treatments-xml/data/03/AB/94/03AB947E202CFFD2A26CFF2DFE122978.xml

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<document id="6D0BD185857C08EBE0AF2EB3B3DBD906" ID-DOI="10.18590/euscorpius.2006.vol2006.iss35.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="12781455" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721358364473" checkinUser="felipe" docAuthor="Baptista, Chris, Santiago-Blay, Jorge A., Soleglad, Michael E. &amp; Fet, Victor" docDate="2006" docId="03AB947E202CFFD2A26CFF2DFE122978" docLanguage="en" docName="Euscorpius.35.1-40.pdf" docOrigin="Euscorpius 35 (35)" docSource="https://mds.marshall.edu/euscorpius/vol2006/iss35/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Archaeobuthus estephani Lourenco 2001" docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="FF92EC06202AFFD4A36AFFEAFFE42F44" masterDocTitle="The Cretaceous scorpion genus, Archaeobuthus, revisited (Scorpiones: Archaeobuthidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="5" updateTime="1721380782877" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="36641FBC810C8C75F0AFE6DD7077336A">The Cretaceous scorpion genus, Archaeobuthus, revisited (Scorpiones: Archaeobuthidae)</mods:title>
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<mods:namePart id="3976F26F77A8F659AA9648BE5B7AA7A1">Baptista, Chris</mods:namePart>
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<mods:namePart id="E8004B7F3CC080F258961D7D35B4EE62">Santiago-Blay, Jorge A.</mods:namePart>
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<mods:namePart id="51ADE045386B4807422683A692259CA1">Soleglad, Michael E.</mods:namePart>
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<mods:namePart id="1DA591A987572F950A3572A8B6DFA5A6">Fet, Victor</mods:namePart>
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<treatment id="03AB947E202CFFD2A26CFF2DFE122978" ID-DOI="http://doi.org/10.5281/zenodo.12781368" ID-Zenodo-Dep="12781368" LSID="urn:lsid:plazi:treatment:03AB947E202CFFD2A26CFF2DFE122978" httpUri="http://treatment.plazi.org/id/03AB947E202CFFD2A26CFF2DFE122978" lastPageNumber="5" pageId="6" pageNumber="5">
<subSubSection id="C31876E3202CFFD2A26CFF2DFD382F9A" box="[262,732,199,222]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="8BBD2568202CFFD2A26CFF2DFD382F9A" blockId="6.[262,732,199,253]" box="[262,732,199,222]" pageId="6" pageNumber="5">
<heading id="D0F59204202CFFD2A26CFF2DFD382F9A" box="[262,732,199,222]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="5" reason="4">
<taxonomicName id="4C025EEB202CFFD2A26CFF2DFD382F9A" authority="Lourenco, 2001" authorityName="Lourenco" authorityYear="2001" box="[262,732,199,222]" class="Arachnida" family="Archaeobuthidae" genus="Archaeobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="estephani">
<emphasis id="B976F97A202CFFD2A26CFF2DFD382F9A" bold="true" box="[262,732,199,222]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A26CFF2DFDFA2F9A" bold="true" box="[262,542,199,222]" italics="true" pageId="6" pageNumber="5">Archaeobuthus estephani</emphasis>
Lourenço, 2001
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C31876E3202CFFD2A20AFF0CFD662FB9" box="[352,642,230,253]" pageId="6" pageNumber="5" type="description">
<paragraph id="8BBD2568202CFFD2A20AFF0CFD662FB9" blockId="6.[262,732,199,253]" box="[352,642,230,253]" pageId="6" pageNumber="5">
(
<figureCitation id="133939ED202CFFD2A20DFF0CFE3F2FB9" box="[359,475,230,253]" captionStart-0="Figure 1" captionStart-1="Figures 25" captionStart-2="Figures 69" captionStart-3="Figures 1017" captionStartId-0="4.[192,268,1901,1924]" captionStartId-1="7.[192,278,1484,1507]" captionStartId-2="8.[192,278,1388,1411]" captionStartId-3="13.[192,278,1737,1760]" captionTargetBox-0="[189,1442,189,1860]" captionTargetBox-1="[193,1426,153,1440]" captionTargetBox-2="[184,1447,183,1347]" captionTargetBox-3="[196,1438,123,1711]" captionTargetId-0="figure-27@4.[192,1439,192,1857]" captionTargetId-1="figure-265@7.[193,1438,153,1440]" captionTargetId-2="figure-8@8.[192,1439,192,1339]" captionTargetId-3="figure-135@13.[195,1438,123,1711]" captionTargetPageId-0="4" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="13" captionText-0="Figure 1: Dorsal view of Archaeobuthus estephani." captionText-1="Figures 25: Pedipalp trichobothrial pattern of Archaeobuthus estephani. 2. Dorsoexternal view of chela. 3. Ventral view of chelal palm. 4. Dorsal view of patella. 