<documentid="1354C86644E9282E98F13D53581B4CDE"ID-CLB-Dataset="8134"ID-DOI="10.11646/zootaxa.4927.1.2"ID-GBIF-Dataset="f8e774a8-36c4-482c-8d53-adc4612ea281"ID-ISSN="1175-5326"ID-Zenodo-Dep="4533849"ID-ZooBank="E4C28E67-3072-4023-BD47-C2FD9CECAAE5"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="admin"IM.taxonomicNames_approvedBy="admin"checkinTime="1613034223463"checkinUser="plazi"docAuthor="Lyu, Zhi-Tong, Zeng, Zhao-Chi, Wang, Jian, Liu, Zu-Yao, Huang, Ya-Qiong, Li, Wen-Zhou & Wang, Ying-Yong"docDate="2021"docId="03B087AFFFB0FFAC628EFA3DFDE3D77E"docLanguage="en"docName="zootaxa.4927.1.2.pdf"docOrigin="Zootaxa 4927 (1)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Panophrys daiyunensis Lyu, Wang & Wang 2021, sp. nov."docType="treatment"docVersion="10"lastPageNumber="23"masterDocId="FF89FFD7FFBCFFA26219FFBFFF8DD303"masterDocTitle="Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus"masterLastPageNumber="40"masterPageNumber="9"pageNumber="21"updateTime="1720383473109"updateUser="admin"zenodo-license-document="CLOSED">
<mods:titleid="84BEE5D3B1D6C382139B203A0F18B42A">Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus</mods:title>
<mods:affiliationid="BDA76EF2342F0653A467B8D1CE49F253">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859</mods:affiliation>
<mods:affiliationid="BE292259B1CF1B22D90FE43E669C0603">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399</mods:affiliation>
<mods:affiliationid="538B38E2EFF4216504963452D85553E2">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & wangj 397 @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 4249 - 7767</mods:affiliation>
<mods:affiliationid="167A2E3FF0CBF709F2627D47B6F07C29">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Institute of Ecology and Evolution, University of Bern, Bern 3012, Switzerland.</mods:affiliation>
<mods:affiliationid="A82C3E09515C0F1C7E119CD7F81088A8">Fujian Daiyun Mountain National Nature Reserve, Quanzhou 362500, PR China & Fujian Wildlife Conservation Center, Fujian Provincial Department of Forestry, Fuzhou 350003, PR China</mods:affiliation>
<mods:affiliationid="7A5738D3C3838075F0EE9C5BE40D94D7">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
<taxonomicNameid="4C194D3AFFB0FFAE628EFA3DFD3DD69E"authority="Lyu, Wang & Wang"authorityName="Lyu, Wang & Wang"authorityYear="2021"box="[151,688,1410,1437]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="12"pageNumber="21"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB0FFAE6335FA78FE13D6DC"ID-CoL="5N6Z"authorityName="Kuhl & Van Hasselt"authorityYear="1822"box="[300,414,1479,1503]"class="Amphibia"family="Megophryidae"genus="Megophrys"kingdom="Animalia"order="Anura"pageId="12"pageNumber="21"phylum="Chordata"rank="genus">
<bibRefCitationid="EF884B48FFB0FFAE63F5FA78FD0ED6DC"author="Liu, Z. Y. & Chen, G. L. & Zhu, T. Q. & Zeng, Z. C. & Lyu, Z. T. & Wang, J. & Messenger, K. & Greenberg, A. J. & Guo, Z. X. & Yang, Z. H. & Shi, S. H. & Wang, Y. Y."box="[492,643,1479,1503]"pageId="12"pageNumber="21"pagination="723 - 731"refId="ref18211"refString="Liu, Z. Y., Chen, G. L., Zhu, T. Q., Zeng, Z. C., Lyu, Z. T., Wang, J., Messenger, K., Greenberg, A. J., Guo, Z. X., Yang, Z. H., Shi, S. H. & Wang, Y. Y. (2018) Prevalence of cryptic species in morphologically uniform taxa-Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution, 127, 723 - 731. https: // doi. org / 10.1016 / j. ympev. 2018.06.020"type="journal article"year="2018">
