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<document id="E4DA30BB58591B90B6D8C0D35197CC01" ID-DOI="10.5281/zenodo.195747" ID-GBIF-Dataset="c371f527-f7c6-456b-ab07-5bef541b10e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="195747" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460232772886" checkinUser="plazi" docAuthor="Saether, Ole A., Andersen, Trond, Pinho, Luiz C. &amp; Mendes, Humberto F." docDate="2010" docId="03BC87900E05FFDAFF44C53FC8D97652" docLanguage="en" docName="zt02497p036.pdf" docOrigin="Zootaxa 2497" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Polypedilum" docType="treatment" docVersion="10" lastPageNumber="28" masterDocId="FF85FFE80E1EFFC1FFD3C738CC1E7200" masterDocTitle="The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n." masterLastPageNumber="36" masterPageNumber="1" pageNumber="28" updateTime="1698239121165" updateUser="plazi">
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<mods:titleInfo id="7E7A25339256BED799D7A0FA0F03E0A2">
<mods:title id="DDE8AEFC53EAFEB779CF193D09B6B800">The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n.</mods:title>
</mods:titleInfo>
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<mods:namePart id="A64EFA5E8B002C46DA268A3A8F7842F4">Saether, Ole A.</mods:namePart>
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<mods:name id="30D5B058147BB9436147D37F1AA02F24" type="personal">
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<mods:roleTerm id="D718D9A7AC86FD9667AF6666FFAF0897">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="616352BD91306417F49162C0528C733F">Andersen, Trond</mods:namePart>
</mods:name>
<mods:name id="F3F83C07A8FAD538A28150A10B3127F7" type="personal">
<mods:role id="88A2D8321023A3BACF281E0821E28717">
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<mods:namePart id="2CF313FAFCB63D79586547D5345EFCB1">Pinho, Luiz C.</mods:namePart>
</mods:name>
<mods:name id="C50F6A62C97E087AAEEA3621F988185E" type="personal">
<mods:role id="394B420623F6EFA70E789B859E188361">
<mods:roleTerm id="F9AB7237BF05635147D6625D35508BEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="813E241766F39CFFFCE4AE738660CEEA">Mendes, Humberto F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A866FFE2F09F88D1B845C522077AB3DB">text</mods:typeOfResource>
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<mods:title id="C30C84435D1FE8DE9AF92BB71B58EDDF">Zootaxa</mods:title>
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<mods:part id="65A3D91AB87707AA9731EE20FC8C8BC8">
<mods:date id="B952146EBE8FB35CA5A8FAA0DC325753">2010</mods:date>
<mods:detail id="B2A70E9BEA40922E4549A350483892E1" type="volume">
<mods:number id="EACCD2EFEDEC16566BF4C613C5A7B35A">2497</mods:number>
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<mods:classification id="C0AD3EE15436100CCD3EC781C295F97A">journal article</mods:classification>
<mods:identifier id="B448808002D437A1A778FE65F4B8B339" type="DOI">10.5281/zenodo.195747</mods:identifier>
<mods:identifier id="D2FDC608727BF484CA55AE996058F161" type="GBIF-Dataset">c371f527-f7c6-456b-ab07-5bef541b10e6</mods:identifier>
<mods:identifier id="44EBC8DFFC50F9BC56C29EC4E5D95788" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2DB46B433EA7F574491790B6929CAE3C" type="Zenodo-Dep">195747</mods:identifier>
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<treatment id="03BC87900E05FFDAFF44C53FC8D97652" ID-DOI="http://doi.org/10.5281/zenodo.5030746" ID-GBIF-Taxon="119387545" ID-Zenodo-Dep="5030746" LSID="urn:lsid:plazi:treatment:03BC87900E05FFDAFF44C53FC8D97652" httpUri="http://treatment.plazi.org/id/03BC87900E05FFDAFF44C53FC8D97652" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="C30F650D0E05FFDAFF44C53FCD337044" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BAA36860E05FFDAFF44C53FCD777021" blockId="27.[151,361,519,580]" box="[151,361,519,545]" pageId="27" pageNumber="28">
