFCF6D6FC36CC29FAE7F84E" box="[939,1298,1917,1939]" gridcol="2" gridrow="31" pageId="2" pageNumber="527">29–30</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CCF3FAE7F860" box="[159,1298,1959,1981]" gridrow="32" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CCF3FDC7F860" box="[159,562,1959,1981]" gridcol="0" gridrow="32" pageId="2" pageNumber="527">Pelvic fin spine</th>
<td id="104C5D92FFFCF6D6FD22CCF3FCA8F860" box="[703,861,1959,1981]" gridcol="1" gridrow="32" pageId="2" pageNumber="527">19.3</td>
<td id="104C5D92FFFCF6D6FC36CCF3FAE7F860" box="[939,1298,1959,1981]" gridcol="2" gridrow="32" pageId="2" pageNumber="527">17</td>
</tr>
<tr id="539D34EEFFFCF6D6FF02CC84FAE7F83B" box="[159,1298,2000,2022]" gridrow="33" pageId="2" pageNumber="527">
<th id="104C5D92FFFCF6D6FF02CC84FDC7F83B" box="[159,562,2000,2022]" gridcol="0" gridrow="33" pageId="2" pageNumber="527">Pelvic fin length</th>
<td id="104C5D92FFFCF6D6FD22CC84FCA8F83B" box="[703,861,2000,2022]" gridcol="1" gridrow="33" pageId="2" pageNumber="527">35.2</td>
<td id="104C5D92FFFCF6D6FC36CC84FAE7F83B" box="[939,1298,2000,2022]" gridcol="2" gridrow="33" pageId="2" pageNumber="527">28–31</td>
</tr>
</table>
</paragraph>
<caption id="B9D26624FFFD092AFF0ACBCCFB66FF72" box="[151,1171,151,175]" pageId="3" pageNumber="528">
<paragraph id="ED1236ACFFFD092AFF0ACBCCFB66FF72" blockId="3.[151,1171,151,175]" box="[151,1171,151,175]" pageId="3" pageNumber="528">
<emphasis id="DFD9EABEFFFD092AFF0ACBCCFEF6FF72" bold="true" box="[151,259,152,175]" pageId="3" pageNumber="528">TABLE 2</emphasis>
. Comparison of coloration features of male
<taxonomicName id="2AAD4D2FFFFD092AFD41CBC3FC47FF72" authority="Gill &amp; Psomadakis, 2018" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[732,946,151,175]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="emma" status="sp. nov.">
<emphasis id="DFD9EABEFFFD092AFD41CBC3FC47FF72" box="[732,946,151,175]" italics="true" pageId="3" pageNumber="528">Pseudanthias emma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4EA57C5FFFD092AFC25CBCCFBFBFF72" box="[952,1038,152,175]" pageId="3" pageNumber="528" rank="species">
<emphasis id="DFD9EABEFFFD092AFC25CBCCFBFBFF72" bold="true" box="[952,1038,152,175]" pageId="3" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="2AAD4D2FFFFD092AFBDCCBCCFB66FF72" box="[1089,1171,151,175]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="pillai">P. pillai</taxonomicName>
</paragraph>
</caption>
<paragraph id="ED1236ACFFFD092AFE78CB9FFAADFCB5" pageId="3" pageNumber="528">
<table id="9FADC40CFFFDF6D6FF02CB9EFA63FC5A" box="[159,1430,202,903]" gridcols="4" gridrows="11" pageId="3" pageNumber="528">
<tr id="539D34EEFFFDF6D6FF02CB9EFA63FF3D" box="[159,1430,202,224]" gridrow="0" pageId="3" pageNumber="528" rowspan-0="1" rowspan-3="1">
<th id="104C5D92FFFDF6D6FE78CB9EFCB6FF3D" box="[485,835,202,224]" gridcol="1" gridrow="0" pageId="3" pageNumber="528">
<taxonomicName id="2AAD4D2FFFFD092AFE78CB9FFDCFFF3D" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[485,570,203,224]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFD092AFE78CB9FFDCFFF3D" box="[485,570,203,224]" italics="true" pageId="3" pageNumber="528">P. emma</emphasis>
</taxonomicName>
</th>
<th id="104C5D92FFFDF6D6FCC7CB9EFB8CFF3D" box="[858,1145,202,224]" gridcol="2" gridrow="0" pageId="3" pageNumber="528">
<taxonomicName id="2AAD4D2FFFFD092AFCC7CB9FFC5DFF3D" box="[858,936,202,224]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFD092AFCC7CB9FFC5DFF3D" box="[858,936,202,224]" italics="true" pageId="3" pageNumber="528">P. pillai</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="539D34EEFFFDF6D6FF02CBA0FA63FEF5" box="[159,1430,244,296]" gridrow="1" pageId="3" pageNumber="528" rowspan-0="1" rowspan-1="1">
<td id="104C5D92FFFDF6D6FCC7CBA0FB8CFEF5" box="[858,1145,244,296]" gridcol="2" gridrow="1" pageId="3" pageNumber="528">Heemstra &amp; Akhilesh 2012: fig. 40</td>
<td id="104C5D92FFFDF6D6FB0FCBA0FA63FEF5" box="[1170,1430,244,296]" gridcol="3" gridrow="1" pageId="3" pageNumber="528">
Krishna