<figureCitationid="13222A3CFFB0FFAE63D8F9B2FD8FD524"box="[449,514,1549,1575]"captionStart="FIGURE 3"captionStartId="15.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-20@15.[151,1436,181,1842]"captionTargetPageId="15"captionText="FIGURE 3. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a001733 of Panophrys daiyunensis sp. nov. in life. (E)–(F) dorsolateral and ventral views of adult female paratype SYS a006000."figureDoi="http://doi.org/10.5281/zenodo.4533856"httpUri="https://zenodo.org/record/4533856/files/figure.png"pageId="12"pageNumber="21">Fig. 3</figureCitation>
<collectingDateid="EFE3E991FFB0FFAE660DF9B2FB26D524"box="[1044,1195,1549,1576]"pageId="12"pageNumber="21"value="2012-05-22">22 May 2012</collectingDate>
<specimenCodeid="DBBF9EC2FFB0FFAE60A3F9C6FCCFD597"box="[698,834,1657,1684]"collectionCode="CIB"country="China"httpUri="http://grbio.org/cool/vxkr-andy"name="Chengdu Institute of Biology"pageId="12"pageNumber="21">CIB 116075</specimenCode>
, SYS a001731–1732), collected at the same time from the same locality as the holotype
<collectingDateid="EFE3E991FFB0FFAE66D1F922FAD3D5B4"box="[1224,1374,1693,1720]"pageId="12"pageNumber="21"value="2017-06-26">26 June 2017</collectingDate>
</date>
from the same locality as the holotype
</materialsCitation>
;
<materialsCitationid="3B713CE4FFB0FFAE600FF97DFE31D427"ID-GBIF-Occurrence="3038228309"collectingDate="2017-06-26"collectorName="Zhi-Tong Lyu & Ying-Yong Wang & Ya-Qiong Huang"country="China"elevation="1200"latitude="25.7101"location="Daiyun Mountain Nature Reserve"longLatPrecision="7"longitude="118.12"pageId="12"pageNumber="21"specimenCount="1"stateProvince="Fujian"typeStatus="paratype">
two adult females (SYS a006000, 6003), collected by
<collectingDateid="EFE3E991FFB0FFAE63E5F95AFD1CD5FC"box="[508,657,1765,1792]"pageId="12"pageNumber="21"value="2017-06-26">26 June 2017</collectingDate>
<taxonomicNameid="4C194D3AFFB0FFAE602DF892FD35D444"authorityName="Lyu, Wang & Wang"authorityYear="2021"box="[564,696,1837,1863]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="12"pageNumber="21"phylum="Chordata"rank="species"species="daiyunensis">
(n = 3) in adult females; (2) tympanum edge distinct, upper margin in contact with supratympanic fold, TD/ED 0.45–0.53; (3) vomerine teeth present; (4) margin of tongue not notched; (5) heels overlapping or meeting; (6) tibio-tarsal articulation reaching just posterior to eye; (7) TIB/SVL 0.36–0.42, FTL/SVL 0.51–0.60; (8) narrow lateral fringes on fingers present, one subarticular tubercle present at the base of each finger, relative finger lengths I = II <IV <III; (9) rudimentary web and narrow lateral fringes on toes present, one subarticular tubercle present at the base of each toe; (10) dorsal body’s skin texture rough, with densely-distributed granules and scattered tubercles, “X” shaped skin ridge on center of dorsum and dorsolateral skin ridges present; (11) a small horn-like tubercle present at edge of upper eyelid; (12) supratympanic fold distinct; (13) dorsal surfaces yellowish brown or reddish brown, a dark triangular marking with light edge between eyes, a dark “X” or “)(”-shaped marking, with light edge, present on center of dorsum; (14) rounded, densely-distributed tubercles present on ventrum (chest and belly); (15) single subgular vocal sac present in males; (16) nuptial pads and/or nuptial spines absent in breeding males.
