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681 lines
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<document id="F86CC708034E01C027F433B67A1C63A7" ID-CLB-Dataset="22389" ID-DOI="10.5852/ejt.2015.140" ID-GBIF-Dataset="6c9cf4da-a611-4100-b63f-95f094d99bed" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785520" ID-ZooBank="A44ED7CD-AEC4-46F8-8361-796DD49D6CF6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622406715" checkinUser="carolina" docAuthor="Ng, Peter K. L. & Bouchet, Philippe" docDate="2015" docId="BF12EE17B276FFAEFD6D956DFE0FFEBD" docLanguage="en" docName="ejt-2015-140.pdf.imf" docOrigin="European Journal of Taxonomy 140" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Actaea grimaldii Bouchet & Isyeb 2015, sp. nov." docType="treatment" docUuid="A736C0FE-D2B4-4256-AB85-4370386B0231" docUuidSource="ZooBank" docVersion="7" lastPageNumber="11" masterDocId="432B966FB274FFA4FFE99466FFA6FFC9" masterDocTitle="Actaea grimaldii, a new species of reef crab from Papua New Guinea (Crustacea, Brachyura, Xanthidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1698838447334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="015627FC63BD0604DBAF59DA71291E3D">Actaea grimaldii, a new species of reef crab from Papua New Guinea (Crustacea, Brachyura, Xanthidae)</mods:title>
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<mods:namePart id="97353B26A04E04FB9E4C77E1A693A2F3">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="EF43251DA13D2EAA441EE351887713C6">Bouchet, Philippe</mods:namePart>
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<mods:typeOfResource id="3180F37672DAB422459A9723236F9051">text</mods:typeOfResource>
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<mods:title id="8EA7C25C3B9CB3A7F8A4D32F0B78C5DD">European Journal of Taxonomy</mods:title>
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<mods:part id="75E18898C9198A424CF14C3BD8AE7685">
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<mods:date id="2A72BCD672695622CB2FCF1ED1B2754D">2015</mods:date>
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<mods:number id="851DC40673A00D9C0D38A722B743A46A">2015-09-29</mods:number>
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<mods:number id="0320CC878887D172481884A926C16B7D">140</mods:number>
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<treatment id="BF12EE17B276FFAEFD6D956DFE0FFEBD" ID-DOI="http://doi.org/10.5281/zenodo.3794589" ID-GBIF-Taxon="163944005" ID-Zenodo-Dep="3794589" LSID="urn:lsid:zoobank.org:act:A736C0FE-D2B4-4256-AB85-4370386B0231" httpUri="http://treatment.plazi.org/id/BF12EE17B276FFAEFD6D956DFE0FFEBD" lastPageId="10" lastPageNumber="11" pageId="2" pageNumber="3">
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<subSubSection id="7FA10C8AB276FFA6FD6D956DFC16FEEF" box="[644,944,267,294]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="37045F01B276FFA6FD6D956DFC16FEEF" blockId="2.[644,944,267,294]" box="[644,944,267,294]" pageId="2" pageNumber="3">
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<heading id="6C4CE86DB276FFA6FD6D956DFC16FEEF" box="[644,944,267,294]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="F0BB2482B276FFA6FD6D956DFCE8FEEF" ID-CoL="9S2S" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[644,846,267,294]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis id="05CF8313B276FFA6FD6D956DFCE8FEEF" bold="true" box="[644,846,267,294]" italics="true" pageId="2" pageNumber="3">Actaea grimaldii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1EFC3E68B276FFA6FCBC956AFC16FEEF" box="[853,944,268,294]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7FA10C8AB276FFA6FE9A9557FCEBFEBB" pageId="2" pageNumber="3" type="description">
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<paragraph id="37045F01B276FFA6FE9A9557FB66FE85" blockId="2.[371,1216,305,370]" box="[371,1216,305,332]" pageId="2" pageNumber="3">
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<uri id="432A5303B276FFA6FE9A9557FB66FE85" box="[371,1216,305,332]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="431D65D4B276FFA6FD799557FB66FE85" box="[656,1216,305,332]" pageId="2" pageNumber="3">A736C0FE-D2B4-4256-AB85-4370386B0231</uuid>
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</uri>
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</paragraph>
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<paragraph id="37045F01B276FFA6FD0E9531FCEBFEBB" blockId="2.[371,1216,305,370]" box="[743,845,343,370]" pageId="2" pageNumber="3">
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<figureCitation id="AF804384B276FFA6FD0E9531FCEBFEBB" box="[743,845,343,370]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="3.[189,232,1860,1886]" captionStartId-1="5.[189,232,1930,1956]" captionStartId-2="6.[189,232,1930,1956]" captionStartId-3="7.[189,232,1930,1956]" captionStartId-4="8.[189,232,1895,1921]" captionTargetBox-0="[189,1399,399,1819]" captionTargetBox-1="[189,1399,296,1890]" captionTargetBox-2="[205,1383,265,1890]" captionTargetBox-3="[189,1399,436,1890]" captionTargetBox-4="[213,1389,376,1821]" captionTargetId-0="figure@3.[189,1399,399,1819]" captionTargetId-1="figure@5.[189,1399,296,1890]" captionTargetId-2="figure@6.[205,1383,265,1890]" captionTargetId-3="figure@7.[189,1399,436,1890]" captionTargetId-4="figure@8.[189,1398,360,1854]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." captionText-1="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." captionText-2="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." captionText-3="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." captionText-4="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3785522" figureDoi-1="http://doi.org/10.5281/zenodo.3785524" figureDoi-2="http://doi.org/10.5281/zenodo.3785526" figureDoi-3="http://doi.org/10.5281/zenodo.3785528" figureDoi-4="http://doi.org/10.5281/zenodo.3785530" httpUri-0="https://zenodo.org/record/3785522/files/figure.png" httpUri-1="https://zenodo.org/record/3785524/files/figure.png" httpUri-2="https://zenodo.org/record/3785526/files/figure.png" httpUri-3="https://zenodo.org/record/3785528/files/figure.png" httpUri-4="https://zenodo.org/record/3785530/files/figure.png" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7FA10C8AB276FFA6FF5495FDFD2CFDC9" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="37045F01B276FFA6FF5495FDFB4BFE7C" blockId="2.[189,1398,411,513]" box="[189,1261,411,437]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="17AB782FB276FFA6FF5495FDFB4BFE7C" box="[189,1261,411,437]" pageId="2" pageNumber="3">
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<taxonomicName id="F0BB2482B276FFA6FF5495FDFDBEFE7C" ID-CoL="9S5L" authorityName="Henderson" authorityYear="1893" box="[189,536,411,437]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="variety" species="peronii" variety="squamosa">
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<emphasis id="05CF8313B276FFA6FF5495FDFECCFE7C" box="[189,362,411,437]" italics="true" pageId="2" pageNumber="3">Actaea peronii</emphasis>
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var.
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<emphasis id="05CF8313B276FFA6FE4A95FAFDBEFE7C" box="[419,536,412,437]" italics="true" pageId="2" pageNumber="3">squamosa</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="B61A7910B276FFA6FDDF95FDFCA6FE7C" author="Calman W. T." box="[566,768,411,437]" page="10" pageId="2" pageNumber="3" year="1900">
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<bibRefCitation id="532A22F0B276FFA6FDDF95FDFCA6FE7C" author="Calman W. T." box="[566,768,411,437]" pageId="2" pageNumber="3" pagination="1 - 50" refId="ref6232" refString="Calman W. T. 1900. On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Zoology, series 2 8 (1): 1 - 50. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman 1900: 10</bibRefCitation>
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</treatmentCitation>
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(not
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<taxonomicName id="F0BB2482B276FFA6FCD795FDFB46FE7C" ID-CoL="9S5L" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[830,1248,411,437]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="squamosa">
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<emphasis id="05CF8313B276FFA6FCD795FDFBAAFE7C" box="[830,1036,411,437]" italics="true" pageId="2" pageNumber="3">Actaea squamosa</emphasis>
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Henderson, 1893
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="37045F01B276FFA6FF5495A7FD2CFDC9" blockId="2.[189,1398,411,513]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="17AB782FB276FFA6FF5495A7FD2CFDC9" pageId="2" pageNumber="3">
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<taxonomicName id="F0BB2482B276FFA6FF5495A7FE0BFE13" ID-CoL="9S5J" authorityName="Borradaile" authorityYear="1902" box="[189,429,449,475]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spinosissima">
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<emphasis id="05CF8313B276FFA6FF5495A7FE0BFE13" box="[189,429,449,475]" italics="true" pageId="2" pageNumber="3">Actaea spinosissima</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="B61A7910B276FFA6FE2595A7FD33FE12" author="Odhner T." box="[460,661,448,475]" page="59" pageId="2" pageNumber="3" year="1925">
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<bibRefCitation id="532A22F0B276FFA6FE2595A7FD33FE12" author="Odhner T." box="[460,661,448,475]" pageId="2" pageNumber="3" pagination="1 - 92" refId="ref7145" refString="Odhner T. 1925. Monographierte Gattungen der Krabbenfamilie Xanthidae. I. Goteborgs Kungliga Vetenskaps-och Vitterhets-Samhalles Handlingar, serie 4 29 (1): 1 - 92." type="journal article" year="1925">Odhner 1925: 59</bibRefCitation>
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</treatmentCitation>
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(part). —
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<treatmentCitation id="B61A7910B276FFA6FCFC95A7FC43FE12" author="Serene R." box="[789,997,448,475]" page="115" pageId="2" pageNumber="3" year="1984">
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<bibRefCitation id="532A22F0B276FFA6FCFC95A7FC43FE12" author="Serene R." box="[789,997,448,475]" pageId="2" pageNumber="3" refId="ref7392" refString="Serene R. 1984. Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Addendum: Crosnier A. Carpiliidae et Menippidae. Faune Tropicale 24, Editions de l'ORSTOM, Paris." type="journal volume" year="1984">Serène 1984: 115</bibRefCitation>
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</treatmentCitation>
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, pl.