<emphasis id="DFD9EABEFFFD092AFB75CBA1FAEEFED7" box="[1256,1307,244,266]" italics="true" pageId="3" pageNumber="528">et al.</emphasis>
2017: fig. 1
</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02CA68FA63FEAD" box="[159,1430,316,368]" gridrow="2" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02CA68FE39FEAD" box="[159,460,316,368]" gridcol="0" gridrow="2" pageId="3" pageNumber="528">Dark spots on scales of upper body and nape</th>
<td id="104C5D92FFFDF6D6FE78CA68FCB6FEAD" box="[485,835,316,368]" gridcol="1" gridrow="2" pageId="3" pageNumber="528">absent</td>
<td id="104C5D92FFFDF6D6FCC7CA68FB8CFEAD" box="[858,1145,316,368]" gridcol="2" gridrow="2" pageId="3" pageNumber="528">present</td>
<td id="104C5D92FFFDF6D6FB0FCA68FA63FEAD" box="[1170,1430,316,368]" gridcol="3" gridrow="2" pageId="3" pageNumber="528">present</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02CAD0FA63FE65" box="[159,1430,388,440]" gridrow="3" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02CAD0FE39FE65" box="[159,460,388,440]" gridcol="0" gridrow="3" pageId="3" pageNumber="528">Sides of body</th>
<td id="104C5D92FFFDF6D6FE78CAD0FCB6FE65" box="[485,835,388,440]" gridcol="1" gridrow="3" pageId="3" pageNumber="528">mostly orange</td>
<td id="104C5D92FFFDF6D6FCC7CAD0FB8CFE65" box="[858,1145,388,440]" gridcol="2" gridrow="3" pageId="3" pageNumber="528">orange with bright pink stripes</td>
<td id="104C5D92FFFDF6D6FB0FCAD0FA63FE65" box="[1170,1430,388,440]" gridcol="3" gridrow="3" pageId="3" pageNumber="528">orange with bright pink stripes</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02CA98FA63FDDD" box="[159,1430,460,512]" gridrow="4" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02CA98FE39FDDD" box="[159,460,460,512]" gridcol="0" gridrow="4" pageId="3" pageNumber="528">Soft part of dorsal fin</th>
<td id="104C5D92FFFDF6D6FE78CA98FCB6FDDD" box="[485,835,460,512]" gridcol="1" gridrow="4" pageId="3" pageNumber="528">orange, bright red distally</td>
<td id="104C5D92FFFDF6D6FCC7CA98FB8CFDDD" box="[858,1145,460,512]" gridcol="2" gridrow="4" pageId="3" pageNumber="528">bright yellow, rays purple or yellow</td>
<td id="104C5D92FFFDF6D6FB0FCA98FA63FDDD" box="[1170,1430,460,512]" gridcol="3" gridrow="4" pageId="3" pageNumber="528">bright yellow, rays purple or yellow</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C940FA63FD95" box="[159,1430,532,584]" gridrow="5" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02C940FE39FD95" box="[159,460,532,584]" gridcol="0" gridrow="5" pageId="3" pageNumber="528">Pink band along upper part of body</th>
<td id="104C5D92FFFDF6D6FE78C940FCB6FD95" box="[485,835,532,584]" gridcol="1" gridrow="5" pageId="3" pageNumber="528">distinct</td>
<td id="104C5D92FFFDF6D6FCC7C940FB8CFD95" box="[858,1145,532,584]" gridcol="2" gridrow="5" pageId="3" pageNumber="528">diffuse</td>
<td id="104C5D92FFFDF6D6FB0FC940FA63FD95" box="[1170,1430,532,584]" gridcol="3" gridrow="5" pageId="3" pageNumber="528">diffuse</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C908FA63FDAF" box="[159,1430,604,626]" gridrow="6" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02C908FE39FDAF" box="[159,460,604,626]" gridcol="0" gridrow="6" pageId="3" pageNumber="528">Orange-red blotch on breast</th>
<td id="104C5D92FFFDF6D6FE78C908FCB6FDAF" box="[485,835,604,626]" gridcol="1" gridrow="6" pageId="3" pageNumber="528">present</td>
<td id="104C5D92FFFDF6D6FCC7C908FB8CFDAF" box="[858,1145,604,626]" gridcol="2" gridrow="6" pageId="3" pageNumber="528">absent</td>
<td id="104C5D92FFFDF6D6FB0FC908FA63FDAF" box="[1170,1430,604,626]" gridcol="3" gridrow="6" pageId="3" pageNumber="528">absent</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C9D1FA63FD05" box="[159,1430,645,728]" gridrow="7" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02C9D1FE39FD05" box="[159,460,645,728]" gridcol="0" gridrow="7" pageId="3" pageNumber="528">Caudal fin</th>
<td id="104C5D92FFFDF6D6FE78C9D1FCB6FD05" box="[485,835,645,728]" gridcol="1" gridrow="7" pageId="3" pageNumber="528">pinkish red on basal third, pinkish hyaline on middle third, and bright red on distal third</td>