5. Dorsal view of femur. Compare to Figs. 69." captionText-2="Figures 69: Pedipalp of Archaeobuthus estephani showing trichobothria; compare to Figs. 25. 6. Dorsoexternal view of chela (also see Fig. 36). 7. Ventral view of chelal palm (also see Fig. 37). 8. Dorsal view of patella (also see Fig. 44). 9. Dorsal view of femur (also see Fig. 45)." captionText-3="Figures 1017: Diagrammatic trichobothrial patterns of pedipalpal chela for Archaeobuthus, Pseudochactas, and representatives of the six buthoid groups suggested by Fet et al. (2005). 10. Archaeobuthus estephani (idealized). 11. Pseudochactas ovchinnikovi (after Soleglad &amp; Fet, 2001, in part). 12. Buthus bonito (after Lourenço &amp; Geniez, 2005, in part). 13. Ananteris luciae (after Lourenço, 1984, in part). 14. Isometrus sankeriensis (after Tikader &amp; Bastawade, 1983, in part). 15. Charmus indicus (after Lourenço, 2000b, in part, and Tikader &amp; Bastawade, 1983, in part). 16. Uroplectes occidentalis (after Vachon, 1950, in part). 17. Tityus shiriana (after González-Sponga, 1991, in part). Closed circles depict location of internal trichobothria; horizontal bars depict location of V1V2 and dbdt series." figureDoi-0="http://doi.org/10.5281/zenodo.12781457" figureDoi-1="http://doi.org/10.5281/zenodo.12781463" figureDoi-2="http://doi.org/10.5281/zenodo.12781465" figureDoi-3="http://doi.org/10.5281/zenodo.12781467" httpUri-0="https://zenodo.org/record/12781457/files/figure.png" httpUri-1="https://zenodo.org/record/12781463/files/figure.png" httpUri-2="https://zenodo.org/record/12781465/files/figure.png" httpUri-3="https://zenodo.org/record/12781467/files/figure.png" pageId="6" pageNumber="5">Figs. 110</figureCitation>
,
<figureCitation id="133939ED202CFFD2A28CFF0CFDE72FB9" box="[486,515,230,253]" captionStart="Figures 1825" captionStartId="14.[192,278,1292,1315]" captionTargetBox="[194,1426,196,1251]" captionTargetId="figure-8@14.[192,1437,192,1251]" captionTargetPageId="14" captionText="Figures 1825: Diagrammatic trichobothrial patterns of pedipalpal patella for Archaeobuthus, Pseudochactas, and representatives of the six buthoid groups suggested by Fet et al. (2005). 18. Archaeobuthus estephani (idealized). 19. Pseudochactas ovchinnikovi (after Soleglad &amp; Fet, 2001, in part). 20. Buthus bonito (after Lourenço &amp; Geniez, 2005, in part). 21. Ananteris luciae (after Lourenço, 1984, in part). 22. Isometrus sankeriensis (after Tikader &amp; Bastawade, 1983, in part). 23. Charmus indicus (after Lourenço, 2000b, in part, and Tikader &amp; Bastawade, 1983, in part). 24. Uroplectes occidentalis (after Vachon, 1950, in part). 25. Tityus shiriana (after González-Sponga, 1991, in part). Vertical bars depict location of external trichobothria." figureDoi="http://doi.org/10.5281/zenodo.12781473" httpUri="https://zenodo.org/record/12781473/files/figure.png" pageId="6" pageNumber="5">18</figureCitation>
,
<figureCitation id="133939ED202CFFD2A164FF0CFDCF2FB9" box="[526,555,230,253]" captionStart="Figures 2633" captionStartId="15.[192,278,1214,1237]" captionTargetBox="[196,1423,194,1166]" captionTargetId="figure-9@15.[196,1423,194,1166]" captionTargetPageId="15" captionText="Figures 2633: Diagrammatic trichobothrial patterns of pedipalpal femur for Archaeobuthus, Pseudochactas, and representatives of the six buthoid groups suggested by Fet et al. (2005). 26. Archaeobuthus estephani (idealized). 27. Pseudochactas ovchinnikovi (after Soleglad &amp; Fet, 2001, in part). 28. Buthus bonito (after Lourenço &amp; Geniez, 2005, in part). 29. Ananteris luciae (after Lourenço, 1984, in part). 30. Isometrus sankeriensis (after Tikader &amp; Bastawade, 1983, in part). 31. Charmus indicus (after Lourenço, 2000b, in part). 32. Uroplectes occidentalis (after Vachon, 1950, in part). 33. Tityus shiriana (after González-Sponga, 1991, in part). Vertical bars depict location of internal and external trichobothria; closed circles depict trichobothria d1, d3, and d4, emphasizing the alpha/beta pattern." figureDoi="http://doi.org/10.5281/zenodo.12781475" httpUri="https://zenodo.org/record/12781475/files/figure.png" pageId="6" pageNumber="5">26</figureCitation>
,