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<specimenCount id="21BD9488B276FFA6FBF095A7FBEAFE12" box="[1049,1100,449,475]" pageId="2" pageNumber="3" type="generic">14F.</specimenCount>
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—
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<treatmentCitation id="B61A7910B276FFA6FB9395A7FA99FE12" author="Davie P. J. F." box="[1146,1343,448,475]" page="511" pageId="2" pageNumber="3" year="2002">
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<bibRefCitation id="532A22F0B276FFA6FB9395A7FA99FE12" author="Davie P. J. F." box="[1146,1343,448,475]" pageId="2" pageNumber="3" refId="ref6358" refString="Davie P. J. F. 2002. Crustacea: Malacostraca. Eucarida. Part 2. Decapoda-Anomura, Brachyura. Zoological Catalogue of Australia 19.3 B, CSIRO Publications, Collingwood." type="book" year="2002">Davie 2002: 511</bibRefCitation>
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</treatmentCitation>
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(not
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<taxonomicName id="F0BB2482B276FFA6FF549581FDDBFDC9" ID-CoL="9S5J" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[189,637,486,513]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spinosissima">
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<emphasis id="05CF8313B276FFA6FF549581FE0AFDC9" box="[189,428,487,513]" italics="true" pageId="2" pageNumber="3">Actaea spinosissima</emphasis>
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<bibRefCitation id="532A22F0B276FFA6FE5A9580FDDBFDC9" author="Borradaile L. A." box="[435,637,486,512]" pageId="2" pageNumber="3" pagination="237 - 271" refId="ref6135" refString="Borradaile L. A. 1902. Marine crustaceans III: The Xanthidae and some other crabs. In: Gardiner J. S. (ed.) The Fauna and Geography of the Maldive and Laccadive Archipelagoes, being the Account of the Work carried on and of the Collections made by an Expedition during the Years 1899 and 1900. Vol. 1. Part. 3: 237 - 271. Cambridge University Press, Cambridge. http: // dx. doi. org / 10.5962 / bhl. title. 10215" type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="7FA10C8AB276FFA6FF54964CFD2DFAD7" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="37045F01B276FFA6FF54964CFE90FD8D" blockId="2.[189,310,554,580]" box="[189,310,554,580]" pageId="2" pageNumber="3">
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<heading id="6C4CE86DB276FFA6FF54964CFE90FD8D" bold="true" box="[189,310,554,580]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
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<emphasis id="05CF8313B276FFA6FF54964CFE90FD8D" bold="true" box="[189,310,554,580]" pageId="2" pageNumber="3">Diagnosis</emphasis>
|
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</heading>
|
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</paragraph>
|
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<paragraph id="37045F01B276FFA6FF54963DFD2DFAD7" blockId="2.[189,1399,603,1310]" pageId="2" pageNumber="3">
|
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Anterior surfaces of carapace regions (1M, 2M, 3M, 2L, 5L) covered with numerous short sharp spines and sharp tubercles of varying sizes, 2M and 2L surface gently convex, not distinctly inflated in frontal view, 2M separated by median longitudinal groove anteriorly, 4M lunate, with sharp granules; 1P flattened, divided into many smaller regions by shallow grooves, with distinct transverse groove on anterior third, confluent with series of flattened granules on 3R, 2R and 1R (
|
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|
|
<figureCitation id="AF804384B276FFA6FB929694FB61FCC4" box="[1147,1223,754,781]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
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,
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<figureCitation id="AF804384B276FFA6FB319695FB58FCC4" box="[1240,1278,755,781]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">2A</figureCitation>
|
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,
|
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|
|
<figureCitation id="AF804384B276FFA6FAE59695FA94FCC4" box="[1292,1330,755,781]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="2" pageNumber="3">3A</figureCitation>
|
|||
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,
|
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|
|
<figureCitation id="AF804384B276FFA6FAA89695FACFFCC4" box="[1345,1385,755,781]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="2" pageNumber="3">4A</figureCitation>
|
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|
); frontal margins lined by sharp granules, 1F and 2F with many sharp spines which project anteriorly, hiding frontal margin from dorsal view (
|
|||
|
|
<figureCitation id="AF804384B276FFA6FD779758FD4FFC91" box="[670,745,830,856]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="AF804384B276FFA6FD109758FCA1FC91" box="[761,775,830,856]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
|
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|
A–B, 3A–B, 4A); anterolateral margin with 4 spines (excluding external orbital spine), each with smaller spine or spines basal to it on sub-branchial surface (
|
|||
|
|
<figureCitation id="AF804384B276FFA6FF2C97EFFEB7FC6D" box="[197,273,905,932]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
|||
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,
|
|||
|
|
<figureCitation id="AF804384B276FFA6FEC897ECFEE1FC6D" box="[289,327,906,932]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">2A</figureCitation>
|
|||
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,
|
|||
|
|
<figureCitation id="AF804384B276FFA6FEBD97ECFEDCFC6D" box="[340,378,906,932]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="2" pageNumber="3">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B276FFA6FE6197ECFE16FC6D" box="[392,432,906,932]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="2" pageNumber="3">4A</figureCitation>
|
|||
|
|
); suborbital and subhepatic regions covered with numerous small, sharp granules (
|
|||
|
|
<figureCitation id="AF804384B276FFA6FF2C97C9FE8EFC00" box="[197,296,943,969]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">Figs 2B</figureCitation>
|
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,
|
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<figureCitation id="AF804384B276FFA6FEDF97C9FEFDFC00" box="[310,347,943,969]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="2" pageNumber="3">3B</figureCitation>
|
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); third maxilliped with outer surfaces pitted (
|
|||
|
|
<figureCitation id="AF804384B276FFA6FC9697C9FC46FC03" box="[895,992,943,970]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="2" pageNumber="3">Fig. 5A</figureCitation>
|
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); outer surfaces of chelipeds with numerous sharp spines and sharp tubercles, with small, sharp granules interspersed, carpus ovate, inner angle with several short, slightly curved spines, outer surface of chela with strong spines, inner surface of chela with median spine and several smaller sharp tubercles and granules (
|
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|
<figureCitation id="AF804384B276FFA6FB849046FB1DFBF2" box="[1133,1211,1056,1083]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
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,
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<figureCitation id="AF804384B276FFA6FB259047FB54FBF2" box="[1228,1266,1057,1083]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">2A</figureCitation>
|
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, C–D, 3A, C–D); ambulatory legs short, outer surfaces of merus, carpus and propodus covered with sharp tubercles or granules, dorsal margins of merus, carpus and dactylus with numerous sharp spines, ventral margin of dactylus with subterminal tubercle, with scattered long, simple setae, some of which extend beyond tip of dactylus (
|
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|
|
<figureCitation id="AF804384B276FFA6FE9D90DEFE18FB1B" box="[372,446,1208,1234]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
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,
|
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<figureCitation id="AF804384B276FFA6FE2490DEFE55FB1B" box="[461,499,1208,1234]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="2" pageNumber="3">2A</figureCitation>
|
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,
|
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<figureCitation id="AF804384B276FFA6FE1690DEFD83FB1B" box="[511,549,1208,1234]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="2" pageNumber="3">3A</figureCitation>
|
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,
|
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|
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<figureCitation id="AF804384B276FFA6FDD890DEFDFEFB1B" box="[561,600,1208,1234]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="2" pageNumber="3">4C</figureCitation>
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); thoracic sternum with surface distinctly pitted (
|
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|
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<figureCitation id="AF804384B276FFA6FB7F90DEFB53FB1B" box="[1174,1269,1208,1234]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="2" pageNumber="3">Figs 1B</figureCitation>
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,
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<figureCitation id="AF804384B276FFA6FAEB90D1FA81FB1B" box="[1282,1319,1207,1234]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="2" pageNumber="3">5B</figureCitation>
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); male abdomen longitudinally narrow, surfaces of sternites 1–3 pitted (
|
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<figureCitation id="AF804384B276FFA6FC5D90BBFBB6FB31" box="[948,1040,1245,1272]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="2" pageNumber="3">Fig. 5C</figureCitation>
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); G1 elongate, gently sinuous, distal part curves upwards (
|
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<figureCitation id="AF804384B276FFA6FDE99165FDE2FAD7" box="[512,580,1283,1310]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
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D–F).
