treatments-xml/data/D2/8E/4D/D28E4D4457E156949D4805D7490BEAE0.xml

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<mods:title id="22C9821098A5B0CBF73DA243B297B77F">Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae)</mods:title>
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<mods:namePart id="B3BAD698A0893433509E6208F055A866">Chang, Wan-Jin</mods:namePart>
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<mods:namePart id="C1EC340D9DC0CE61D997505EA9DE39F9">Li, Shuqiang</mods:namePart>
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<treatment id="D28E4D4457E156949D4805D7490BEAE0" ID-GBIF-Taxon="162174134" LSID="urn:lsid:zoobank.org:act:8B33B61D-7B2C-4632-A1D8-9A6F335B04F3" httpUri="http://treatment.plazi.org/id/D28E4D4457E156949D4805D7490BEAE0" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Psilodercidae" scope_order="Araneae">
<subSubSection id="6D755F45A4A9170758FB0F64AD80A741" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="D275FBED6C51011824F4D6AE09D99A8A" pageId="0" pageNumber="1">
<taxonomicName id="19DEFDB2B380DE000F9E11266F0C03FD" ID-CoL="98ZTS" LSID="D28E4D44-57E1-5694-9D48-05D7490BEAE0" authority="Chang &amp; Li, 2020" authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera hponensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hponensis" status="sp. nov.">Leclercera hponensis</taxonomicName>
<taxonomicNameLabel id="D41ED96EC126561C98643D2E407EB2E9" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="1EAB5D2B243E9AB957EC76FB9B39A0D2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">Figs 25</figureCitation>
<figureCitation id="2203535C2B0DEC389E2A8CB220AE30A2" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">, 26</figureCitation>
<figureCitation id="65527E18D39AB47A49FB47A6EBE13DD6" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">, 56F</figureCitation>
<figureCitation id="536B8E62F982A80069BA64B2687647AB" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">, 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C86CFD040D3C71DDB5BA324D13B040F3" pageId="0" pageNumber="1" type="types">
<paragraph id="5A057AE941CAA83A84FAD0C913B8A1DA" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="07796FE6DB03A02651F4D7C6792ECB5E" pageId="0" pageNumber="1">
<emphasis id="B9D1390193FA1BA2147AC20DDFF23493" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂ (IZCAS), Myanmar, Kachin State, Putao Town, Hponkanrazi Wildlife Sanctuary Roadside between Camp 2 to Camp 3, 
<geoCoordinate id="ADEBFB04DF29DCA9CCBFE5D215481DD4" degrees="27" direction="north" minutes="37.1500" orientation="latitude" precision="1" value="27.619167">27°37.1500'N</geoCoordinate>
, 
<geoCoordinate id="02DF369241554C92288B9415550FFAD7" degrees="96" direction="east" minutes="58.9170" orientation="longitude" precision="1" value="96.98195">96°58.9170'E</geoCoordinate>
, elevation ca 2806 m, 16.XII.2016, J. Wu leg. 
<emphasis id="34FA7A5C8AB91FD4EDBF344AE15ED727" bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
: 1♀ (IZCAS), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="84096E1F3DA0A1A60BF6A3B6A4831386" pageId="0" pageNumber="1" type="etymology">
<paragraph id="6CB6C08FB3B9692DAF73E7F255AF2EDA" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="89235C295F7362C754A4C5A970F33095" pageId="0" pageNumber="1">The species name is an adjective referring to the type locality.</paragraph>
</subSubSection>
<subSubSection id="B277FD828875BE5EB2799655A79B2295" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="3A1736887BD2210F0FB9A88B3AAFA444" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="9D519D63E98E026F98082873C867F4AB" pageId="0" pageNumber="1">
Males of 
<taxonomicName id="33608B562CB9779BA7D7FA2FC89878DA" lsidName="L. hponensis" pageId="0" pageNumber="1" rank="species" species="hponensis">
<emphasis id="7C424BDA170CF4FD5BE8B8FF837F3F97" italics="true" pageId="0" pageNumber="1">L. hponensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from congeners by the presence of a three-branched, laminar apophysis on the distal end of the bulb (Fig. 