<td id="104C5D92FFFDF6D6FCC7C9D1FB8CFD05" box="[858,1145,645,728]" gridcol="2" gridrow="7" pageId="3" pageNumber="528">magenta with posterior margin bright yellow</td>
<td id="104C5D92FFFDF6D6FB0FC9D1FA63FD05" box="[1170,1430,645,728]" gridcol="3" gridrow="7" pageId="3" pageNumber="528">magenta with posterior margin bright yellow</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C9B8FA63FCDF" box="[159,1430,748,770]" gridrow="8" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02C9B8FE39FCDF" box="[159,460,748,770]" gridcol="0" gridrow="8" pageId="3" pageNumber="528">Anal fin</th>
<td id="104C5D92FFFDF6D6FE78C9B8FCB6FCDF" box="[485,835,748,770]" gridcol="1" gridrow="8" pageId="3" pageNumber="528">pale pink to white, with distal</td>
<td id="104C5D92FFFDF6D6FCC7C9B8FB8CFCDF" box="[858,1145,748,770]" gridcol="2" gridrow="8" pageId="3" pageNumber="528">bright pinkish purple with</td>
<td id="104C5D92FFFDF6D6FB0FC9B8FA63FCDF" box="[1170,1430,748,770]" gridcol="3" gridrow="8" pageId="3" pageNumber="528">bright pinkish purple</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C85EFA63FCE2" box="[159,1430,778,831]" gridrow="9" pageId="3" pageNumber="528" rowspan-0="1">
<td id="104C5D92FFFDF6D6FE78C85EFB8CFCE2" box="[485,1145,778,831]" colspan="2" colspanRight="1" gridcol="1" gridrow="9" pageId="3" pageNumber="528">portion of soft part of fin bright red some yellow, pale pink anteriorly</td>
<td id="104C5D92FFFDF6D6FB0FC85EFA63FCE2" box="[1170,1430,778,831]" gridcol="3" gridrow="9" pageId="3" pageNumber="528">with some yellow, pale pink anteriorly</td>
</tr>
<tr id="539D34EEFFFDF6D6FF02C806FA63FC5A" box="[159,1430,850,903]" gridrow="10" pageId="3" pageNumber="528">
<th id="104C5D92FFFDF6D6FF02C806FE39FC5A" box="[159,460,850,903]" gridcol="0" gridrow="10" pageId="3" pageNumber="528">Pelvic fin</th>
<td id="104C5D92FFFDF6D6FE78C806FCB6FC5A" box="[485,835,850,903]" gridcol="1" gridrow="10" pageId="3" pageNumber="528">pale pink to white with distal part abruptly bright red</td>
<td id="104C5D92FFFDF6D6FCC7C806FB8CFC5A" box="[858,1145,850,903]" gridcol="2" gridrow="10" pageId="3" pageNumber="528">pale yellow to pale pink</td>
<td id="104C5D92FFFDF6D6FB0FC806FA63FC5A" box="[1170,1430,850,903]" gridcol="3" gridrow="10" pageId="3" pageNumber="528">pale yellowish pink</td>
</tr>
</table>
</paragraph>
<paragraph id="ED1236ACFFFD092AFF5AC893FB70F9FD" blockId="3.[151,1437,967,2037]" pageId="3" pageNumber="528">
Colour of male
<typeStatus id="3216880EFFFD092AFE1DC893FE17FC3D" box="[384,482,967,992]" pageId="3" pageNumber="528" type="holotype">holotype</typeStatus>
when freshly dead (described from two photos, one with a white background that shows fin coloration better but is not of publishable quality, and
<figureCitation id="75962A29FFFD092AFC1AC8B8FC1DFBD9" box="[903,1000,1004,1029]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[151,1436,1123,1873]" captionTargetId="figure@4.[151,1436,1123,1873]" captionTargetPageId="4" captionText="FIGURE 1. Pseudanthias emma sp. nov., SAIAB 203722, 107 mm SL, male holotype, off Tanintharyi coast, Myanmar, Andaman Sea, Indian Ocean. Note that filamentous tips of caudal fin were cropped during photography. Photo by Oddgeir Alvheim." httpUri="https://zenodo.org/record/1457511/files/figure.png" pageId="3" pageNumber="528">Figure 1</figureCitation>
): head pink, pale pink ventrally, with orange bar extending almost vertically from about 1 o’clock position of orbital rim over nape; two orange stripes, one narrow and from front of eye to snout tip, the other broad and from behind lower half of eye to pectoral-fin base, the stripes bordered ventrally by continuous bright pink (on head) to silvery pink (on pectoral-fin base) stripe; iris bright yellow, with short black crescent near anterior border of eye; body mostly orange, with bright pink band from dorsal-fin origin along dorsal part of body to upper edge of caudal peduncle, the band broader beneath middle of dorsal fin, extending ventrally almost to lateral line; lower part of body silvery white anteriorly on breast, becoming pale pink on lower part of caudal peduncle, separated from