<taxonomicNameid="4C194D3AFFB1FFAF6079FE40FCE9D11A"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[608,868,511,537]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF66B3FE40FAAED11A"authorityName="Rao & Yang"authorityYear="1997"box="[1194,1315,511,537]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="genus">
<tableCitationid="C69B0302FFB1FFAF633BFD9CFEFAD13E"box="[290,375,547,573]"captionStart="TABLE 3"captionStartId="8.[151,239,468,494]"captionTargetPageId="8"captionText="TABLE 3. Diagnostic characters separating all 51 species of the genus Panophrys."httpUri="http://table.plazi.org/id/DF666631FFB4FFAA628EFE6BFB81D2ED"pageId="13"pageNumber="22"tableUuid="DF666631FFB4FFAA628EFE6BFB81D2ED">Table 3</tableCitation>
<taxonomicNameid="4C194D3AFFB1FFAF62EFFDF8FEE2D162"authorityName="Rao & Yang"authorityYear="1997"box="[246,367,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="genus">
<taxonomicNameid="4C194D3AFFB1FFAF61CAFDF8FB2ED162"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="2005"box="[979,1187,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="huangshanensis">
<taxonomicNameid="4C194D3AFFB1FFAF677FFDF8FF73D186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu & Jiang"baseAuthorityYear="2017"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="lishuiensis">
<taxonomicNameid="4C194D3AFFB1FFAF6312FDD4FE2AD186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Messenger & Dahn"baseAuthorityYear="2019"box="[267,423,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="ombrophila">
<taxonomicNameid="4C194D3AFFB1FFAF63FBFDD4FDF6D187"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Wu, Peng, Shi, Lu & Wu"baseAuthorityYear="2020"box="[482,635,619,644]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="xianjuensis">
<taxonomicNameid="4C194D3AFFB1FFAF6091FDD4FC06D186"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[648,907,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF6036FD68FCB9D1F2"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[559,820,727,753]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF6634FD68FB71D1F2"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="2005"box="[1069,1276,727,753]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="huangshanensis">
in female), vomerine teeth present (vs absent), tongue not notched (vs notched), narrow lateral fringes and rudimentary web on toes present (vs both absent), and round light patches on the shoulder absent (vs present).
<taxonomicNameid="4C194D3AFFB1FFAF6092FCFCFC1DD05E"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[651,912,835,861]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
by its vomerine teeth present (vs absent), tongue not notched (vs notched), heels overlapping or meeting (vs not meeting), and rudimentary web on toes present (vs absent).
<taxonomicNameid="4C194D3AFFB1FFAF60DDFC34FC44D0A6"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[708,969,907,933]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF66A7FC34FADDD0A6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu & Jiang"baseAuthorityYear="2017"box="[1214,1360,907,933]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="lishuiensis">
<taxonomicNameid="4C194D3AFFB1FFAF662AFC6CFAB4D0EE"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1075,1337,979,1005]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF633AFC48FE4DD712"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Messenger & Dahn"baseAuthorityYear="2019"box="[291,448,1015,1041]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="ombrophila">
by its vomerine teeth present (vs absent), heels overlapping or meeting (vs not meeting), and narrow lateral fringes and rudimentary web on toes present (vs both absent).