|
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</paragraph>
|
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</subSubSection>
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|
|
<subSubSection id="7FA10C8AB276FFA6FF549121FBF2F9CA" pageId="2" pageNumber="3" type="etymology">
|
|||
|
|
<paragraph id="37045F01B276FFA6FF549121FEE2FAA8" blockId="2.[189,324,1351,1377]" box="[189,324,1351,1377]" pageId="2" pageNumber="3">
|
|||
|
|
<heading id="6C4CE86DB276FFA6FF549121FEE2FAA8" bold="true" box="[189,324,1351,1377]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
|||
|
|
<emphasis id="05CF8313B276FFA6FF549121FEE2FAA8" bold="true" box="[189,324,1351,1377]" pageId="2" pageNumber="3">Etymology</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B276FFA6FF54911EFBF2F9CA" blockId="2.[189,1399,1400,1540]" pageId="2" pageNumber="3">
|
|||
|
|
The species name is in honour of His Serene Highness Albert II, Prince of
|
|||
|
|
<collectingCountry id="4FAC1F91B276FFA6FB8C911FFB6EFA5A" box="[1125,1224,1401,1427]" name="Monaco" pageId="2" pageNumber="3">Monaco</collectingCountry>
|
|||
|
|
, patron of the PAPUA NIUGINI Expedition and several other biodiversity expeditions of the “Our Planet Reviewed” programme conducted by MNHN and Pro-Natura International. The red and white colour pattern of the new species also alludes to the colours associated with the House of Grimaldi.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="7FA10C8AB276FFA0FF549248FBF9FCD1" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="materials_examined">
|
|||
|
|
<paragraph id="37045F01B276FFA6FF549248FECAF98E" blockId="2.[189,364,1581,1608]" box="[189,364,1581,1608]" pageId="2" pageNumber="3">
|
|||
|
|
<heading id="6C4CE86DB276FFA6FF549248FECAF98E" bold="true" box="[189,364,1581,1608]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
|||
|
|
<emphasis id="05CF8313B276FFA6FF549248FECAF98E" bold="true" box="[189,364,1581,1608]" pageId="2" pageNumber="3">Type material</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<materialsCitation id="87D3555CB276FFA6FF039239FA8EF90D" ID-GBIF-Occurrence="2610058770" collectingDate="2012-12-04" collectionCode="MNHN" country="Papua New Guinea" elevation="12" latitude="-5.116667" location="Expedition" longLatPrecision="130" longitude="145.82333" municipality="Expedition" pageId="2" pageNumber="3" specimenCode="MNHN IU 2013-921" specimenCount="1" specimenCount-male="1" stateProvince="Madang District" typeStatus="holotype">
|
|||
|
|
<paragraph id="37045F01B276FFA6FF039239FEFAF9B0" blockId="2.[189,1399,1631,1733]" box="[234,348,1631,1657]" pageId="2" pageNumber="3">
|
|||
|
|
<emphasis id="05CF8313B276FFA6FF039239FEFAF9B0" bold="true" box="[234,348,1631,1657]" pageId="2" pageNumber="3">
|
|||
|
|
<typeStatus id="E800E1A3B276FFA6FF039239FEFAF9B0" box="[234,348,1631,1657]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B276FFA6FF5492E3FA8EF90D" blockId="2.[189,1399,1631,1733]" pageId="2" pageNumber="3">
|
|||
|
|
<collectingCountry id="4FAC1F91B276FFA6FF5492E3FE46F957" box="[189,480,1668,1695]" name="Papua New Guinea" pageId="2" pageNumber="3">PAPUA NEW GUINEA</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="F57F91E3B276FFA6FE0592E2FD09F957" box="[492,687,1668,1694]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="3">Madang District</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="D760C57BB276FFA6FD5592E2FC9BF957" box="[700,829,1668,1694]" pageId="2" pageNumber="3">Expedition</collectingMunicipality>
|
|||
|
|
PAPUA NIUGINI, stn PB35, outer slope,
|
|||
|
|
<locationDeviation id="26B64623B276FFA6FADE92E2FE8CF90C" pageId="2" pageNumber="3">south of Sek Is</locationDeviation>
|
|||
|
|
.,
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FED292CCFE17F90C" box="[315,433,1706,1733]" degrees="05" direction="south" minutes="07.0" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-5.116667">05°07.0’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FE5092CCFDE5F90C" box="[441,579,1706,1733]" degrees="145" direction="east" minutes="49.4" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="145.82333">145°49.4’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="F043F2E4B276FFA6FDA792CCFD2AF90C" box="[590,652,1706,1733]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="12.0">
|
|||
|
|
<elevation id="BC96B832B276FFA6FDA792CCFD2AF90C" box="[590,652,1706,1733]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="12.0">12 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="430579C1B276FFA6FD7092CCFC8DF90D" box="[665,811,1706,1733]" pageId="2" pageNumber="3" value="2012-12-04">
|
|||
|
|
<collectingDate id="53418029B276FFA6FD7092CCFC8DF90D" box="[665,811,1706,1733]" pageId="2" pageNumber="3" value="2012-12-04">4 Dec. 2012</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B276FFA6FCDF92CCFCC2F90C" box="[822,868,1706,1733]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
|
|||
|
|
, 6.2 ×
|
|||
|
|
<quantity id="F043F2E4B276FFA6FC5D92CCFBA8F90C" box="[948,1038,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="3" unit="mm" value="4.4">4.4 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<specimenCode id="671DF77AB276FFA6FBF492CCFABAF90D" box="[1053,1308,1706,1733]" pageId="2" pageNumber="3">MNHN IU 2013-921</specimenCode>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</materialsCitation>
|
|||
|
|
<paragraph id="37045F01B276FFA6FF039288FEC1F8C1" blockId="2.[189,1399,1774,2027]" box="[234,359,1774,1800]" pageId="2" pageNumber="3">
|
|||
|
|
<emphasis id="05CF8313B276FFA6FF039288FEC1F8C1" bold="true" box="[234,359,1774,1800]" pageId="2" pageNumber="3">
|
|||
|
|
<typeStatus id="E800E1A3B276FFA6FF039288FEC1F8C1" box="[234,359,1774,1800]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B276FFA6FF549372FCA6F822" blockId="2.[189,1399,1774,2027]" pageId="2" pageNumber="3">
|
|||
|
|
<collectingCountry id="4FAC1F91B276FFA6FF549372FE44F8E7" box="[189,482,1812,1838]" name="Papua New Guinea" pageId="2" pageNumber="3">PAPUA NEW GUINEA</collectingCountry>
|
|||
|
|
: same data as
|
|||
|
|
<typeStatus id="E800E1A3B276FFA6FD799372FD5CF8E7" box="[656,762,1812,1838]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B276FFA6FCEE9372FC91F8E7" box="[775,823,1812,1838]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
|
|||
|
|
, 5.1 ×
|
|||
|
|
<quantity id="F043F2E4B276FFA6FC649372FC4EF8E7" box="[909,1000,1812,1838]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="3" unit="mm" value="3.8">3.8 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FC109372FB90F8E7" box="[1017,1078,1812,1838]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="3">ZRC</collectionCode>
|
|||
|
|
, ex
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FB809372FB65F8E7" box="[1129,1219,1812,1838]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FB259372FB4DF8E7" box="[1228,1259,1812,1838]" country="USA" name="Indiana University" pageId="2" pageNumber="3">IU</collectionCode>
|
|||
|
|
2013-920);
|
|||
|
|
<collectingRegion id="F57F91E3B276FFA6FF54935CFE27F89D" box="[189,385,1850,1876]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="3">Madang District</collectingRegion>
|
|||
|
|
, Expedition PAPUA NIUGINI, stn PR17, west of Kranket Is.,
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FB82935FFB46F89D" box="[1131,1248,1849,1876]" degrees="05" direction="south" minutes="11.8" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-5.1966667">05°11.8’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FB01935FFAD4F89D" box="[1256,1394,1849,1876]" degrees="145" direction="east" minutes="48.8" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="145.81334">145°48.8’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="F043F2E4B276FFA6FF549339FF5FF8B3" box="[189,249,1887,1914]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="15.0">15 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="430579C1B276FFA6FEEA9306FE06F8B3" box="[259,416,1888,1914]" pageId="2" pageNumber="3" value="2012-11-11">11 Nov. 2012</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B276FFA6FE409306FE75F8B3" box="[425,467,1888,1914]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
|
|||
|
|
, 10.2 ×
|
|||
|
|
<quantity id="F043F2E4B276FFA6FDC69306FD21F8B3" box="[559,647,1888,1914]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="3" unit="mm" value="7.8">7.8 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FD7D9306FD77F8B3" box="[660,721,1888,1914]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="3">ZRC</collectionCode>
|
|||
|
|
, ex
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FD159306FCF0F8B3" box="[764,854,1888,1914]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FCB29306FCDCF8B3" box="[859,890,1888,1914]" country="USA" name="Indiana University" pageId="2" pageNumber="3">IU</collectionCode>
|
|||
|
|
2013-1245); Expedition PAPUA NIUGINI, stn PR05, Kranket Is.,
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FE2393E3FDE6F869" box="[458,576,1925,1952]" degrees="05" direction="south" minutes="11.3" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-5.1883335">05°11.3’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FDAE93E3FD77F869" box="[583,721,1925,1952]" degrees="145" direction="east" minutes="49.6" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="145.82666">145°49.6’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<date id="430579C1B276FFA6FD3793E0FCD4F856" box="[734,882,1925,1952]" pageId="2" pageNumber="3" value="2012-11-07">7 Nov. 2012</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B276FFA6FC9793E3FC0DF869" box="[894,939,1925,1952]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
|
|||
|
|
, 8.5 ×
|
|||
|
|