orange part of body by narrow silvery white stripe; a large (eye-sized) orange-red blotch on breast, between bases of pectoral and pelvic fins; spinous part of dorsal fin bright orange-yellow with fins spines pink, and distal part of fin behind seventh spine narrowly bright red; soft portion of dorsal fin orange with distal third to half of fin bright red; anal fin pale pink to white, with distal half of soft part of fin bright red; caudal fin narrowly bright pink dorsally and pale pink ventrally, the remainder of fin pinkish red on basal third, pinkish hyaline on middle third, and bright red on distal third, with filamentous tips pale pink to hyaline; pelvic fin pale pink to white with distal tip abruptly bright red; orange stripe on pectoral fin base extends on to fin, becoming reddish orange, this bordered dorsally and ventrally by narrow bright pink (dorsally) or silvery pink to white (ventrally) stripes, the remainder of fin pinkish hyaline.
</paragraph>
<paragraph id="ED1236ACFFFD092AFF5ACD78FABBF998" blockId="3.[151,1437,967,2037]" box="[199,1358,1580,1605]" pageId="3" pageNumber="528">Colour in preservative: Head and body tan, with no apparent markings; fins pale tan, without markings.</paragraph>
<paragraph id="ED1236ACFFFD092AFF5ACD1BFF3AF925" blockId="3.[151,1437,967,2037]" pageId="3" pageNumber="528">
<emphasis id="DFD9EABEFFFD092AFF5ACD1BFE00F9B5" bold="true" box="[199,501,1615,1640]" pageId="3" pageNumber="528">Habitat and distribution.</emphasis>
Known only from the
<typeStatus id="3216880EFFFD092AFD67CD1BFCAAF9B5" box="[762,863,1615,1640]" pageId="3" pageNumber="528" type="holotype">holotype</typeStatus>
, trawled in proximity of a deep coral reef (
<quantity id="2A559B49FFFD092AFAD7CD1BFA61F9B5" box="[1354,1428,1615,1640]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="528" unit="m" value="105.0">105 m</quantity>
) off the south east coast of
<collectingCountry id="95BA763CFFFD092AFE5FCD21FDDAF950" box="[450,559,1653,1677]" name="Myanmar" pageId="3" pageNumber="528">Myanmar</collectingCountry>
in the Andaman Sea (eastern Indian Ocean). The chaetodontids
<taxonomicName id="2AAD4D2FFFFD092AFA97CD21FEF3F96D" authority="(Norman)" authorityName="Norman" baseAuthorityName="Norman" baseAuthorityYear="1939" class="Actinopterygii" family="Chaetodontidae" genus="Roa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="jayakari">
<emphasis id="DFD9EABEFFFD092AFA97CD21FA69F951" box="[1290,1436,1652,1676]" italics="true" pageId="3" pageNumber="528">Roa jayakari</emphasis>
(Norman)
</taxonomicName>
and
<taxonomicName id="2AAD4D2FFFFD092AFED0CDCCFD14F96D" authority="(Linnaeus)" authorityName="Linnaeus" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[333,737,1687,1712]" class="Actinopterygii" family="Chaetodontidae" genus="Heniochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="DFD9EABEFFFD092AFED0CDCCFDA0F972" box="[333,597,1688,1712]" italics="true" pageId="3" pageNumber="528">Heniochus acuminatus</emphasis>
(Linnaeus)
</taxonomicName>
, the priacanthid
<taxonomicName id="2AAD4D2FFFFD092AFC29CDCCFA93F96D" authority="(Valenciennes)" authorityName="Valenciennes" baseAuthorityName="Valenciennes" baseAuthorityYear="1862" box="[948,1382,1687,1712]" class="Actinopterygii" family="Priacanthidae" genus="Pristigenys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="refulgens">
<emphasis id="DFD9EABEFFFD092AFC29CDCCFB5FF96D" box="[948,1194,1688,1712]" italics="true" pageId="3" pageNumber="528">Pristigenys refulgens</emphasis>
(Valenciennes)
</taxonomicName>
, the lethrinid
<taxonomicName id="2AAD4D2FFFFD092AFF60CDE9FDB4F908" authority="(Smith)" authorityName="Smith" baseAuthorityName="Smith" baseAuthorityYear="1957" box="[253,577,1724,1749]" class="Actinopterygii" family="Lethrinidae" genus="Wattsia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="mossambica">
<emphasis id="DFD9EABEFFFD092AFF60CDE9FE10F909" box="[253,485,1724,1748]" italics="true" pageId="3" pageNumber="528">Wattsia mossambica</emphasis>
(Smith)
</taxonomicName>
as well as several other reef-associated fish species were caught in the same haul.
