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<document id="AE3444215DCE79B18862DDCCAF42C328" ID-DOI="10.5281/zenodo.175948" ID-GBIF-Dataset="02bb7dce-7590-4f45-8aa9-c6280f2e8b36" ID-ISSN="1175-5326" ID-Zenodo-Dep="175948" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459825277757" checkinUser="plazi" docAuthor="Liu, Xingyue, Hayashi, Fumio &amp; Yang, Ding" docDate="2007" docId="03CF87E5FFFCFFEAFF46AD1AFCDDFE13" docLanguage="en" docName="zt01439p046.pdf" docOrigin="Zootaxa 1439" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Protohermes yangi Liu, Hayashi &amp; Yang, 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFF6FF9DFFD4FFC0FFD1A94DFFEDFFD9" masterDocTitle="Systematics of the Protohermes costalis species-group (Megaloptera: Corydalidae)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="41" updateTime="1698222167997" updateUser="plazi">
<mods:mods id="C12D262E4FC8A6C01089EAA074B5837C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EEEDFD2E7AE3CDF85D737BFE250E61FE">
<mods:title id="F90FF5CC6DE16547C67B356400F341AE">Systematics of the Protohermes costalis species-group (Megaloptera: Corydalidae)</mods:title>
</mods:titleInfo>
<mods:name id="F41AC220A0416985FC7A5E029BC89D08" type="personal">
<mods:role id="1E65E0671B9051B9935BEA14C7126920">
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<mods:namePart id="94BA082AD21807D564CBEA1B0E6FBFDB">Liu, Xingyue</mods:namePart>
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<mods:namePart id="7C656BC5EC1315D0855DC0CD3322FA46">Hayashi, Fumio</mods:namePart>
</mods:name>
<mods:name id="879742436CFDAEAFB9AE6D5AA62C04D5" type="personal">
<mods:role id="60F6E3EAA1F07A261C0989F29D504058">
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<mods:namePart id="D1CB6B1FD61164F9B65E905EE1D0C658">Yang, Ding</mods:namePart>
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<mods:typeOfResource id="44F854B7020ACD4EAF3DF81B205CB36B">text</mods:typeOfResource>
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<mods:title id="FA0CCF88BC5B3A7EC294A1A6EA9DE08C">Zootaxa</mods:title>
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<mods:part id="21A4DE1FA1B3ADCD45F88C0FE5C1EA67">
<mods:date id="25340D5ABDA50B5E0782CF2B56CE3C33">2007</mods:date>
<mods:detail id="E33E09DE1EFBB8AE02F0C02DD6301192" type="volume">
<mods:number id="2F85D6E833AC521A46DAF0698E1703B2">1439</mods:number>
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<mods:classification id="08AE7A9B0C9A1E3B8C97CD44DD3EC930">journal article</mods:classification>
<mods:identifier id="BE6D513933D2D47D28E305D5980545B9" type="DOI">10.5281/zenodo.175948</mods:identifier>
<mods:identifier id="8A9D788306C5BBA8040755AD10146BF6" type="GBIF-Dataset">02bb7dce-7590-4f45-8aa9-c6280f2e8b36</mods:identifier>
<mods:identifier id="B1D69999DD955345860C05408A89571D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="73F0873A2406C120293B60A4E693D247" type="Zenodo-Dep">175948</mods:identifier>
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<treatment id="03CF87E5FFFCFFEAFF46AD1AFCDDFE13" ID-DOI="http://doi.org/10.5281/zenodo.6248253" ID-GBIF-Taxon="119356602" ID-Zenodo-Dep="6248253" LSID="urn:lsid:plazi:treatment:03CF87E5FFFCFFEAFF46AD1AFCDDFE13" httpUri="http://treatment.plazi.org/id/03CF87E5FFFCFFEAFF46AD1AFCDDFE13" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C37C6578FFFCFFE8FF46AD1AFE93FB4D" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BD936F3FFFCFFE8FF46AD1AFE0DFBA8" blockId="40.[151,480,1111,1172]" box="[151,480,1111,1137]" pageId="40" pageNumber="41">
<heading id="D091819FFFFCFFE8FF46AD1AFE0DFBA8" bold="true" box="[151,480,1111,1137]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B912EAE1FFFCFFE8FF46AD1AFE0DFBA8" bold="true" box="[151,480,1111,1137]" pageId="40" pageNumber="41">
<taxonomicName id="4C664D70FFFCFFE8FF46AD1AFE91FBA8" ID-CoL="4N5K6" box="[151,380,1111,1137]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B912EAE1FFFCFFE8FF46AD1AFE91FBA8" bold="true" box="[151,380,1111,1137]" italics="true" pageId="40" pageNumber="41">Protohermes yangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221579AFFFCFFE8FE52AD1AFE0DFBA8" box="[387,480,1111,1137]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936F3FFFCFFE8FF46AD37FE93FB4D" blockId="40.[151,480,1111,1172]" box="[151,382,1146,1172]" pageId="40" pageNumber="41">