<quantity id="F043F2E4B276FFA6FC1693E3FBFCF869" box="[1023,1114,1925,1952]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="3" unit="mm" value="6.2">6.2 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FB8093E3FB63F856" box="[1129,1221,1925,1951]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FB2493E0FB4AF869" box="[1229,1260,1926,1952]" country="USA" name="Indiana University" pageId="2" pageNumber="3">IU</collectionCode>
|
|||
|
|
2013-222); Expedition PAPUA NIUGINI, stn PB28, east of Kranket Is.,
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FC9593CDFC54F80F" box="[892,1010,1963,1990]" degrees="05" direction="south" minutes="11.9" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-5.1983333">05°11.9’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="528F39C6B276FFA6FC1193CDFB27F80F" box="[1016,1153,1963,1990]" degrees="145" direction="east" minutes="49.6" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="145.82666">145°49.6’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="F043F2E4B276FFA6FB6593CDFB6EF80C" box="[1164,1224,1963,1989]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="10.0">10 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="430579C1B276FFA6FB3A93CDFAD4F80C" box="[1235,1394,1963,1989]" pageId="2" pageNumber="3" value="2012-11-24">24 Nov. 2012</date>
|
|||
|
|
, 1 young
|
|||
|
|
<specimenCount id="21BD9488B276FFA6FECD93B7FE9CF822" box="[292,314,2001,2027]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
|
|||
|
|
, 6.7 ×
|
|||
|
|
<quantity id="F043F2E4B276FFA6FE6593B7FE40F822" box="[396,486,2001,2027]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="3" unit="mm" value="4.9">4.9 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FE1C93B7FDF7F822" box="[501,593,2001,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
|
|||
|
|
<collectionCode id="51AAC7C4B276FFA6FDB193B7FDD1F822" box="[600,631,2001,2027]" country="USA" name="Indiana University" pageId="2" pageNumber="3">IU</collectionCode>
|
|||
|
|
2013-759).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="63C40F89B277FFA7FF549322FAF3F822" ID-DOI="http://doi.org/10.5281/zenodo.3785522" ID-Zenodo-Dep="3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1860,1886]" targetBox="[189,1399,399,1819]" targetPageId="3">
|
|||
|
|
<paragraph id="37045F01B277FFA7FF549322FAF3F822" blockId="3.[189,1399,1860,2027]" pageId="3" pageNumber="4">
|
|||
|
|
<emphasis id="05CF8313B277FFA7FF549322FEADF897" bold="true" box="[189,267,1860,1886]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
|||
|
|
<taxonomicName id="F0BB2482B277FFA7FEFA9322FE7CF897" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[275,474,1860,1886]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
|||
|
|
<emphasis id="05CF8313B277FFA7FEFA9322FE7CF897" box="[275,474,1860,1886]" italics="true" pageId="3" pageNumber="4">Actaea grimaldii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="1EFC3E68B277FFA7FE0B9322FDE6F897" box="[482,576,1860,1886]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, Papua New Guinea, colours in life.
|
|||
|
|
<emphasis id="05CF8313B277FFA7FC1D9322FB8AF897" bold="true" box="[1012,1068,1860,1886]" pageId="3" pageNumber="4">A–B</emphasis>
|
|||
|
|
. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920).
|
|||
|
|
<emphasis id="05CF8313B277FFA7FDA99301FDF3F848" bold="true" box="[576,597,1895,1921]" pageId="3" pageNumber="4">C</emphasis>
|
|||
|
|
. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921).
|
|||
|
|
<emphasis id="05CF8313B277FFA7FB289301FB70F848" bold="true" box="[1217,1238,1895,1921]" pageId="3" pageNumber="4">D</emphasis>
|
|||
|
|
. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245).
|
|||
|
|
<emphasis id="05CF8313B277FFA7FCEA93ECFCB1F86D" bold="true" box="[771,791,1930,1956]" pageId="3" pageNumber="4">E</emphasis>
|
|||
|
|
. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222).
|
|||
|
|
<emphasis id="05CF8313B277FFA7FF5493C8FF69F801" bold="true" box="[189,207,1966,1992]" pageId="3" pageNumber="4">F</emphasis>
|
|||
|
|
. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759).
|
|||
|
|
<emphasis id="05CF8313B277FFA7FCC393CBFCC0F801" bold="true" box="[810,870,1965,1992]" pageId="3" pageNumber="4">G–H</emphasis>
|
|||
|
|
. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF54956DFE75FEEC" blockId="4.[189,467,267,293]" box="[189,467,267,293]" pageId="4" pageNumber="5">
|
|||
|
|
<heading id="6C4CE86DB270FFA0FF54956DFE75FEEC" bold="true" box="[189,467,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF54956DFE75FEEC" bold="true" box="[189,467,267,293]" pageId="4" pageNumber="5">Comparative material</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF03955CFD10FE9D" blockId="4.[189,1399,314,518]" box="[234,694,314,340]" pageId="4" pageNumber="5">
|
|||
|
|
<taxonomicName id="F0BB2482B270FFA0FF03955CFD10FE9D" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[234,694,314,340]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="spinosissima">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF03955CFD10FE9D" bold="true" box="[234,694,314,340]" pageId="4" pageNumber="5">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF03955CFE78FE9D" bold="true" box="[234,478,314,340]" italics="true" pageId="4" pageNumber="5">Actaea spinosissima</emphasis>
|
|||
|
|
<bibRefCitation id="532A22F0B270FFA0FE0C955CFD10FE9D" author="Borradaile L. A." box="[485,694,314,340]" pageId="4" pageNumber="5" pagination="237 - 271" refId="ref6135" refString="Borradaile L. A. 1902. Marine crustaceans III: The Xanthidae and some other crabs. In: Gardiner J. S. (ed.) The Fauna and Geography of the Maldive and Laccadive Archipelagoes, being the Account of the Work carried on and of the Collections made by an Expedition during the Years 1899 and 1900. Vol. 1. Part. 3: 237 - 271. Cambridge University Press, Cambridge. http: // dx. doi. org / 10.5962 / bhl. title. 10215" type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF54953BFDB4FE55" blockId="4.[189,1399,314,518]" pageId="4" pageNumber="5">
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FF54953BFEFFFEBE" box="[189,345,349,375]" name="Maldives" pageId="4" pageNumber="5">MALDIVES</collectingCountry>
|
|||
|
|
: Mahlos Atoll, 23 fathoms, J.S. Gardiner coll.,
|
|||
|
|
<typeStatus id="E800E1A3B270FFA0FC7D953BFC58FEBE" box="[916,1022,349,375]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B270FFA0FBE29538FB85FEB1" box="[1035,1059,350,376]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
|
|||
|
|
, 5.1 ×
|
|||
|
|
<quantity id="F043F2E4B270FFA0FB93953BFB71FEB1" box="[1146,1239,349,376]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="5" unit="mm" value="3.4">3.4 mm</quantity>
|
|||
|
|
(juvenile, no gonopods) (
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FEA195E7FE39FE52" box="[328,415,385,411]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="4" pageNumber="5">CUMZ</collectionCode>
|
|||
|
|
I.63106).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF5495C3FED4FDCF" blockId="4.[189,1399,314,518]" pageId="4" pageNumber="5">
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FF5495C3FD1BFE76" box="[189,701,421,447]" name="British Indian Ocean Territory" pageId="4" pageNumber="5">BRITISH INDIAN OCEAN TERRITORY</collectingCountry>
|
|||
|
|
: Great Chagos Bank, stn
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FC0795C2FBBEFE77" box="[1006,1048,420,446]" name="Switzerland" pageId="4" pageNumber="5">CH</collectingCountry>
|
|||
|
|
0666, outer reef, among dead branching coral heads,
|
|||
|
|
<quantity id="F043F2E4B270FFA0FE3995AEFDA8FE2A" box="[464,526,456,483]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</quantity>
|
|||
|
|
, Brothers Island, C. Head & H. Koldewey coll.,
|
|||
|
|
<date id="430579C1B270FFA0FBB895AEFB49FE2B" box="[1105,1263,456,482]" pageId="4" pageNumber="5" value="2013-02-28">28 Feb. 2013</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B270FFA0FB1595AEFA8DFE2A" box="[1276,1323,456,483]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
|
|||
|
|
, 8.9 ×
|
|||
|
|
<quantity id="F043F2E4B270FFA0FF54958AFEB1FDCF" box="[189,279,492,518]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="5" unit="mm" value="6.6">6.6 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FECF958AFEC3FDCF" box="[294,357,492,518]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="5">ZRC</collectionCode>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF03964BFD2EFD8E" blockId="4.[189,1398,557,691]" box="[234,648,557,584]" pageId="4" pageNumber="5">
|
|||
|
|
<taxonomicName id="F0BB2482B270FFA0FF03964BFD2EFD8E" authority="(Heller, 1861)" baseAuthorityName="Heller" baseAuthorityYear="1861" box="[234,648,557,584]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="polyacantha">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF03964BFD2EFD8E" bold="true" box="[234,648,557,584]" pageId="4" pageNumber="5">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF03964BFE7FFD81" bold="true" box="[234,473,557,584]" italics="true" pageId="4" pageNumber="5">Actaea polyacantha</emphasis>
|
|||
|
|
(Heller, 1861)
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549637FB10FD7A" blockId="4.[189,1398,557,691]" pageId="4" pageNumber="5">
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FF549637FEBDFDA2" box="[189,283,593,619]" name="Guam" pageId="4" pageNumber="5">GUAM</collectingCountry>
|
|||
|
|
: southwest of Orote Peninsula, G. Paulay coll.,
|
|||
|
|
<date id="430579C1B270FFA0FCA39637FBB4FDA2" box="[842,1042,593,620]" pageId="4" pageNumber="5" value="2000-04-15" valueMax="2000-04-18" valueMin="2000-04-15">15–18 Apr. 2000</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B270FFA0FBF49637FBEFFDA5" box="[1053,1097,593,620]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FBB09637FB32FDA2" box="[1113,1172,593,619]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="5">ZRC</collectionCode>
|
|||
|
|
2000.731).