</paragraph>
<paragraph id="ED1236ACFFFD092DFF5ACC50FE89FEC0" blockId="3.[151,1437,967,2037]" lastBlockId="4.[151,1437,151,1040]" lastPageId="4" lastPageNumber="529" pageId="3" pageNumber="528">
<emphasis id="DFD9EABEFFFD092AFF5ACC50FE98F8C0" bold="true" box="[199,365,1796,1821]" pageId="3" pageNumber="528">Comparisons.</emphasis>
<taxonomicName id="2AAD4D2FFFFD092AFEE8CC50FDADF8C1" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[373,600,1796,1820]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFD092AFEE8CC50FDADF8C1" box="[373,600,1796,1820]" italics="true" pageId="3" pageNumber="528">Pseudanthias emma</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="2AAD4D2FFFFD092AFCAECC51FB17F8C1" authority="Heemstra &amp; Akhilesh (2012)" authorityName="Heemstra &amp; Akhilesh" authorityYear="2012" box="[819,1250,1796,1821]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFD092AFCAECC51FC79F8C1" box="[819,908,1796,1820]" italics="true" pageId="3" pageNumber="528">P. pillai</emphasis>
<bibRefCitation id="893C4B5DFFFD092AFC08CC51FB17F8C1" box="[917,1250,1796,1821]" pageId="3" pageNumber="528" refId="ref5073" refString="Heemstra, P. C. &amp; Akhilesh, K. V. (2012) A review of the anthiine fish genus Pseudanthias (Perciformes: Serranidae) of the western Indian Ocean, with description of a new species and a key to the species. aqua, International Journal of Ichthyology, 18, 121 - 164." type="journal article">Heemstra &amp; Akhilesh (2012)</bibRefCitation>
</taxonomicName>
in general form and coloration. The latter species was described on the basis of two specimens from off Charakkadu,
<collectingRegion id="2F69F84EFFFD092AFA97CC73FAA2F89D" box="[1290,1367,1831,1856]" country="India" name="Kerala" pageId="3" pageNumber="528">Kerala</collectingRegion>
,
<collectingCountry id="95BA763CFFFD092AFAFFCC73FA69F89D" box="[1378,1436,1831,1856]" name="India" pageId="3" pageNumber="528">India</collectingCountry>
(
<geoCoordinate id="8899506BFFFD092AFF3DCC18FEFBF8B9" box="[160,270,1868,1892]" direction="north" orientation="latitude" pageId="3" pageNumber="528" precision="925" value="10.5">10°30’ N</geoCoordinate>
,
<geoCoordinate id="8899506BFFFD092AFE81CC18FE8BF8B8" box="[284,382,1868,1893]" direction="east" orientation="longitude" pageId="3" pageNumber="528" precision="925" value="75.4">75°24’E</geoCoordinate>
). Subsequently,
<bibRefCitation id="893C4B5DFFFD092AFDA6CC18FCDBF8B9" box="[571,814,1868,1893]" pageId="3" pageNumber="528" refId="ref5275" refString="Krishna, N. M., Rao, V. G. &amp; Venu, D. (2017) Pseudanthias vizagensis sp. nov., a new anthiine fish (subfamily: Anthiinae), genus Pseudanthias, from India. Journal of Experimental Zoology, India, 20 (1), 213 - 216." type="journal article">
Krishna
<emphasis id="DFD9EABEFFFD092AFD03CC19FD2CF8B9" box="[670,729,1868,1892]" italics="true" pageId="3" pageNumber="528">et al.</emphasis>
(2017)
</bibRefCitation>
described
<taxonomicName id="2AAD4D2FFFFD092AFC33CC19FBB5F8B9" authorityName="Krishna, Rao &amp; Venu" authorityYear="2017" box="[942,1088,1869,1892]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="vizagensis">
<emphasis id="DFD9EABEFFFD092AFC33CC19FBB5F8B9" box="[942,1088,1869,1892]" italics="true" pageId="3" pageNumber="528">P. vizagensis</emphasis>
</taxonomicName>
on the basis of
<specimenCount id="FBABFD25FFFD092AFA9CCC18FA69F8B8" box="[1281,1436,1868,1893]" pageId="3" pageNumber="528" type="generic">44 specimens</specimenCount>