<figureCitation id="133939ED202CFFD2A15CFF0CFD9E2FB9" box="[566,634,230,253]" captionStart-0="Figure 36" captionStart-1="Figure 37" captionStart-10="Figure 46" captionStart-11="Figure 47" captionStart-12="Figure 48" captionStart-13="Figure 49" captionStart-14="Figure 50" captionStart-15="Figure 51" captionStart-16="Figure 52" captionStart-17="Figure 53" captionStart-18="Figure 54" captionStart-2="Figure 38" captionStart-3="Figure 39" captionStart-4="Figure 40" captionStart-5="Figure 41" captionStart-6="Figure 42" captionStart-7="Figure 43" captionStart-8="Figure 44" captionStart-9="Figure 45" captionStartId-0="23.[192,268,1863,1886]" captionStartId-1="24.[192,268,1863,1886]" captionStartId-10="33.[192,268,1863,1886]" captionStartId-11="34.[192,268,1863,1886]" captionStartId-12="35.[192,268,1862,1885]" captionStartId-13="36.[192,268,1886,1909]" captionStartId-14="37.[192,268,1862,1885]" captionStartId-15="38.[192,268,1862,1885]" captionStartId-16="39.[192,268,1862,1885]" captionStartId-17="40.[192,268,1857,1880]" captionStartId-18="41.[192,268,1863,1886]" captionStartId-2="25.[192,268,1860,1883]" captionStartId-3="26.[192,268,1863,1886]" captionStartId-4="27.[192,268,1876,1899]" captionStartId-5="28.[192,268,1863,1886]" captionStartId-6="29.[192,268,1862,1885]" captionStartId-7="30.[192,268,1862,1885]" captionStartId-8="31.[192,268,1862,1885]" captionStartId-9="32.[192,268,1862,1885]" captionTargetBox-0="[189,1442,165,1836]" captionTargetBox-1="[187,1445,163,1837]" captionTargetBox-10="[189,1442,165,1836]" captionTargetBox-11="[189,1442,165,1836]" captionTargetBox-12="[189,1443,165,1835]" captionTargetBox-13="[189,1443,189,1859]" captionTargetBox-14="[189,1443,165,1835]" captionTargetBox-15="[189,1443,165,1835]" captionTargetBox-16="[189,1443,165,1835]" captionTargetBox-17="[189,1442,165,1836]" captionTargetBox-18="[189,1442,165,1836]" captionTargetBox-2="[189,1442,165,1836]" captionTargetBox-3="[189,1442,165,1836]" captionTargetBox-4="[189,1443,165,1835]" captionTargetBox-5="[189,1442,165,1836]" captionTargetBox-6="[189,1443,165,1835]" captionTargetBox-7="[189,1443,165,1835]" captionTargetBox-8="[189,1443,165,1835]" captionTargetBox-9="[189,1443,165,1835]" captionTargetId-0="figure-6@23.[192,1439,168,1833]" captionTargetId-1="figure-6@24.[192,1440,168,1832]" captionTargetId-10="figure-6@33.[192,1439,168,1833]" captionTargetId-11="figure-6@34.[192,1439,168,1833]" captionTargetId-12="figure-6@35.[192,1440,168,1832]" captionTargetId-13="figure-6@36.[192,1440,192,1856]" captionTargetId-14="figure-6@37.[192,1440,168,1832]" captionTargetId-15="figure-6@38.[192,1440,168,1832]" captionTargetId-16="figure-6@39.[192,1440,168,1832]" captionTargetId-17="figure-6@40.[192,1439,168,1833]" captionTargetId-18="figure-7@41.[192,1439,168,1833]" captionTargetId-2="figure-7@25.[192,1439,168,1833]" captionTargetId-3="figure-6@26.[192,1439,168,1833]" captionTargetId-4="figure-7@27.[192,1440,168,1832]" captionTargetId-5="figure-6@28.[192,1439,168,1833]" captionTargetId-6="figure-6@29.[192,1440,168,1832]" captionTargetId-7="figure-6@30.[192,1440,168,1832]" captionTargetId-8="figure-6@31.[192,1440,168,1832]" captionTargetId-9="figure-6@32.[192,1440,168,1832]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-10="33" captionTargetPageId-11="34" captionTargetPageId-12="35" captionTargetPageId-13="36" captionTargetPageId-14="37" captionTargetPageId-15="38" captionTargetPageId-16="39" captionTargetPageId-17="40" captionTargetPageId-18="41" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionTargetPageId-5="28" captionTargetPageId-6="29" captionTargetPageId-7="30" captionTargetPageId-8="31" captionTargetPageId-9="32" captionText-0="Figure 36: Archaeobuthus estephani. Dorsoexternal view of pedipalpal chela showing trichobothria Eb1, Eb2, Est, Et1, eb, est, et, and db." captionText-1="Figure 37: Archaeobuthus estephani. Ventral view of pedipalp chelal palm showing trichobothria Eb1, Eb2, V1 (areola only), Est, and Et1." captionText-10="Figure 46: Archaeobuthus estephani. Dorsal view of left pe
)
</paragraph>
</subSubSection>
<subSubSection id="C31876E3202CFFD2A3AAFEF7FE122978" pageId="6" pageNumber="5" type="diagnosis">
<paragraph id="8BBD2568202CFFD2A3AAFEF7FDFF2E70" blockId="6.[192,802,285,461]" box="[192,539,285,308]" pageId="6" pageNumber="5">DIAGNOSIS. As for the family.</paragraph>
<paragraph id="8BBD2568202CFFD2A3AAFED1FEF22E35" blockId="6.[192,802,285,461]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A3AAFED1FEC62E16" box="[192,290,315,338]" italics="true" pageId="6" pageNumber="5">
<typeStatus id="54B99BCA202CFFD2A3AAFED1FEC62E16" box="[192,290,315,338]" pageId="6" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
. Gender and biological age of the specimen unclear.