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FF549613FE4CFD46" box="[189,490,629,655]" name="Papua New Guinea" pageId="4" pageNumber="5">PAPUA NEW GUINEA</collectingCountry>
|
|||
|
|
: Expedition PAPUA NIUGINI, stn PB17, outer slope, Sek Is.,
|
|||
|
|
<geoCoordinate id="528F39C6B270FFA0FAE99612FAD0FD46" box="[1280,1398,628,655]" degrees="05" direction="south" minutes="04.9" orientation="latitude" pageId="4" pageNumber="5" precision="92" value="-5.0816665">05°04.9’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="528F39C6B270FFA0FF5496FEFEE0FD7A" box="[189,326,664,691]" degrees="145" direction="east" minutes="49.3" orientation="longitude" pageId="4" pageNumber="5" precision="92" value="145.82167">145°49.3’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="F043F2E4B270FFA0FEBB96FEFE36FD7A" box="[338,400,664,691]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="5" unit="m" value="26.0">26 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="430579C1B270FFA0FE7496FEFD98FD7B" box="[413,574,664,691]" pageId="4" pageNumber="5" value="2012-11-14">14 Nov. 2012</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="21BD9488B270FFA0FDA396FEFDD0FD7A" box="[586,630,664,691]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
|
|||
|
|
, 6.4 ×
|
|||
|
|
<quantity id="F043F2E4B270FFA0FD2196FEFC84FD7A" box="[712,802,664,691]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="5" unit="mm" value="4.9">4.9 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FCDB96FEFCC9FD7B" box="[818,879,664,690]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="5">ZRC</collectionCode>
|
|||
|
|
, ex
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FC7796FFFC5EFD7A" box="[926,1016,665,691]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FC1696FFFBB8FD7A" box="[1023,1054,665,691]" country="USA" name="Indiana University" pageId="4" pageNumber="5">IU</collectionCode>
|
|||
|
|
2013-1255).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF0396BCFD40FD3D" blockId="4.[189,1119,730,792]" box="[234,742,730,756]" pageId="4" pageNumber="5">
|
|||
|
|
<taxonomicName id="F0BB2482B270FFA0FF0396BCFD40FD3D" authority="(H. Milne Edwards, 1834)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[234,742,730,756]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="peronii">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF0396BCFD40FD3D" bold="true" box="[234,742,730,756]" pageId="4" pageNumber="5">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF0396BCFE3DFD3D" bold="true" box="[234,411,730,756]" italics="true" pageId="4" pageNumber="5">Actaea peronii</emphasis>
|
|||
|
|
(H. Milne Edwards, 1834)
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549698FBF9FCD1" blockId="4.[189,1119,730,792]" box="[189,1119,765,792]" pageId="4" pageNumber="5">
|
|||
|
|
<collectingCountry id="4FAC1F91B270FFA0FF549698FECEFCD1" box="[189,360,766,792]" name="Australia" pageId="4" pageNumber="5">AUSTRALIA</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="F57F91E3B270FFA0FE9A9698FDE1FCDE" box="[371,583,765,792]" country="Australia" name="New South Wales" pageId="4" pageNumber="5">New South Wales</collectingRegion>
|
|||
|
|
, M. Ward coll.,
|
|||
|
|
<specimenCount id="21BD9488B270FFA0FCEA9698FCE3FCD1" box="[771,837,766,792]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="51AAC7C4B270FFA0FCBD9698FC29FCD1" box="[852,911,766,792]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="5">ZRC</collectionCode>
|
|||
|
|
1969.12.27.3–4).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="7FA10C8AB270FFADFF549759FBD2F822" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="description">
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549759FEE8FC90" blockId="4.[189,334,831,857]" box="[189,334,831,857]" pageId="4" pageNumber="5">
|
|||
|
|
<heading id="6C4CE86DB270FFA0FF549759FEE8FC90" bold="true" box="[189,334,831,857]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
|||
|
|
<emphasis id="05CF8313B270FFA0FF549759FEE8FC90" bold="true" box="[189,334,831,857]" pageId="4" pageNumber="5">Description</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549708FCBFFA9E" blockId="4.[189,1399,878,1367]" pageId="4" pageNumber="5">
|
|||
|
|
Regions well demarcated, separated by distinct grooves, anterior surfaces (notably 1M, 2M, 3M, 2L, 5L) covered with numerous spines and sharp tubercles of varying sizes; 2M and 2L surface gently convex, does not appear inflated in frontal view; 2M separated by median longitudinal groove anteriorly; 3M subtriangular, with sharp spines; 4M lunate, with sharp granules; 1P flattened, divided into many smaller regions by shallow grooves, with distinct transverse groove on anterior third, confluent with series of flattened granules on 3R, 2R and 1R; 2O transversely narrow, low; 2P raised, surface rounded (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FACD9046FAC9FBF3" box="[1316,1391,1056,1082]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FF549022FF45FB97" box="[189,227,1092,1118]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FF069022FEB3FB97" box="[239,277,1092,1118]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FECB9022FEECFB97" box="[290,330,1092,1118]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="4" pageNumber="5">4A</figureCitation>
|
|||
|
|
). Frontal margin bilobate, lobes separated by broad, U-shaped cleft; margins lined by sharp granules; 1F and 2F with many sharp spines which project anteriorly, hiding frontal margin from dorsal view (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FEE090EDFEF1FB6C" box="[265,343,1163,1189]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FE8390EDFEDEFB6C" box="[362,376,1163,1189]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
|
|||
|
|
A–B, 3A–B, 4A). Supraorbital margin with 3 distinct spines; orbit short, ovate; eye completely filling orbit; peduncle short, stout, with several dorsal granules; cornea large, with granule basally (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FECD90B4FED7FB25" box="[292,369,1234,1260]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
|
|||
|
|
A–B, 3A–B, 4A–B). Anterolateral margin arcuate, with 4 spines (excluding external orbital spine), each with smaller basal spine or spines on sub-branchial surface; posterolateral margin gently concave, surfaces covered with small, flattened granules; posterior carapace gently sinuous, lined with small, flattened granules (
|
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|
|
<figureCitation id="AF804384B270FFA0FDCF915BFDD6FA9E" box="[550,624,1341,1367]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FD96915BFD03FA9E" box="[639,677,1341,1367]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2A</figureCitation>
|
|||
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,
|
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|
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<figureCitation id="AF804384B270FFA0FD58915BFD71FA9E" box="[689,727,1341,1367]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3A</figureCitation>
|
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|
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,
|
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|
|
<figureCitation id="AF804384B270FFA0FD0D915BFCAAFA9E" box="[740,780,1341,1367]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="4" pageNumber="5">4A</figureCitation>
|
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|
).