trawled off Visakhapatnam in the Bay of Bengal (stated coordinates
<geoCoordinate id="8899506BFFFD092AFC00CC3BFC00F855" box="[925,1013,1903,1928]" direction="north" orientation="latitude" pageId="3" pageNumber="528" precision="925" value="17.166666">17°10’N</geoCoordinate>
,
<geoCoordinate id="8899506BFFFD092AFB9CCC24FB95F855" box="[1025,1120,1904,1928]" direction="east" orientation="longitude" pageId="3" pageNumber="528" precision="925" value="18.166666">18°10’E</geoCoordinate>
, which is clearly erroneous; possibly
<geoCoordinate id="8899506BFFFD092AFF66CCC0FEADF871" box="[251,344,1940,1964]" direction="east" orientation="longitude" pageId="3" pageNumber="528" precision="925" value="83.166664">83°10’E</geoCoordinate>
was the intended longitude value). The description of
<taxonomicName id="2AAD4D2FFFFD092AFC2ACCC1FBB4F871" authorityName="Krishna, Rao &amp; Venu" authorityYear="2017" box="[951,1089,1941,1964]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="vizagensis">
<emphasis id="DFD9EABEFFFD092AFC2ACCC1FBB4F871" box="[951,1089,1941,1964]" italics="true" pageId="3" pageNumber="528">P. vizagensis</emphasis>
</taxonomicName>
is problematic, not the least of which for the listing of all
<specimenCount id="FBABFD25FFFD092AFE41CCECFD8DF80D" box="[476,632,1975,2000]" pageId="3" pageNumber="528" type="generic">44 specimens</specimenCount>
as “
<typeStatus id="3216880EFFFD092AFD2DCCE3FCD1F80D" box="[688,804,1975,2000]" pageId="3" pageNumber="528" type="holotype">holotypes</typeStatus>
”. Although
<bibRefCitation id="893C4B5DFFFD092AFC2ECCE3FB5CF80D" box="[947,1193,1975,2000]" pageId="3" pageNumber="528" refId="ref5275" refString="Krishna, N. M., Rao, V. G. &amp; Venu, D. (2017) Pseudanthias vizagensis sp. nov., a new anthiine fish (subfamily: Anthiinae), genus Pseudanthias, from India. Journal of Experimental Zoology, India, 20 (1), 213 - 216." type="journal article">
Krishna
<emphasis id="DFD9EABEFFFD092AFB8ACCEDFBA1F80D" box="[1047,1108,1976,2000]" italics="true" pageId="3" pageNumber="528">et al.</emphasis>
(2017)
</bibRefCitation>
compared their new species with
<taxonomicName id="2AAD4D2FFFFD092AFEB5CC89FE8AF829" box="[296,383,2012,2036]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="528" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFD092AFEB5CC89FE8AF829" box="[296,383,2012,2036]" italics="true" pageId="3" pageNumber="528">P. pillai</emphasis>
</taxonomicName>
, purported differences in coloration and fin shapes are erroneous, and apparent differences in meristic characters (17 versus 19 pectoral-fin rays; 44–45 versus 36–38 lateral-line scales) may reflect different or inaccurate counting methods. We here suggest that the two nominal species are synonyms, though re-examination of the
<taxonomicName id="2AAD4D2FFFFA092DFF7ACBB5FE83FF2A" authorityName="Krishna, Rao &amp; Venu" authorityYear="2017" box="[231,374,224,248]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="vizagensis">
<emphasis id="DFD9EABEFFFA092DFF7ACBB5FE83FF2A" box="[231,374,224,248]" italics="true" pageId="4" pageNumber="529">P. vizagensis</emphasis>
</taxonomicName>
<typeStatus id="3216880EFFFA092DFE1BCBB4FE43FF25" box="[390,438,224,248]" pageId="4" pageNumber="529">type</typeStatus>
specimens (here interpreted as
<typeStatus id="3216880EFFFA092DFCB3CBB4FC67FF25" box="[814,914,224,248]" pageId="4" pageNumber="529" type="syntype">syntypes</typeStatus>
) will be necessary to reconcile the apparent meristic differences.