(
<figureCitation id="135D2A76FFFCFFE8FF4EAD37FEEFFB4D" box="[159,258,1146,1172]" captionStart="FIGURES 16 – 23" captionStartId="14.[151,269,1433,1457]" captionTargetBox="[156,1426,190,1415]" captionTargetId="figure@14.[760,1432,796,1058]" captionTargetPageId="14" captionText="FIGURES 16 – 23. Habitus of the species in the P. costalis group. 16. P. niger Yang &amp; Yang (Holotype), male. 17. P. o r i - entalis sp. nov. (Holotype), male. 18. P. similis Yang &amp; Yang (Holotype), male. 19. P. s i n e n s i s Yang &amp; Yang (Holotype), male. 20. P. stigmosus sp. nov. (Holotype), male. 21. P. triangulatus sp. nov. (Holotype), male. 22. P. y a n g i sp. nov. (Holotype), male. 23. P. yunnanensis Yang &amp; Yang (Holotype), male. Scale bars = 5 mm." httpUri="https://zenodo.org/record/175952/files/figure.png" pageId="40" pageNumber="41">Figs. 22</figureCitation>
,
<figureCitation id="135D2A76FFFCFFE8FEDFAD37FE9BFB4D" box="[270,374,1146,1172]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="40" pageNumber="41">135–141</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C6578FFFCFFE9FF46AD86FB83F95A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8BD936F3FFFCFFE8FF46AD86FD12FAED" blockId="40.[151,1437,1226,2017]" pageId="40" pageNumber="41">
<emphasis id="B912EAE1FFFCFFE8FF46AD86FEE3FB3D" box="[151,270,1227,1252]" italics="true" pageId="40" pageNumber="41">Diagnosis</emphasis>
. Head with two blackish markings on each side of vertex; pronotum bivittate black; wings grayish brown with distinct whitish markings; male ninth sternum with deep V-shaped posterior incision; male tenth tergum strongly and roundly produced distolaterally.
</paragraph>
<paragraph id="8BD936F3FFFCFFE8FF16AC0FFC12FA5D" blockId="40.[151,1437,1226,2017]" pageId="40" pageNumber="41">
Body length
<quantity id="4C9E9B16FFFCFFE8FEB2AC0FFE09FA85" box="[355,484,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="40" pageNumber="41" unit="mm" value="32.5" valueMax="34.0" valueMin="31.0">31–34 mm</quantity>
(male) and
<quantity id="4C9E9B16FFFCFFE8FDA9AC0FFD14FA85" box="[632,761,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="40" pageNumber="41" unit="mm" value="37.5" valueMax="40.0" valueMin="35.0">35–40 mm</quantity>
(female); forewing length
<quantity id="4C9E9B16FFFCFFE8FBEFAC0FFB52FA85" box="[1086,1215,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="40" pageNumber="41" unit="mm" value="38.5" valueMax="40.0" valueMin="37.0">37–40 mm</quantity>
(male) and
<quantity id="4C9E9B16FFFCFFE8FA82AC0FFF28FA5D" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.5" metricValueMin="3.8" pageId="40" pageNumber="41" unit="mm" value="41.5" valueMax="45.0" valueMin="38.0">38–45 mm</quantity>
(female), hindwing length
<quantity id="4C9E9B16FFFCFFE8FDD5AC27FD6EFA5D" box="[516,643,1386,1412]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="40" pageNumber="41" unit="mm" value="34.0" valueMax="35.0" valueMin="33.0">33–35 mm</quantity>
(male) and
<quantity id="4C9E9B16FFFCFFE8FCDEAC27FC63FA5D" box="[783,910,1386,1412]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="40" pageNumber="41" unit="mm" value="38.0" valueMax="41.0" valueMin="35.0">35–41 mm</quantity>
(female).
</paragraph>
<paragraph id="8BD936F3FFFCFFE8FF16ACDFFF34F9AD" blockId="40.[151,1437,1226,2017]" pageId="40" pageNumber="41">
Head (
<figureCitation id="135D2A76FFFCFFE8FEC7ACDFFE91FA75" box="[278,380,1426,1452]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="40" pageNumber="41">Fig. 135</figureCitation>
) yellowish brown without post-ocular spine; vertex with two blackish markings on each side, lateral marking band-like, medial marking punctuate. Compound eyes grayish brown; ocelli yellow, darkly margined medially. Antennae black, scapes and pedicel brown with tip and basal half pale yellow. Mouthparts yellowish brown; distal three joints of maxillary palpus and distal three joints of labial palpus brown, mandible with distal half dark. Occiput yellowish brown with one pair of blackish markings on lateral sides.