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549118FEC3F982" blockId="4.[189,1399,1406,1611]" pageId="4" pageNumber="5">
|
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|
|
Suborbital margin with 5 or 6 small spines; suborbital and subhepatic regions covered with numerous small, sharp granules; pterygostomian region almost smooth (
|
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|
|
<figureCitation id="AF804384B270FFA0FC4391C4FBB6FA75" box="[938,1040,1442,1468]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1H</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FBC991C4FBE3FA75" box="[1056,1093,1442,1468]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2B</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FBBD91C4FBDFFA75" box="[1108,1145,1442,1468]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3B</figureCitation>
|
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|
). Antennules folding laterally (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FEC691A0FE34FA29" box="[303,402,1478,1504]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2B</figureCitation>
|
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,
|
|||
|
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<figureCitation id="AF804384B270FFA0FE4991A0FE63FA29" box="[416,453,1478,1504]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3B</figureCitation>
|
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|
|
). Antenna with basal article quadrate, with 1 large and 1 small, sharp granule, mobile; flagellum short, at base of orbital hiatus (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FCCB918FFC21F9CA" box="[802,903,1513,1539]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2B</figureCitation>
|
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,
|
|||
|
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<figureCitation id="AF804384B270FFA0FC7E918FFC1AF9CA" box="[919,956,1513,1539]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3B</figureCitation>
|
|||
|
|
). Epistome transversely rectangular; anterior margin granulated; posterior margin with prominent median triangle, lateral margins sinuous (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FF2C9256FE80F982" box="[197,294,1584,1611]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2B</figureCitation>
|
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|
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FEDB9257FEF1F982" box="[306,343,1585,1611]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3B</figureCitation>
|
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|
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).
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B270FFA0FF549214FAC5F91D" blockId="4.[189,1399,1650,1748]" pageId="4" pageNumber="5">
|
|||
|
|
Third maxilliped relatively short; outer surfaces pitted; ischium subrectangular, with shallow but distinct submedian sulcus, inner margin dentate; merus quadrate, distolateral angle not distinctly auricular; exopod stout, distal end reaching to distolateral edge of merus, flagellum elongate, as wide as merus (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FB1192DFFAF0F91D" box="[1272,1366,1721,1748]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="4" pageNumber="5">Fig. 5A</figureCitation>
|
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).
|
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</paragraph>
|
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|
<paragraph id="37045F01B270FFADFF54929DFD75FEA5" blockId="4.[189,1399,1787,2027]" lastBlockId="9.[189,1399,267,364]" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5">
|
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|
|
Chelipeds slightly asymmetrical; outer surfaces with numerous sharp spines and sharp tubercles, with small, sharp granules interspersed; inner and outer surfaces with scattered short, simple setae (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FACA9379FAC9F8F0" box="[1315,1391,1823,1849]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
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,
|
|||
|
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<figureCitation id="AF804384B270FFA0FF549324FF45F895" box="[189,227,1858,1884]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2A</figureCitation>
|
|||
|
|
, C–D, 3A, C–D). Basis-ischium lined with small granules. Merus short, rounded; dorso-distal spine largest. Carpus ovate, inner angle with several short, curved spines (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FC3B9300FBBAF849" box="[978,1052,1894,1920]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FBC09300FBE9F849" box="[1065,1103,1894,1920]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">2A</figureCitation>
|
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,
|
|||
|
|
<figureCitation id="AF804384B270FFA0FBB29300FB24F849" box="[1115,1154,1894,1920]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="4" pageNumber="5">3A</figureCitation>
|
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|
). Chelae short, stout, palm longer than fingers, not pigmented black or dark brown; inner surface with median spine and several smaller sharp tubercles and granules (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FD2493CBFCB0F80E" box="[717,790,1965,1991]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
|
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|
|
C–D, 3C–D). Pollex with 2 low, longitudinal ridges, 1 submedian groove; cutting edge with 3 or 4 large, blade-like teeth; tip subspatuliform (
|
|||
|
|
<figureCitation id="AF804384B270FFA0FB0293B7FA91F822" box="[1259,1335,2001,2027]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
|
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C–D, 3C–D). Dactylus with 1 distinct low, longitudinal ridge, 1 submedian groove; upper margin with 2–4 spines on proximal half, cutting edge with 2 larger teeth on proximal half, distal half with 2 or 3 low teeth; tip subspatuliform (
|
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|
|
<figureCitation id="AF804384B27DFFADFE029534FD93FEA5" box="[491,565,338,364]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="9" pageNumber="10">Figs 2</figureCitation>
|
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|
C–D, 3C–D).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="63C40F89B271FFA1FF5493ECFE97F822" ID-DOI="http://doi.org/10.5281/zenodo.3785524" ID-Zenodo-Dep="3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1930,1956]" targetBox="[189,1399,296,1890]" targetPageId="5">
|
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|
|
<paragraph id="37045F01B271FFA1FF5493ECFE97F822" blockId="5.[189,1399,1930,2027]" pageId="5" pageNumber="6">
|
|||
|
|
<emphasis id="05CF8313B271FFA1FF5493ECFEAFF86D" bold="true" box="[189,265,1930,1956]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
|
|||
|
|
<taxonomicName id="F0BB2482B271FFA1FEE693EDFE72F86D" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[271,468,1930,1957]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
|||
|
|
<emphasis id="05CF8313B271FFA1FEE693EDFE72F86D" box="[271,468,1930,1957]" italics="true" pageId="5" pageNumber="6">Actaea grimaldii</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="1EFC3E68B271FFA1FE3393EDFD90F86C" box="[474,566,1931,1957]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea.
|
|||
|
|
<emphasis id="05CF8313B271FFA1FF5493CBFF74F80E" bold="true" box="[189,210,1965,1991]" pageId="5" pageNumber="6">A</emphasis>
|
|||
|
|
. Dorsal view of habitus.
|
|||
|
|
<emphasis id="05CF8313B271FFA1FE1593C8FDB6F801" bold="true" box="[508,528,1966,1992]" pageId="5" pageNumber="6">B</emphasis>
|
|||
|
|
. Frontal view of carapace.
|
|||
|
|
<emphasis id="05CF8313B271FFA1FCBB93CBFCC1F80E" bold="true" box="[850,871,1965,1991]" pageId="5" pageNumber="6">C</emphasis>
|
|||
|
|
. Outer view of right chela.
|
|||
|
|
<emphasis id="05CF8313B271FFA1FB4793C8FB65F801" bold="true" box="[1198,1219,1966,1992]" pageId="5" pageNumber="6">D</emphasis>
|
|||
|
|
. Outer view of left chela.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="63C40F89B272FFA2FF5493ECFE59F822" ID-DOI="http://doi.org/10.5281/zenodo.3785526" ID-Zenodo-Dep="3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1930,1956]" targetBox="[205,1383,265,1890]" targetPageId="6">
|
|||
|
|
<paragraph id="37045F01B272FFA2FF5493ECFE59F822" blockId="6.[189,1399,1930,2027]" pageId="6" pageNumber="7">
|
|||
|
|
<emphasis id="05CF8313B272FFA2FF5493ECFEADF86D" bold="true" box="[189,267,1930,1956]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
|
|||
|
|
<taxonomicName id="F0BB2482B272FFA2FEFA93EDFE7FF86D" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[275,473,1930,1957]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
|||
|
|
<emphasis id="05CF8313B272FFA2FEFA93EDFE7FF86D" box="[275,473,1930,1957]" italics="true" pageId="6" pageNumber="7">Actaea grimaldii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="1EFC3E68B272FFA2FE0893EDFD98F86C" box="[481,574,1931,1957]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea.
|
|||
|
|
<emphasis id="05CF8313B272FFA2FE8C93CBFEDCF80E" bold="true" box="[357,378,1965,1991]" pageId="6" pageNumber="7">A</emphasis>
|
|||
|
|
. Dorsal view of habitus.
|
|||
|
|
<emphasis id="05CF8313B272FFA2FD5B93C8FD60F801" bold="true" box="[690,710,1966,1992]" pageId="6" pageNumber="7">B</emphasis>
|
|||
|
|
. Frontal view of carapace.
|
|||
|
|
<emphasis id="05CF8313B272FFA2FBFC93CBFB8CF80E" bold="true" box="[1045,1066,1965,1991]" pageId="6" pageNumber="7">C</emphasis>
|
|||
|
|
. Outer view of right chela.
|
|||
|
|
<emphasis id="05CF8313B272FFA2FF5493B7FF74F822" bold="true" box="[189,210,2001,2027]" pageId="6" pageNumber="7">D</emphasis>
|
|||
|
|
. Outer view of left chela.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="63C40F89B273FFA3FF5493ECFED3F822" ID-DOI="http://doi.org/10.5281/zenodo.3785528" ID-Zenodo-Dep="3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1930,1956]" targetBox="[189,1399,436,1890]" targetPageId="7">
|
|||
|
|
<paragraph id="37045F01B273FFA3FF5493ECFED3F822" blockId="7.[189,1399,1930,2027]" pageId="7" pageNumber="8">
|
|||
|
|
<emphasis id="05CF8313B273FFA3FF5493ECFEADF86D" bold="true" box="[189,267,1930,1956]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
|
|||
|
|
<taxonomicName id="F0BB2482B273FFA3FEFA93EDFE7FF86D" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[275,473,1930,1957]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
|||
|
|
<emphasis id="05CF8313B273FFA3FEFA93EDFE7FF86D" box="[275,473,1930,1957]" italics="true" pageId="7" pageNumber="8">Actaea grimaldii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="1EFC3E68B273FFA3FE0893EDFD98F86C" box="[481,574,1931,1957]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea.