</paragraph>
<paragraph id="ED1236ACFFFA092DFF5ACA73FEADFCC9" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="529">
Comparison of certain morphometric data provided by
<bibRefCitation id="893C4B5DFFFA092DFCACCA73FBEFFE9D" box="[817,1050,295,320]" pageId="4" pageNumber="529" refId="ref5275" refString="Krishna, N. M., Rao, V. G. &amp; Venu, D. (2017) Pseudanthias vizagensis sp. nov., a new anthiine fish (subfamily: Anthiinae), genus Pseudanthias, from India. Journal of Experimental Zoology, India, 20 (1), 213 - 216." type="journal article">
Krishna
<emphasis id="DFD9EABEFFFA092DFC0CCA7DFC3CFE9D" box="[913,969,296,320]" italics="true" pageId="4" pageNumber="529">et al.</emphasis>
(2017)
</bibRefCitation>
with their figure 1A indicates that their data are either not compatible or not sufficiently accurate for comparison with
<taxonomicName id="2AAD4D2FFFFA092DFBF4CA19FB32FEB9" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[1129,1223,333,356]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFBF4CA19FB32FEB9" box="[1129,1223,333,356]" italics="true" pageId="4" pageNumber="529">P. emma</emphasis>
</taxonomicName>
(e.g., head length 40.44–40.65 stated vs ca 36 % SL from figure, predorsal length 38.20–39.56 vs ca 32 % SL, greatest body depth 39.56–44.94 vs ca 37 % SL). We therefore base our morphometric comparison solely on data provided by
<bibRefCitation id="893C4B5DFFFA092DFF0ACAECFE1FFE0D" box="[151,490,439,464]" pageId="4" pageNumber="529" refId="ref5073" refString="Heemstra, P. C. &amp; Akhilesh, K. V. (2012) A review of the anthiine fish genus Pseudanthias (Perciformes: Serranidae) of the western Indian Ocean, with description of a new species and a key to the species. aqua, International Journal of Ichthyology, 18, 121 - 164." type="journal article">Heemstra &amp; Akhilesh (2012)</bibRefCitation>
, though acknowledge that larger sample sizes are needed for proper comparison. Nonetheless,
<taxonomicName id="2AAD4D2FFFFA092DFEB0CA89FE7CFE29" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[301,393,477,500]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFEB0CA89FE7CFE29" box="[301,393,477,500]" italics="true" pageId="4" pageNumber="529">P. emma</emphasis>
</taxonomicName>
appears to differ substantially from
<taxonomicName id="2AAD4D2FFFFA092DFCBECA89FC8CFE29" box="[803,889,476,500]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFA092DFCBECA89FC8CFE29" box="[803,889,476,500]" italics="true" pageId="4" pageNumber="529">P. pillai</emphasis>
</taxonomicName>
in having a deeper caudal peduncle, longer third dorsal-fin spine, shorter head, broader interorbital, and shorter soft dorsal-fin rays (
<tableCitation id="A02F0317FFFA092DFBD0CAABFB57FDC5" box="[1101,1186,511,536]" captionStart="TABLE 1" captionStartId="2.[151,239,500,523]" captionText="TABLE 1. Morphometric values for Pseudanthias emma sp. nov. expressed as percentage SL, with comparable data for P. pillai (after Heemstra &amp; Akhilesh 2012). * indicates where data for P. pillai is lacking or not comparable." pageId="4" pageNumber="529">Table 1</tableCitation>
). Although similar in coloration, the two species differ in various details (
<tableCitation id="A02F0317FFFA092DFD7EC970FCCEFDE1" box="[739,827,548,573]" captionStart="TABLE 2" captionStartId="3.[151,239,152,175]" captionText="TABLE 2. Comparison of coloration features of male Pseudanthias emma sp. nov. and P. pillai" pageId="4" pageNumber="529">Table 2</tableCitation>
). They also differ in caudal-fin shape. In
<taxonomicName id="2AAD4D2FFFFA092DFA8FC971FA87FDE1" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[1298,1394,549,572]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFA8FC971FA87FDE1" box="[1298,1394,549,572]" italics="true" pageId="4" pageNumber="529">P. emma</emphasis>
</taxonomicName>
the caudal fin is weakly concave with the second-to outermost branched rays dorsally and ventrally expanded with long filaments, whereas in
<taxonomicName id="2AAD4D2FFFFA092DFE4CC939FDDFFD59" box="[465,554,620,644]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFA092DFE4CC939FDDFFD59" box="[465,554,620,644]" italics="true" pageId="4" pageNumber="529">P. pillai</emphasis>
</taxonomicName>
it is weakly convex centrally, with little or no elongation of outer rays (not lunate as stated by