<heading id="D0E281EA0E05FFDAFF44C53FCD777021" bold="true" box="[151,361,519,545]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B961EA940E05FFDAFF44C53FCD777021" bold="true" box="[151,361,519,545]" pageId="27" pageNumber="28">
<taxonomicName id="4C154D050E05FFDAFF44C53FCD307021" ID-CoL="63MXN" ID-ENA="54968" box="[151,302,519,545]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E05FFDAFF44C53FCD307021" bold="true" box="[151,302,519,545]" italics="true" pageId="27" pageNumber="28">Polypedilum</emphasis>
</taxonomicName>
spp.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36860E05FFDAFF44C512CD337044" blockId="27.[151,361,519,580]" box="[151,301,554,580]" pageId="27" pageNumber="28">
(
<figureCitation id="132E2A030E05FFDAFF4CC512CD3B7044" box="[159,293,554,580]" captionStart="FIGURES 44 – 45" captionStartId="27.[151,271,1927,1951]" captionTargetBox="[248,1326,1177,1889]" captionTargetId="figure@27.[236,1351,1162,1903]" captionTargetPageId="27" captionText="FIGURES 44 – 45. Polypedilum (Uresipedilum) spp., females. 44 – genitalia, ventral view of paratype of Polypedilum (Uresipedilum) pedatum Townes, 1945, from Missouri. 45 – genitalia, ventral view of paratype of P. (U.) pedatum from Louisiana." httpUri="https://zenodo.org/record/195757/files/figure.png" pageId="27" pageNumber="28">Figs 44–45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F650D0E05FFDAFF44C54DC8D97652" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BAA36860E05FFDAFF44C54DCF7D70FF" blockId="27.[151,1437,629,1106]" pageId="27" pageNumber="28">
<emphasis id="B961EA940E05FFDAFF44C54DCDC5708F" bold="true" box="[151,475,629,655]" pageId="27" pageNumber="28">
Material examined.
<collectingCountry id="F30276160E05FFDAFE44C54DCDC8708F" box="[407,470,629,655]" name="United States of America" pageId="27" pageNumber="28">USA</collectingCountry>
:
</emphasis>
Missouri, Atherton,
<specimenCount id="9D13FD0F0E05FFDAFD04C54DCF21708F" box="[727,831,629,655]" pageId="27" pageNumber="28" type="female">2 female</specimenCount>
<typeStatus id="54AE88240E05FFDAFC94C54DCFA7708F" box="[839,953,629,655]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C154D050E05FFDAFC31C54DC8CA708E" box="[994,1236,629,654]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="pedatum" subSpecies="pedatum">
<emphasis id="B961EA940E05FFDAFC31C54DC8CA708E" box="[994,1236,629,654]" italics="true" pageId="27" pageNumber="28">P. pedatum pedatum</emphasis>
</taxonomicName>
,
<date id="FFAB10460E05FFDAFB37C54DC954708F" box="[1252,1354,629,655]" pageId="27" pageNumber="28" value="1916-05-07">7.v.1916</date>
&amp;
<date id="FFAB10460E05FFDAFAA2C54DCCCA70B4" pageId="27" pageNumber="28" value="1922-05-07">7.v. 1922</date>
, C.F. Adams (Cornell University, Dept. of Entomology,
<typeStatus id="54AE88240E05FFDAFCBFC5A2CFAF70B4" box="[876,945,666,692]" pageId="27" pageNumber="28">Types</typeStatus>
No. 2671.2, 2671.3); Louisiana, opposite Orange, Sabine River Ferry,
<specimenCount id="9D13FD0F0E05FFDAFE29C5F8CE7A70DA" box="[506,612,704,730]" pageId="27" pageNumber="28" type="female">1 female</specimenCount>
<typeStatus id="54AE88240E05FFDAFDBEC5F8CEC870DA" box="[621,726,704,730]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C154D050E05FFDAFCD2C5F8CFE770D9" box="[769,1017,704,729]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="pedatum" subSpecies="pedatum">
<emphasis id="B961EA940E05FFDAFCD2C5F8CFE770D9" box="[769,1017,704,729]" italics="true" pageId="27" pageNumber="28">P. pedatum pedatum</emphasis>
</taxonomicName>
,
<date id="FFAB10460E05FFDAFBD9C5F8C88E70DA" box="[1034,1168,704,730]" pageId="27" pageNumber="28" value="1917-06-20">20.vi.1917</date>
, Cornell Biol. Exped. (Cornell University, Dept. of Entomology,
<typeStatus id="54AE88240E05FFDAFD5EC5DDCED970FF" box="[653,711,741,767]" pageId="27" pageNumber="28">Type</typeStatus>
No. 2671.1).