<taxonomicNameid="4C194D3AFFB1FFAF662AFBA4FAB4D736"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1075,1337,1051,1077]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF633CFB80FE32D75B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Wu, Peng, Shi, Lu & Wu"baseAuthorityYear="2020"box="[293,447,1087,1112]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="xianjuensis">
<taxonomicNameid="4C194D3AFFB1FFAF673AFB38FE96D7C6"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF62E9FB70FEC2D7EA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che"baseAuthorityYear="2019"box="[240,335,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="angka">
<taxonomicNameid="4C194D3AFFB1FFAF6345FB70FD86D7EA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye, Fei & Xie"baseAuthorityYear="2007"box="[348,523,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="baolongensis">
<taxonomicNameid="4C194D3AFFB1FFAF6001FB70FD5ED7EA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[536,723,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="binchuanensis">
<taxonomicNameid="4C194D3AFFB1FFAF60F8FB70FC0FD7EA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Fei & Ye"baseAuthorityYear="2009"box="[737,898,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="binlingensis">
<taxonomicNameid="4C194D3AFFB1FFAF6189FB70FBB9D7EA"baseAuthorityName="Inger & Romer"baseAuthorityYear="1961"box="[912,1076,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="brachykolos">
<taxonomicNameid="4C194D3AFFB1FFAF6658FB70FB79D7EA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Nguyen, Pham, Nguyen, Luong & Ziegler"baseAuthorityYear="2020"box="[1089,1268,1231,1257]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="caobangensis">
<taxonomicNameid="4C194D3AFFB1FFAF6718FB70FF20D60F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen"baseAuthorityYear="1994"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="caudoprocta">
<taxonomicNameid="4C194D3AFFB1FFAF62A2FB4CFEE8D60E"baseAuthorityName="Rao & Yang"baseAuthorityYear="1997"box="[187,357,1267,1293]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daweimontis">
<taxonomicNameid="4C194D3AFFB1FFAF636BFB4CFDABD60E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[370,550,1267,1293]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="fansipanensis">
<taxonomicNameid="4C194D3AFFB1FFAF602CFB4CFD75D60E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[565,760,1267,1293]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="hoanglienensis">
<taxonomicNameid="4C194D3AFFB1FFAF611FFB4CFC09D60E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu, Lyu, Zeng & Wang"baseAuthorityYear="2017"box="[774,900,1267,1293]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="insularis">
<taxonomicNameid="4C194D3AFFB1FFAF618BFB4CFC62D60F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang & Wu"baseAuthorityYear="2020"box="[914,1007,1267,1292]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="jiangi">
<taxonomicNameid="4C194D3AFFB1FFAF61E4FB4CFB38D60E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="1983"box="[1021,1205,1267,1293]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="jingdongensis">
<taxonomicNameid="4C194D3AFFB1FFAF66DBFB4CFAF7D60F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang"baseAuthorityYear="2012"box="[1218,1402,1267,1292]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="jinggangensis">
<taxonomicNameid="4C194D3AFFB1FFAF6791FB4CFEADD632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Li, Xu, Liu, Jiang, Wei & Wang"baseAuthorityYear="2018"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="leishanensis">
<taxonomicNameid="4C194D3AFFB1FFAF6336FAA8FE22D632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou"baseAuthorityYear="2017"box="[303,431,1303,1329]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="liboensis">
<taxonomicNameid="4C194D3AFFB1FFAF63A7FAA8FE72D632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Yang"baseAuthorityYear="2014"box="[446,511,1303,1329]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="lini">
<taxonomicNameid="4C194D3AFFB1FFAF6016FAA8FD1DD632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang & Zhao"baseAuthorityYear="2020"box="[527,656,1303,1329]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="mirabilis">
<taxonomicNameid="4C194D3AFFB1FFAF6114FAA8FC39D632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng & Wang"baseAuthorityYear="2019"box="[781,948,1303,1329]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="nankunensis">
<taxonomicNameid="4C194D3AFFB1FFAF61DAFAA8FBACD632"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Zhao"baseAuthorityYear="2014"box="[963,1057,1303,1329]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="obesa">