|
|||
|
|
<emphasis id="05CF8313B273FFA3FE8293CBFE26F80E" bold="true" box="[363,384,1965,1991]" pageId="7" pageNumber="8">A</emphasis>
|
|||
|
|
. Dorsal view of carapace.
|
|||
|
|
<emphasis id="05CF8313B273FFA3FD3093C8FD4BF801" bold="true" box="[729,749,1966,1992]" pageId="7" pageNumber="8">B</emphasis>
|
|||
|
|
. Thoracic sternum showing vulvae.
|
|||
|
|
<emphasis id="05CF8313B273FFA3FB5F93CBFB6DF80E" bold="true" box="[1206,1227,1965,1991]" pageId="7" pageNumber="8">C</emphasis>
|
|||
|
|
. Right fourth ambulatory leg.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="63C40F89B27CFFACFF549301FAD1F822" ID-DOI="http://doi.org/10.5281/zenodo.3785530" ID-Zenodo-Dep="3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1895,1921]" targetBox="[213,1389,376,1821]" targetPageId="8">
|
|||
|
|
<paragraph id="37045F01B27CFFACFF549301FAD1F822" blockId="8.[189,1399,1895,2027]" pageId="8" pageNumber="9">
|
|||
|
|
<emphasis id="05CF8313B27CFFACFF549301FEAFF848" bold="true" box="[189,265,1895,1921]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
|
|||
|
|
<taxonomicName id="F0BB2482B27CFFACFEE6930EFE72F848" authority="Bouchet & Isyeb, 2015" authorityName="Bouchet & Isyeb" authorityYear="2015" box="[271,468,1895,1922]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
|||
|
|
<emphasis id="05CF8313B27CFFACFEE6930EFE72F848" box="[271,468,1895,1922]" italics="true" pageId="8" pageNumber="9">Actaea grimaldii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="1EFC3E68B27CFFACFE33930EFD90F84B" box="[474,566,1896,1922]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFF5493ECFF74F86D" bold="true" box="[189,210,1930,1956]" pageId="8" pageNumber="9">A</emphasis>
|
|||
|
|
. Right third maxilliped (setae not drawn).
|
|||
|
|
<emphasis id="05CF8313B27CFFACFD3593ECFD56F86D" bold="true" box="[732,752,1930,1956]" pageId="8" pageNumber="9">B</emphasis>
|
|||
|
|
. Anterior thoracic sternum (sternites 1–4) and telson.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFF5493CBFF74F80E" bold="true" box="[189,210,1965,1991]" pageId="8" pageNumber="9">C</emphasis>
|
|||
|
|
. Abdominal somites 1–6.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFDE893C8FDB0F801" bold="true" box="[513,534,1966,1992]" pageId="8" pageNumber="9">D</emphasis>
|
|||
|
|
. Ventral view of left G1.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFCD393C8FCE8F801" bold="true" box="[826,846,1966,1992]" pageId="8" pageNumber="9">E</emphasis>
|
|||
|
|
. Ventral view of distal part of left G1.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFAE093C8FABDF801" bold="true" box="[1289,1307,1966,1992]" pageId="8" pageNumber="9">F</emphasis>
|
|||
|
|
. Dorsal view of distal part of left G1.
|
|||
|
|
<emphasis id="05CF8313B27CFFACFDFF93B6FD8BF823" bold="true" box="[534,557,2000,2026]" pageId="8" pageNumber="9">G</emphasis>
|
|||
|
|
. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF5495F2FCB5FCD9" blockId="9.[189,1399,404,784]" pageId="9" pageNumber="10">
|
|||
|
|
Ambulatory legs short, stout; second pair longest; fourth pair shortest (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFC1395F2FBE2FE67" box="[1018,1092,404,430]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFBBB95F2FBDCFE67" box="[1106,1146,404,430]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="9" pageNumber="10">3A</figureCitation>
|
|||
|
|
). Basis-ischium lined with small granules. Outer surfaces of merus, carpus and propodus covered with sharp tubercles or granules (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEDA95BCFE27FE3C" box="[307,385,474,501]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFE7B95BDFE1FFE3C" box="[402,441,475,501]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="9" pageNumber="10">2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFE2195BDFE48FE3C" box="[456,494,475,501]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="9" pageNumber="10">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFE1795BDFD83FE3C" box="[510,549,475,501]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
|||
|
|
). Merus with dorsal margin armed with numerous sharp spines which gradually become smaller proximally, distal one largest; ventral margin lined with small, sharp granules (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFF2C9647FEB6FDF2" box="[197,272,545,571]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEF79647FEE2FDF2" box="[286,324,545,571]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="9" pageNumber="10">2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEB89647FED1FDF2" box="[337,375,545,571]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="9" pageNumber="10">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFE6D9647FE0CFDF2" box="[388,426,545,571]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
|||
|
|
). Dorsal margins of carpus and propodus with several sharp spines; ventral margin of carpus with sharp granules; ventral margin of propodus with sharp spines (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFB9F9623FB62FD96" box="[1142,1220,581,607]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFB3F9623FB5BFD96" box="[1238,1277,581,607]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="9" pageNumber="10">2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFAE49623FA95FD96" box="[1293,1331,581,607]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="9" pageNumber="10">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFAAA9623FACCFD96" box="[1347,1386,581,607]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
|||
|
|
). Dactylus gently curved, surface covered with small, sharp granules; ventral margin with relatively larger subterminal tubercle; tip corneous; surfaces with scattered long, simple setae, those on second to third legs may be very long, extending well beyond tip of dactylus (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFC4196C9FC52FD00" box="[936,1012,687,713]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFBED96C9FB8CFD00" box="[1028,1066,687,713]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,296,1890]" captionTargetId="figure@5.[189,1399,296,1890]" captionTargetPageId="5" captionText="Fig. 2. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785524" httpUri="https://zenodo.org/record/3785524/files/figure.png" pageId="9" pageNumber="10">2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFBD196C9FBFBFD00" box="[1080,1117,687,713]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[205,1383,265,1890]" captionTargetId="figure@6.[205,1383,265,1890]" captionTargetPageId="6" captionText="Fig. 3. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of habitus. B. Frontal view of carapace. C. Outer view of right chela. D. Outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.3785526" httpUri="https://zenodo.org/record/3785526/files/figure.png" pageId="9" pageNumber="10">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFB8596C9FB34FD00" box="[1132,1170,687,713]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
|||
|
|
). Dactylo-propodal lock well developed; distal margin of propodus with rounded median projection that overlaps ball-like swelling on proximal margin of dactylus (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFD409690FCA0FCD9" box="[681,774,758,784]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF549751FB4CFC75" blockId="9.[189,1399,823,956]" pageId="9" pageNumber="10">
|
|||
|
|
Thoracic sternum with surface distinctly pitted (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFD099751FC9AFC98" box="[736,828,823,849]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1B</figureCitation>
|
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|
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,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFCAE9751FCCBFC9B" box="[839,877,823,850]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">5B</figureCitation>
|
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|
|
). Thoracic sternites 1 and 2 completely fused; sternites 2 and 3 separated by distinct transverse suture; sternites 3 and 4 completely fused except for lateral cleft (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEB99718FE09FC51" box="[336,431,894,920]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1B</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFE549718FE44FC50" box="[445,482,894,921]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">5B</figureCitation>
|
|||
|
|
). Sternoabdominal cavity deep, reaching to median part of sternite 4 (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFAFB9718FAD7FC51" box="[1298,1393,894,920]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Figs 1B</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="AF804384B27DFFADFF5497C7FF44FC75" box="[189,226,929,956]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">5C</figureCitation>
|
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|
|
).