</paragraph>
<paragraph id="8BD936F3FFFCFFE8FF16AFCFFB13F805" blockId="40.[151,1437,1226,2017]" pageId="40" pageNumber="41">
Prothorax (
<figureCitation id="135D2A76FFFCFFE8FE9DAFCFFE5FF945" box="[332,434,1666,1692]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="40" pageNumber="41">Fig. 135</figureCitation>
) yellowish brown; pronotum with one blackish vitta on each side, vittae sometimes narrowed medially. Meso- and metathorax pale brown, and mesonotum dorsally with pale yellow center. Thoracic pilosity yellow, much longer on meso- and metathorax. Legs pale yellow to brown with dense yellowish short setae; wide apical portion of tibiae and tarsi black; tarsal claws reddish brown. Forewings (
<figureCitation id="135D2A76FFFCFFE8FB2BAFB7FAA2F8CD" box="[1274,1359,1786,1812]" captionStart="FIGURES 16 – 23" captionStartId="14.[151,269,1433,1457]" captionTargetBox="[156,1426,190,1415]" captionTargetId="figure@14.[760,1432,796,1058]" captionTargetPageId="14" captionText="FIGURES 16 – 23. Habitus of the species in the P. costalis group. 16. P. niger Yang &amp; Yang (Holotype), male. 17. P. o r i - entalis sp. nov. (Holotype), male. 18. P. similis Yang &amp; Yang (Holotype), male. 19. P. s i n e n s i s Yang &amp; Yang (Holotype), male. 20. P. stigmosus sp. nov. (Holotype), male. 21. P. triangulatus sp. nov. (Holotype), male. 22. P. y a n g i sp. nov. (Holotype), male. 23. P. yunnanensis Yang &amp; Yang (Holotype), male. Scale bars = 5 mm." httpUri="https://zenodo.org/record/175952/files/figure.png" pageId="40" pageNumber="41">Fig. 22</figureCitation>
) grayish brown, with 1 subtriangular whitish marking at base, 2 large and 5–6 small whitish markings at middle and 1 extremely enlarged round whitish marking on apical 1/3; costal cellules with distinct brownish stripes. Hindwings pale brown with basal half mostly hyaline, with 1 round whitish marking on apical 1/3. Veins yellowish brown, except veins on apical half pale brown. Rs 8 to 10-branched, last branch bifurcate or trifurcate; 9–11 crossveins between R1 and Rs; M1+2 6 to 7-branched, M3+4 2 to 3-branched; 1A 3-branched.
</paragraph>
<caption id="DF19667BFFFDFFE9FF46AD6DFB2FFBA4" httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42" targetBox="[158,1417,203,1020]" targetPageId="41">
<paragraph id="8BD936F3FFFDFFE9FF46AD6DFB2FFBA4" blockId="41.[151,1436,1056,1149]" pageId="41" pageNumber="42">
<emphasis id="B912EAE1FFFDFFE9FF46AD6DFE9AFBE1" bold="true" box="[151,375,1056,1080]" pageId="41" pageNumber="42">FIGURES 135–141.</emphasis>
<taxonomicName id="4C664D70FFFDFFE9FEACAD6CFDAAFBE1" box="[381,583,1057,1080]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B912EAE1FFFDFFE9FEACAD6CFDAAFBE1" box="[381,583,1057,1080]" italics="true" pageId="41" pageNumber="42">Protohermes yangi</emphasis>
</taxonomicName>
,
<emphasis id="B912EAE1FFFDFFE9FD85AD6DFD44FBE1" bold="true" box="[596,681,1056,1080]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A221579AFFFDFFE9FD85AD6DFD44FBE1" box="[596,681,1056,1080]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BD936F3FFFDFFE9FF16ADFCFB83F95A" blockId="41.[151,1437,1201,2027]" pageId="41" pageNumber="42">
Abdomen dark with yellowish short setae. Male ninth tergum (
<figureCitation id="135D2A76FFFDFFE9FC13ADFCFBC4FB12" box="[962,1065,1201,1227]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Fig. 136</figureCitation>
) subquadrate, with trapezoidal anterior margin and shallowly incised posterior margin. Ninth sternum (
<figureCitation id="135D2A76FFFDFFE9FC20AD94FBBAFB2A" box="[1009,1111,1241,1267]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Fig. 137</figureCitation>
) broad with straight lateral margin; posterior margin with deep V-shaped incision, forming two large triangular processes; central portion inflated. Ninth gonostylus (
<figureCitation id="135D2A76FFFDFFE9FE09AC64FDD1FA9A" box="[472,572,1321,1347]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Fig. 138</figureCitation>
) unguiform and strongly curved dorsomedially. Tenth tergum (
<figureCitation id="135D2A76FFFDFFE9FACEAC64FF29FAB2" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Figs. 136– 137</figureCitation>