<bibRefCitation id="893C4B5DFFFA092DFEECC9DBFDA5FD75" box="[369,592,655,680]" pageId="4" pageNumber="529" refId="ref5275" refString="Krishna, N. M., Rao, V. G. &amp; Venu, D. (2017) Pseudanthias vizagensis sp. nov., a new anthiine fish (subfamily: Anthiinae), genus Pseudanthias, from India. Journal of Experimental Zoology, India, 20 (1), 213 - 216." type="journal article">
Krishna
<emphasis id="DFD9EABEFFFA092DFE4FC9C5FDFEFD75" box="[466,523,656,680]" italics="true" pageId="4" pageNumber="529">et al.</emphasis>
2017
</bibRefCitation>
). The two species differ in the number of lower gill rakers (
<quantity id="2A559B49FFFA092DFB64C9C4FAC1FD75" box="[1273,1332,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="4" pageNumber="529" unit="in" value="22.0">22 in</quantity>
<taxonomicName id="2AAD4D2FFFFA092DFAA1C9C5FA69FD75" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[1340,1436,657,680]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFAA1C9C5FA69FD75" box="[1340,1436,657,680]" italics="true" pageId="4" pageNumber="529">P. emma</emphasis>
</taxonomicName>
versus
<quantity id="2A559B49FFFA092DFF75C9E0FEBBFD10" box="[232,334,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.366" metricValueMin="7.112" pageId="4" pageNumber="529" unit="in" value="28.5" valueMax="29.0" valueMin="28.0">28–29 in</quantity>
<taxonomicName id="2AAD4D2FFFFA092DFECAC9E1FE45FD11" box="[343,432,692,716]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFA092DFECAC9E1FE45FD11" box="[343,432,692,716]" italics="true" pageId="4" pageNumber="529">P. pillai</emphasis>
</taxonomicName>
). Finally,
<taxonomicName id="2AAD4D2FFFFA092DFDB5C9E1FD7CFD11" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[552,649,693,716]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFDB5C9E1FD7CFD11" box="[552,649,693,716]" italics="true" pageId="4" pageNumber="529">P. emma</emphasis>
</taxonomicName>
possibly differs from
<taxonomicName id="2AAD4D2FFFFA092DFC11C9E1FC10FD11" box="[908,997,692,716]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="pillai">
<emphasis id="DFD9EABEFFFA092DFC11C9E1FC10FD11" box="[908,997,692,716]" italics="true" pageId="4" pageNumber="529">P. pillai</emphasis>
</taxonomicName>
in having more lateral-line scales (42 versus 36-38), though this assumes the high numbers (44–45) reported for the
<typeStatus id="3216880EFFFA092DFB99C98CFB93FD2D" box="[1028,1126,728,752]" pageId="4" pageNumber="529" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="2AAD4D2FFFFA092DFB16C98DFAECFD32" authorityName="Krishna, Rao &amp; Venu" authorityYear="2017" box="[1163,1305,728,752]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="vizagensis">
<emphasis id="DFD9EABEFFFA092DFB16C98DFAECFD32" box="[1163,1305,728,752]" italics="true" pageId="4" pageNumber="529">P. vizagensis</emphasis>
</taxonomicName>
by Krishna
<emphasis id="DFD9EABEFFFA092DFF0AC9A9FF3AFCC9" box="[151,207,764,788]" italics="true" pageId="4" pageNumber="529">et al.</emphasis>
are in error.
</paragraph>
<paragraph id="ED1236ACFFFA092DFF5AC874FF1AFC30" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="529">
<taxonomicName id="2AAD4D2FFFFA092DFF5AC874FE45FCE5" authorityName="Gill &amp; Psomadakis" authorityYear="2018" box="[199,432,800,824]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="529" phylum="Chordata" rank="species" species="emma">
<emphasis id="DFD9EABEFFFA092DFF5AC874FE45FCE5" box="[199,432,800,824]" italics="true" pageId="4" pageNumber="529">Pseudanthias emma</emphasis>
</taxonomicName>
is distinguished from all other congeners in having the following combination of characters: dorsal-fin rays X,16, the third spine prolonged; pectoral rays 18; lateral-line scales 42; gill rakers 11 + 22; no papillae on posterior margin on orbit; soft part of dorsal fin with low scaly sheath; subopercle and interopercle indistinctly serrated; and caudal fin weakly concave centrally, with elongate filaments extending from second to uppermost and second to lowermost branched rays. The fresh coloration is also distinctive for the species.
</paragraph>
</subSubSection>
<subSubSection id="A5B76527FFFA092DFF5AC8A3FA68FBCD" box="[199,1437,1015,1040]" pageId="4" pageNumber="529" type="etymology">
<paragraph id="ED1236ACFFFA092DFF5AC8A3FA68FBCD" blockId="4.[151,1437,151,1040]" box="[199,1437,1015,1040]" pageId="4" pageNumber="529">
<emphasis id="DFD9EABEFFFA092DFF5AC8A3FEBBFBCD" bold="true" box="[199,334,1015,1040]" pageId="4" pageNumber="529">Etymology.</emphasis>
The species epithet is for the second author’s daughter Emma. It is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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