<figureCitation id="BF121FFF52FD891FDED2484F93B93BB4" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">26B</figureCitation>
) (vs. the absence of a three-branched laminar apophysis in congeners), a spheroid bulb (vs. the bulb of congeners have other shapes), tibia with a retrolateral apophysis bearing a spine that is three times shorter than the apophysis, the entire apophysis, including the spine, is two times longer than the tegulum (Fig. 
<figureCitation id="9904327D520C923BCC7021BF1870CF40" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">26D</figureCitation>
) (vs. the absence of such a combination of a tibia apophysis and spine in congeners); the female can be differentiated from congeners by the pair of stalked spermathecae with a triangular distal part (Fig. 
<figureCitation id="7A08F89647662875BA5A540C4739AC28" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">25A</figureCitation>
).
</paragraph>
<caption id="93283048FFD7A374F3B84BC3609C5424" doi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
<paragraph id="4E025842F22F5C6B6D6BAA5959B78CA7" pageId="0" pageNumber="1">
<emphasis id="C991FBFE6376F5B09B9CA129C2DC981A" bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
<taxonomicName id="269F256FC1799014653CDA2B5E9DCBCB" authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera hponensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hponensis">
<emphasis id="F269A6243F08E9FB44669F9E30117600" italics="true" pageId="0" pageNumber="1">Leclercera hponensis</emphasis>
</taxonomicName>
sp. nov., male holotype and female paratype. 
<emphasis id="A48383C1CEE3AD4ABDAB120AF157BE9E" bold="true" pageId="0" pageNumber="1">A</emphasis>
Endogyne, dorsal view 
<emphasis id="42912365E6EFA9A35B8C1A8911294E51" bold="true" pageId="0" pageNumber="1">B</emphasis>
female epigastric area, ventral view 
<emphasis id="C7163F83A5DCBC892D4D8D1FF31DF4A5" bold="true" pageId="0" pageNumber="1">C</emphasis>
male habitus, dorsal view 
<emphasis id="047F300AFFC2E7922486D49AEA8CB322" bold="true" pageId="0" pageNumber="1">D</emphasis>
female habitus, dorsal view 
<emphasis id="DA2005FA37C71E24187E4C7037C9884B" bold="true" pageId="0" pageNumber="1">E</emphasis>
female habitus, ventral view. Abbreviation: SP = spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7A0E525515F36FD7B8226EB6C038F363" pageId="0" pageNumber="1" type="description">
<paragraph id="70007BB758E92C6AB948AB7DFFDD4828" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="B9941BB182F5042DF24A92E5917FD298" pageId="0" pageNumber="1">
<emphasis id="23B51F53D729D418E245216573A1FB19" bold="true" pageId="0" pageNumber="1">Male</emphasis>
(Holotype). Total length 2.30; carapace 0.80 long, 0.88 wide; abdomen 1.50 long, 0.75 wide. Carapace round and brown, with dark brown longitudinal median band (Fig. 
<figureCitation id="2BF7B07B788FC102AB85ABFDE20F9385" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">25C</figureCitation>
). Chelicerae brown (Fig. 
<figureCitation id="D8233FD4229A81A2AAE8206E34BD3607" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">56F</figureCitation>
). Clypeus light brown. Endites dark brown, light brown basally. Labium dark brown. Sternum light brown. Abdomen elongated, antero-dorsally with dark brown spots, posterior with dark brown stripes medially, antero-ventrally with dark brown elliptical patch, posterior with indistinct dark brown pattern. Legs uniformly brown; measurements: I 8.97 (2.53, 0.31, 2.81, 2.19, 1.13), II 7.10 (1.94, 0.31, 2.19, 1.72, 0.94), III 5.25 (1.44, 0.31, 1.47, 1.25, 0.78), IV 7.97 (2.06, 0.31, 2.13, 2.38, 1.09). Palp (Fig. 