|
|||
|
|
<collectingRegion id="F57F91E3B27DFFADFF1E97C7FE95FC72" box="[247,307,929,955]" country="Maldives" name="Maale" pageId="9" pageNumber="10">Male</collectingRegion>
|
|||
|
|
abdominal locking mechanism a knob-like process, on anterior third of sternite 5.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF549785FC0AFB42" blockId="9.[189,1399,995,1163]" pageId="9" pageNumber="10">
|
|||
|
|
<collectingRegion id="F57F91E3B27DFFADFF549785FF5FFC34" box="[189,249,995,1021]" country="Maldives" name="Maale" pageId="9" pageNumber="10">Male</collectingRegion>
|
|||
|
|
abdomen longitudinally narrow; surfaces of sternites 1–5 pitted (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFBC59785FB29FC37" box="[1068,1167,995,1022]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5C</figureCitation>
|
|||
|
|
). Somites 1 and 2 trapezoidal, completely covering surfaces of thoracic sternum between coxae of last pair of ambulatory legs, sternite 8 completely covered; somites 3–5 completely fused, median sutures not discernible; lateral margins concave; somite 6 rectangular, lateral margins gently sinuous, slightly longer than telson; telson triangular, lateral margins gently sinuous, tip rounded (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFCAB9016FC39FB42" box="[834,927,1136,1163]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5C</figureCitation>
|
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|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF5490D4FE97FADD" blockId="9.[189,1398,1202,1300]" pageId="9" pageNumber="10">
|
|||
|
|
G1 elongate, gently sinuous, distal part curves upwards, subdistal surfaces with numerous long setae and spinules (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEC490B3FED6FB39" box="[301,368,1237,1264]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
|||
|
|
D–F). G2 short, c. quarter of length of G1; proximal part sinuous, distal part spatuliform (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFF2F909FFE82FADD" box="[198,292,1273,1300]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5G</figureCitation>
|
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|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF03915DFDA7FA9D" blockId="9.[189,1399,1338,1613]" box="[234,513,1338,1365]" pageId="9" pageNumber="10">
|
|||
|
|
<emphasis id="05CF8313B27DFFADFF03915DFDA7FA9D" bold="true" box="[234,513,1338,1365]" pageId="9" pageNumber="10">Females and variation</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF549138FB4CF985" blockId="9.[189,1399,1338,1613]" pageId="9" pageNumber="10">
|
|||
|
|
The females agree with the males in almost all non-sexual characters. A female measuring 6.7 ×
|
|||
|
|
<quantity id="F043F2E4B27DFFADFAF79138FAD1FAB0" box="[1310,1399,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="10" unit="mm" value="4.9">4.9 mm</quantity>
|
|||
|
|
(MNHN IU 2013-759) is still immature, with the abdomen triangular in shape. An adult female, 10.2 ×
|
|||
|
|
<quantity id="F043F2E4B27DFFADFF5491C3FEB0FA76" box="[189,278,1445,1471]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="10" unit="mm" value="7.8">7.8 mm</quantity>
|
|||
|
|
(pleopods fully setose, ZRC, ex MNHN IU 2013-1245), has an ovate abdomen that covers about half the surface of the thoracic sternum. The vulvae are small, positioned on the anterior half of sternite 6 submedially and each has a distinct opercular cover in the form of a narrow plate (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFB4D918DFAA7F9CF" box="[1188,1281,1515,1542]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[189,1399,436,1890]" captionTargetId="figure@7.[189,1399,436,1890]" captionTargetPageId="7" captionText="Fig. 4. Actaea grimaldii sp. nov., paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245), Papua New Guinea. A. Dorsal view of carapace. B. Thoracic sternum showing vulvae. C. Right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.3785528" httpUri="https://zenodo.org/record/3785528/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
|
|||
|
|
). There is hardly any variation in the diagnostic characters, although some of the specimens (e.g., ZRC, ex MNHN IU 2013-1245) are twice the size of the smaller crabs (e.g., ZRC, ex MNHN IU 2013-920).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27DFFADFF549212FB17F930" blockId="9.[189,1399,1652,1785]" pageId="9" pageNumber="10">
|
|||
|
|
The long, simple setae on the ambulatory legs are easily broken, and while obvious in fresh specimens (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFF2F92F1FEAEF978" box="[198,264,1687,1713]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
|||
|
|
), they are often lost in preserved material. The
|
|||
|
|
<typeStatus id="E800E1A3B27DFFADFCC592F1FC35F978" box="[812,915,1687,1713]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
|||
|
|
male has a developed G1 (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFB2092F1FAADF97B" box="[1225,1291,1687,1714]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
|||
|
|
D–F) but does not appear to be fully mature. While the setae on the subdistal part of the G1 are long (
|
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<figureCitation id="AF804384B27DFFADFB0692DDFA95F91F" box="[1263,1331,1723,1750]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[213,1389,376,1821]" captionTargetId="figure@8.[189,1398,360,1854]" captionTargetPageId="8" captionText="Fig. 5. Actaea grimaldii sp. nov., holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921), Papua New Guinea. A. Right third maxilliped (setae not drawn). B. Anterior thoracic sternum (sternites 1–4) and telson. C. Abdominal somites 1–6. D. Ventral view of left G1. E. Ventral view of distal part of left G1. F. Dorsal view of distal part of left G1. G. Left G2. Scale bars: A, D, G = 0.5 mm; B–C = 1.0 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785530" httpUri="https://zenodo.org/record/3785530/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
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E–F), they are simple and not plumose as is typical for actaeine gonopods (cf.
|
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<bibRefCitation id="532A22F0B27DFFADFBE592B8FB02F930" author="Guinot D." box="[1036,1188,1758,1785]" pageId="9" pageNumber="10" refId="ref6661" refString="Guinot D. 1976. Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La superfamille des Bellioidea et trois sous-familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum national d'Histoire naturelle, Serie A 97, Museum national d'Histoire naturelle, Paris." type="journal volume" year="1976">Guinot 1976</bibRefCitation>
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).
|
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</paragraph>
|
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<paragraph id="37045F01B27DFFADFF039346FEE4F8F3" blockId="9.[189,1399,1824,2027]" box="[234,322,1824,1850]" pageId="9" pageNumber="10">
|
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|
|
<emphasis id="05CF8313B27DFFADFF039346FEE4F8F3" bold="true" box="[234,322,1824,1850]" pageId="9" pageNumber="10">Colour</emphasis>
|
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|
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</paragraph>
|
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|
|
<paragraph id="37045F01B27DFFADFF549322FBD2F822" blockId="9.[189,1399,1824,2027]" pageId="9" pageNumber="10">
|
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|
|
In life (
|
|||
|
|
<figureCitation id="AF804384B27DFFADFEFD9325FEFEF894" box="[276,344,1859,1885]" captionStart="Fig" captionStartId="3.[189,232,1860,1886]" captionTargetBox="[189,1399,399,1819]" captionTargetId="figure@3.[189,1399,399,1819]" captionTargetPageId="3" captionText="Fig. 1. Actaea grimaldii sp. nov., Papua New Guinea, colours in life. A–B. Paratype, ♂, 5.1 × 3.8 mm (ZRC, ex MNHN IU 2013-920). C. Holotype, ♂, 6.2 × 4.4 mm (MNHN IU 2013-921). D. Paratype, ♀, 10.2 × 7.8 mm (ZRC, ex MNHN IU 2013-1245). E. Paratype, ♀, 8.5 × 6.2 mm (MNHN IU 2013-222). F. Paratype, ♀, 6.7 × 4.9 mm (MNHN IU 2013-759). G–H. Paratype, ♀, 6.4 × 4.9 mm (ZRC, ex MNHN IU 2013-1255). A–G, dorsal views of habitus; B, ventral view of habitus; H, frontal view of habitus." figureDoi="http://doi.org/10.5281/zenodo.3785522" httpUri="https://zenodo.org/record/3785522/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
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|
), the colour is a striking bright orange to red across the median part of the carapace, with the lateral regions white. There may be patches of white on the gastric and adjacent regions in smaller specimens. The legs are banded red and white, and the chelipeds are red and white, with parts yellowishwhite in colour. The fingers of the chela are reddish-orange basally and white at the tips. The ventral surfaces are white, but abdominal somites 1 and 2 may have large patches of red.
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="7FA10C8AB27EFFAEFF54956DFE0FFEBD" pageId="10" pageNumber="11" type="distribution">
|
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|
|
<paragraph id="37045F01B27EFFAEFF54956DFEF0FEEC" blockId="10.[189,342,267,293]" box="[189,342,267,293]" pageId="10" pageNumber="11">
|
|||
|
|
<heading id="6C4CE86DB27EFFAEFF54956DFEF0FEEC" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
|||
|
|
<emphasis id="05CF8313B27EFFAEFF54956DFEF0FEEC" bold="true" box="[189,342,267,293]" pageId="10" pageNumber="11">Distribution</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="37045F01B27EFFAEFF549551FE0FFEBD" blockId="10.[189,1398,311,373]" pageId="10" pageNumber="11">
|
|||
|
|
This species is known for certain from the
|
|||
|
|
<collectingCountry id="4FAC1F91B27EFFAEFD389551FCE5FE98" box="[721,835,311,337]" name="Indonesia" pageId="10" pageNumber="11">Moluccas</collectingCountry>
|
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|
|
and
|
|||
|
|
<collectingCountry id="4FAC1F91B27EFFAEFC6A955EFBD7FE98" box="[899,1137,311,338]" name="Papua New Guinea" pageId="10" pageNumber="11">Papua New Guinea</collectingCountry>
|
|||
|
|
, and is probably also present in
|
|||
|
|
<collectingCountry id="4FAC1F91B27EFFAEFEDF953CFE03FEBD" box="[310,421,346,372]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|