</paragraph>
<paragraph id="8BBD2568202CFFD2A3AAFE93FDA92EEA" blockId="6.[192,802,285,461]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A3AAFE93FED02ED4" box="[192,308,377,400]" italics="true" pageId="6" pageNumber="5">Repository</emphasis>
. Deposited in the private collection of Antoun Estephan (Frankfurt,
<collectingCountry id="F31565F8202CFFD2A2B0FE7DFDA62EEA" box="[474,578,407,430]" name="Germany" pageId="6" pageNumber="5">Germany</collectingCountry>
).
</paragraph>
<paragraph id="8BBD2568202CFFD2A3AAFE5CFDB82E89" blockId="6.[192,802,285,461]" box="[192,604,438,461]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A3AAFE5CFEE02E89" box="[192,260,438,461]" italics="true" pageId="6" pageNumber="5">Range</emphasis>
. Lebanese amber, 135120 Ma.
</paragraph>
<paragraph id="8BBD2568202CFFD2A3AAFE19FE062C79" blockId="6.[192,802,499,829]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A3AAFE19FEB82D4E" box="[192,348,499,522]" italics="true" pageId="6" pageNumber="5">Measurements</emphasis>
(in mm). Carapace. Anterior width 1.06? (right lateral edge obscured). Width of carapace at lateral eyes 0.89; median length 0.98. Carapace + mesosoma length 2.82. Chelicerae. Width at the base 0.29. Right pedipalp. Femur length 1.06, width 0.27; patella length 1.21, width 0.37; chela length 2.10, width 0.34; chela palm length 0.82, width 0.34; movable finger length 1.28. Legs. Leg III tarsus length (dorsal 0.34, ventral 0.41), tarsus basal width 0.09. Metasoma I length/width (L/W) 0.53/0.44, 1.2. Total length: extrapolated to about 6.0 (after Lourenço, 2001).
</paragraph>
<paragraph id="8BBD2568202CFFD2A3AAFC89FE122978" blockId="6.[192,803,867,1596]" pageId="6" pageNumber="5">
<emphasis id="B976F97A202CFFD2A3AAFC89FEA22C3E" box="[192,326,867,890]" italics="true" pageId="6" pageNumber="5">Description.</emphasis>
Yellowish brown, with reddish brown patches on pedipalp. Weakly to moderately granular. Anterior margin of carapace with a moderate median concavity. Anterior median superciliary carinae vestigial; other carinae obsolete. All furrows moderate to weak. Median ocular tubercle distinctly anterior to the center of carapace. Median eyes separated by almost twice their diameter. With two pairs of lateral eyes, the most distal pair (the third) appears to be missing. Pedipalps: right pedipalp well visible, see below for detailed discussion of trichobothria. Chelicerae: dentition was not fully visible but the ventral distal tine of movable finger is longer than the dorsal distal denticle. Sternum not observable. Pecten not available, only a few scattered teeth observable. Without tibial spurs on legs IIIIV; with pedal spurs; tarsal armature reduced. Tergites weakly granular. Tergites I to VI with three longitudinal carinae, moderate to strong. Tergite VII pentacarinate. Sternites IIII not clearly visible, IVVII rather smooth, with median longitudinal punctations; small oval to circular stigma visible on sternites V and VI. Metasomal segments II to V (and telson) missing. Metasomal segment I with 10 carinae; ventral carinae parallel; dorsolateral with spinoid granules.
</paragraph>
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