<figureCitation id="515261ADDF563B5AA63F0B99B61726BA" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">26A-D</figureCitation>
): femur slender, four times longer than patella; patella not swollen; tibia 1.5 times shorter than femur, with retrolateral apophysis anteriorly bearing a spine, two times longer than tegulum, spine three times shorter than apophysis (Fig. 
<figureCitation id="934A91F0751D21F202FE1EABCC2D797A" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">26D</figureCitation>
); cymbium three times shorter than femur; bulb brown, spheroid, with embolus and laminar apophysis arising distally; three-branched, laminar apophyses almost equal in length but shorter than embolus; embolus laminar, longer than all other laminar apophyses of its own (Fig. 
<figureCitation id="2C045DFE02B31EB795BEB9B3213C5565" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Leclercera hponensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1">26B</figureCitation>
).
</paragraph>
<paragraph id="DA79E538661A1FB7625E58C8B1800FE9" pageId="0" pageNumber="1">
<emphasis id="D73B1B5F9789540313CE9EE1C5668168" bold="true" pageId="0" pageNumber="1">Female</emphasis>
(Paratype). General features and coloration similar to those of male (Fig. 
<figureCitation id="B467BA2E1749862E09CF1FFBE86E65D1" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">25D, E</figureCitation>
). Measurements: total length 2.40; carapace 0.84 long, 0.94 wide; abdomen 1.56 long, 1.19 wide. Leg measurements: I 7.75 (2.06, 0.31, 2.44, 1.88, 1.06), II 6.47 (1.72, 0.31, 1.94, 1.56, 0.94), III 4.53 (1.25, 0.31, 1.13, 1.09, 0.75), IV 6.44 (1.72, 0.31, 1.88, 1.56, 0.97). Epigastric area (Fig. 
<figureCitation id="0DEC59D6EA68FA7F9A26FA39DE66D3AF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">25B</figureCitation>
): dark brown patch delimiting a horizontal light brown slit. Endogyne (Fig. 
<figureCitation id="B64FBA2E678DB2798A25C41671C8D490" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Leclercera hponensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure25" httpUri="https://binary.pensoft.net/fig/380967" pageId="0" pageNumber="1">25A</figureCitation>
): a pair of stalked spermathecae with anterior stalks and posteriorly a triangular distal part.
</paragraph>
</subSubSection>
<subSubSection id="A747E0C94250BBBCD47653D6B765F1CD" pageId="0" pageNumber="1" type="distribution">
<paragraph id="C0F7524DD8E83C6D868C6AC30CAF1903" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="4D9357A7C512438AE0325CFFA73193DD" pageId="0" pageNumber="1">
Known only from the type locality (Fig. 
<figureCitation id="52AEF2101681FFF2E70D64DB052DEEEA" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">58</figureCitation>
).
</paragraph>
<caption id="933BAF0D47BD6FE0F042D2E90D1C8A0A" doi="10.3897/zookeys.913.48650.figure26" httpUri="https://binary.pensoft.net/fig/380968" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
<paragraph id="F48DDAF06A3DE4FC4F116EFEF05693A8" pageId="0" pageNumber="1">
<emphasis id="D42F2EABC9A5B5600139998622124E0B" bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
<taxonomicName id="ECEEAF38D274A64A2C2471D2D7186FA5" authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera hponensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hponensis">
<emphasis id="7EC64E6D9D6E9B5CF543A9CF5C5AE620" italics="true" pageId="0" pageNumber="1">Leclercera hponensis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="AF580128362EF90DD25B10292B237EAD" bold="true" pageId="0" pageNumber="1">A</emphasis>
Palp, ventral view 
<emphasis id="F9BEC3CDA05F1A70B7250341A085ECAA" bold="true" pageId="0" pageNumber="1">B</emphasis>
bulb, ventral view 
<emphasis id="E9F1C8C971B03EC42D252EC945445C4F" bold="true" pageId="0" pageNumber="1">C</emphasis>
palp, prolateral view 
<emphasis id="27BF3F4FE4871E7540BBE731FE22F3E5" bold="true" pageId="0" pageNumber="1">D</emphasis>
palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>