<taxonomicNameid="4C194D3AFFB1FFAF62C2FA84FE0BD656"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Ye & Fei"baseAuthorityYear="2008"box="[219,390,1339,1365]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="sangzhiensis">
<taxonomicNameid="4C194D3AFFB1FFAF638CFA84FDD1D656"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tian & Sun"baseAuthorityYear="1995"box="[405,604,1339,1365]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="shuichengensis">
<taxonomicNameid="4C194D3AFFB1FFAF6072FA84FD57D657"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu & Hu"baseAuthorityYear="1973"box="[619,730,1339,1364]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="spinata">
<taxonomicNameid="4C194D3AFFB1FFAF60F3FA84FC45D656"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen, Mo & Li"baseAuthorityYear="2010"box="[746,968,1339,1365]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="tuberogranulatus">
<taxonomicNameid="4C194D3AFFB1FFAF61CEFA84FB08D657"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[983,1157,1339,1364]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="wugongensis">
<taxonomicNameid="4C194D3AFFB1FFAF668DFA84FAF5D656"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[1172,1400,1339,1365]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="wuliangshanensis">
<taxonomicNameid="4C194D3AFFB1FFAF6791FA84FEA8D67A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="wushanensis">
<taxonomicNameid="4C194D3AFFB1FFAF632AFAE0FE63D67B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[307,494,1375,1400]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="xiangnanensis">
<taxonomicNameid="4C194D3AFFB1FFAF6035FAE0FD60D67B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[556,749,1375,1400]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="yangmingensis">
<taxonomicNameid="4C194D3AFFB1FFAF62DEFA3CFE46D69E"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[199,459,1411,1437]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFB1FFAF6759FA18FE8AD6E7"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFB1FFAF608AFA74FCA3D6E6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[659,814,1483,1509]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFB1FFAF62ACFA50FEE1D50A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[181,364,1519,1545]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="mufumontana">
<taxonomicNameid="4C194D3AFFB1FFAF6114FA50FC35D50A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[781,952,1519,1545]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="nanlingensis">
<taxonomicNameid="4C194D3AFFB1FFAF634EF9ACFDABD52E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Deng, Liu, Wu & Liu"baseAuthorityYear="2019"box="[343,550,1555,1581]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="shunhuangensis">
<taxonomicNameid="4C194D3AFFB1FFAF669AF9ACFB50D52F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Jin"baseAuthorityYear="2014"box="[1155,1245,1555,1580]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="acuta">
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<taxonomicNameid="4C194D3AFFB1FFAF6054F988FC8CD552"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[589,769,1591,1617]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="mufumontana">
<taxonomicNameid="4C194D3AFFB1FFAF6127F988FB81D552"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Deng, Liu, Wu & Liu"baseAuthorityYear="2019"box="[830,1036,1591,1617]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="shunhuangensis">
<taxonomicNameid="4C194D3AFFB1FFAF628EF9E4FF62D576"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Liu"baseAuthorityYear="2014"box="[151,239,1627,1653]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="cheni">
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<taxonomicNameid="4C194D3AFFB1FFAF63BDF9E4FDC0D576"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[420,589,1627,1653]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="nanlingensis">
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<taxonomicNameid="4C194D3AFFB1FFAF61DEF9C0FB92D59A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Liu"baseAuthorityYear="2014"box="[967,1055,1663,1689]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="13"pageNumber="22"phylum="Chordata"rank="species"species="cheni">
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; head width shorter than head length, HDW/HDL 0.97; snout rounded in dorsal view, projecting, sloping posteriorly to mouth in profile, protruding well beyond margin of lower jaw; dorsal surface of head flat; eye small, ED/HDL 0.39; nostril obliquely ovoid; pupil vertical; canthus rostralis well developed, curved above nostril; loreal region sloping; internasal distance larger than interorbital distance; tympanum distinct, small, upper margin in contact with supratympanic fold; choanae large ovoid, situated at base of maxilla; vomerine teeth present; margin of tongue not notched.