</paragraph>
<paragraph id="8BAA36860E05FFDAFF15C432C8D97652" blockId="27.[151,1437,629,1106]" pageId="27" pageNumber="28">
Three female
<typeStatus id="54AE88240E05FFDAFEBBC432CDC67124" box="[360,472,778,804]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
from Missouri and Louisiana although belonging to
<taxonomicName id="4C154D050E05FFDAFBECC432C8CD7123" box="[1087,1235,778,803]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E05FFDAFBECC432C8CD7123" box="[1087,1235,778,803]" italics="true" pageId="27" pageNumber="28">Polypedilum</emphasis>
</taxonomicName>
cannot belong to
<taxonomicName id="4C154D050E05FFDAFF44C408CD027149" box="[151,284,816,841]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pedatum">
<emphasis id="B961EA940E05FFDAFF44C408CD027149" box="[151,284,816,841]" italics="true" pageId="27" pageNumber="28">P. pedatum</emphasis>
</taxonomicName>
as they have a quite different ventrolateral lobe of gonapophysis VIII. They have R4+5 strongly curved at apex and ending at wing tip and M is bare. Thus according to the description by Townes they should have been listed under
<taxonomicName id="4C154D050E05FFDAFE77C446CE897197" box="[420,663,893,919]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="pedatum" subSpecies="excelsius">
<emphasis id="B961EA940E05FFDAFE77C446CDAD7197" box="[420,435,894,919]" italics="true" pageId="27" pageNumber="28">P</emphasis>
.
<emphasis id="B961EA940E05FFDAFE12C446CE897197" box="[449,663,894,919]" italics="true" pageId="27" pageNumber="28">pedatum excelsius</emphasis>
</taxonomicName>
and not under
<taxonomicName id="4C154D050E05FFDAFC95C446C8277197" box="[838,1081,893,919]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="pedatum" subSpecies="pedatum">
<emphasis id="B961EA940E05FFDAFC95C446CF4A7197" box="[838,852,894,919]" italics="true" pageId="27" pageNumber="28">P</emphasis>
.
<emphasis id="B961EA940E05FFDAFCB7C446C8277197" box="[868,1081,894,919]" italics="true" pageId="27" pageNumber="28">pedatum pedatum</emphasis>
</taxonomicName>
. However, all three specimens have a spine on the fore tibial scale and thus do not belong to
<taxonomicName id="4C154D050E05FFDAFC52C49BC81071BC" box="[897,1038,930,956]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="excelsius">
<emphasis id="B961EA940E05FFDAFC52C49BCF8E71BC" box="[897,912,931,956]" italics="true" pageId="27" pageNumber="28">P</emphasis>
.
<emphasis id="B961EA940E05FFDAFC4CC49BC81071BC" box="[927,1038,931,956]" italics="true" pageId="27" pageNumber="28">excelsius</emphasis>
</taxonomicName>
. The three specimens differ from each other both in the genitalia and in the front tibial scale and apparently consist of at least two different species. The genitalia of both are illustrated in
<figureCitation id="132E2A030E05FFDAFD3EC4D5CFB87607" box="[749,934,1005,1031]" captionStart="FIGURES 44 – 45" captionStartId="27.[151,271,1927,1951]" captionTargetBox="[248,1326,1177,1889]" captionTargetId="figure@27.[236,1351,1162,1903]" captionTargetPageId="27" captionText="FIGURES 44 – 45. Polypedilum (Uresipedilum) spp., females. 44 – genitalia, ventral view of paratype of Polypedilum (Uresipedilum) pedatum Townes, 1945, from Missouri. 45 – genitalia, ventral view of paratype of P. (U.) pedatum from Louisiana." httpUri="https://zenodo.org/record/195757/files/figure.png" pageId="27" pageNumber="28">Figures 44–45</figureCitation>
. Both have the highly unusual, rounded ventrolateral lobe with weak or no microtrichia as in
<taxonomicName id="4C154D050E05FFDAFCC4C32BC84F762C" box="[791,1105,1043,1068]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="marcondesi" subGenus="Probolum">
<emphasis id="B961EA940E05FFDAFCC4C32BC84F762C" box="[791,1105,1043,1068]" italics="true" pageId="27" pageNumber="28">P. (Probolum) marcondesi</emphasis>
</taxonomicName>
and not the brush-like lobe typical for other
<taxonomicName id="4C154D050E05FFDAFE88C300CDF17651" box="[347,495,1080,1105]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E05FFDAFE88C300CDF17651" box="[347,495,1080,1105]" italics="true" pageId="27" pageNumber="28">Polypedilum</emphasis>
</taxonomicName>
. They thus could very well belong to the subgenus
<emphasis id="B961EA940E05FFDAFB9FC300C8DE7651" box="[1100,1216,1080,1105]" italics="true" pageId="27" pageNumber="28">Probolum</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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