</paragraph>
<paragraphid="8BA636B9FFB1FFAC62DEF820FF72D3D6"blockId="13.[151,1437,150,2013]"lastBlockId="14.[151,1437,151,1149]"lastPageId="14"lastPageNumber="23"pageId="13"pageNumber="22">Lower arm length 0.23 of SVL and hand length 0.24 of SVL; relative finger lengths I = II <IV <III; tip of finger rounded, slightly dilated; one subarticular tubercle present at base of each finger; fingers without webs, narrow lateral fringes present on fingers; outer and inner metacarpal tubercles distinct, inner metacarpal tubercle observably enlarged.</paragraph>
<paragraphid="8BA636B9FFB2FFAC62DEFF60FDF2D28A"blockId="14.[151,1437,151,1149]"pageId="14"pageNumber="23">Shank length 0.46 of SVL and foot 0.63 of SVL; tibio-tarsal articulation reaches posterior margin of eye when hindlimb stretched alongside body; heels overlapping when hindlimbs held at right angles to body; relative toe lengths I <II <V <III <IV; tips of toes rounded, slightly dilated; one subarticular tubercle present at base of each toe; rudimentary web and lateral fringes present between toes; tarsal folds absent; inner metatarsal tubercle long, ovoid, and outer metatarsal tubercle absent.</paragraph>
<paragraphid="8BA636B9FFB2FFAC62DEFE2CFE34D162"blockId="14.[151,1437,151,1149]"pageId="14"pageNumber="23">Dorsal body’s skin texture rough with densely-distributed granules and scattered raised tubercles; “X” shaped skin ridge on center of dorsum; dorsolateral skin ridges present; a small horn-like tubercle present at edge of upper eyelid; supratympanic fold distinct, curving from posterior corner of eye, posteroventrally to above insertion of arm. Ventral surface of throat smooth; rounded, densely-distributed tubercles present on ventral chest and belly; raised, densely-distributed tubercles on ventral thigh; pectoral gland large, closer to axilla; single large femoral gland on posterior surface of thigh.</paragraph>
Dorsal surface yellowish brown; a dark incomplete triangular marking with light edge between eyes; a dark “X” shaped marking with light edge on center of dorsum; dark patches on dorsal upper arms and hindlimbs; dark stripes below eyes and at lateral tip of snout; iris reddish brown. Ventral surface dark brown; three dark longitudinal stripes on the throat, middle one distinctly shorter; red spots on the chest and densely-distributed tiny white spots on belly and ventral thigh; palms and soles purplish brown, tips of digits greyish white, metacarpal and metatarsal tubercles orange red; pectoral glands and femoral glands white.
<tableCitationid="C69B0302FFB2FFAC613BFCFCFCF4D05E"box="[802,889,835,861]"captionStart="TABLE 4"captionStartId="14.[151,239,1196,1220]"captionTargetPageId="14"captionText="TABLE 4. Measurements (in mm) of the type series of Panophrys daiyunensis sp. nov., * for the holotype."httpUri="http://table.plazi.org/id/DF666631FFB2FFAC628EFB13FA9CD7C7"pageId="14"pageNumber="23"tableUuid="DF666631FFB2FFAC628EFB13FA9CD7C7">Table 4</tableCitation>
. All specimens were similar in morphology. Females are larger than males. SYS a006000 (
<figureCitationid="13222A3CFFB2FFAC609BFCD8FD78D082"box="[642,757,871,897]"captionStart="FIGURE 3"captionStartId="15.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-20@15.[151,1436,181,1842]"captionTargetPageId="15"captionText="FIGURE 3. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a001733 of Panophrys daiyunensis sp. nov. in life. (E)–(F) dorsolateral and ventral views of adult female paratype SYS a006000."figureDoi="http://doi.org/10.5281/zenodo.4533856"httpUri="https://zenodo.org/record/4533856/files/figure.png"pageId="14"pageNumber="23">Fig. 3E, F</figureCitation>
) has reddish brown dorsal surface with “)(”-shaped marking and gray ventral surface with unclear marking. Without “X” or “)(”-shaped marking on center of dorsum in SYS a006002.
<taxonomicNameid="4C194D3AFFB2FFAC6069FC6CFCF9D0EE"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[624,884,979,1005]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="14"pageNumber="23"phylum="Chordata"rank="species"species="daiyunensis"status="sp. nov.">
. This toad inhabits streams surrounded by moist subtropical secondary evergreen broadleaved forests, and is common from May to June. Males call actively on leaves of bushes or rocks near streams during this period. All females found in June were gravid with oocytes but tadpoles have not been found.