) short and subcylindrical, ventrally with one feebly developed tufted tubercle at posteromedial corner; posterolateral corner distinctly and roundly produced; tip densely setose and sometimes slightly incised. Tenth sternum (
<figureCitation id="135D2A76FFFDFFE9FED6ACECFE81FA62" box="[263,364,1441,1467]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Fig. 139</figureCitation>
) arched, anteriorly with one subtriangular median process; posterior incision small and Vshaped; lateral lobes digitiform and slightly incurved. Female eighth sternum (
<figureCitation id="135D2A76FFFDFFE9FBFDAC84FB3BFA3A" box="[1068,1238,1481,1507]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Figs. 140–141</figureCitation>
) strongly sclerotized and subtriangular in lateral view, posterior margin elevated without incision, forming a pair of triangular processes; ninth segment (
<figureCitation id="135D2A76FFFDFFE9FE07AF54FDD1F9EA" box="[470,572,1561,1587]" captionStart="FIGURES 135 – 141" captionStartId="41.[151,269,1056,1080]" captionTargetBox="[158,1417,203,1020]" captionTargetId="figure@41.[151,1436,194,1032]" captionTargetPageId="41" captionText="FIGURES 135 – 141. Protohermes yangi, sp. nov. 135. Male head and pronotum, dorsal view. 136. Male genitalia, dorsal view. 137. Male genitalia, ventral view. 138. Male ninth gonostylus, caudal view. 139. Male tenth sternum, ventral view. 140. Female genitalia, lateral view. 141. Female eighth sternum, ventral view. Scale lines = 1.0 mm." httpUri="https://zenodo.org/record/175969/files/figure.png" pageId="41" pageNumber="42">Fig. 140</figureCitation>
) laterally inflated forming a pair of small suboval sac-like lobes; ninth gonocoxite valvate and membranous with one digitiform process at tip; tenth tergum short, with posterior margin medially incised, leaving one subtriangular upper and one round lower lobe.
</paragraph>
</subSubSection>
<subSubSection id="C37C6578FFFDFFE9FF16AFDCFB30F842" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BD936F3FFFDFFE9FF16AFDCFB30F842" blockId="41.[151,1437,1201,2027]" pageId="41" pageNumber="42">
<emphasis id="B912EAE1FFFDFFE9FF16AFDCFE92F973" box="[199,383,1681,1706]" italics="true" pageId="41" pageNumber="42">
<typeStatus id="54DD8851FFFDFFE9FF16AFDCFF13F973" box="[199,254,1681,1706]" pageId="41" pageNumber="42">Type</typeStatus>
materials.
</emphasis>
<typeStatus id="54DD8851FFFDFFE9FE5BAFDCFE1AF972" box="[394,503,1681,1707]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="F3717663FFFDFFE9FDF9AFDCFD59F972" box="[552,692,1681,1707]" name="Vietnam" pageId="41" pageNumber="42">VIETNAM</collectingCountry>
: Mt. Pia Oac,
<date id="FFD81033FFFDFFE9FCB8AFDCFC09F972" box="[873,996,1681,1707]" pageId="41" pageNumber="42" value="1998-05-14">14.V.1998</date>
, F. Hayashi (
<collectionCode id="ED77AE36FFFDFFE9FB56AFDCFB26F972" LSID="urn:lsid:biocol.org:col:33074" box="[1159,1227,1681,1707]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
).
<typeStatus id="54DD8851FFFDFFE9FB32AFDCFAB4F972" box="[1251,1369,1681,1707]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
: 3 ɗ,
<collectingCountry id="F3717663FFFDFFE9FF46AFF4FF14F90A" box="[151,249,1721,1747]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guangxi, Tianlin, Langping,
<quantity id="4C9E9B16FFFDFFE9FD8BAFF4FD5AF90A" box="[602,695,1721,1747]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="41" pageNumber="42" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FFD81033FFFDFFE9FD12AFF4FCD1F90A" box="[707,828,1721,1747]" pageId="41" pageNumber="42" value="2002-06-02">2.VI.2002</date>
, Jianwen Liu (
<collectionCode id="ED77AE36FFFDFFE9FC3BAFF4FBC3F90A" LSID="urn:lsid:biocol.org:col:33074" box="[1002,1070,1721,1747]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); 2 ɗ,
<collectingCountry id="F3717663FFFDFFE9FBACAFF4FB33F90A" box="[1149,1246,1721,1747]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guangxi, Tianlin, Langping,
<quantity id="4C9E9B16FFFDFFE9FE92AFACFE4DF922" box="[323,416,1761,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="41" pageNumber="42" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FFD81033FFFDFFE9FE7AAFACFDC9F922" box="[427,548,1761,1787]" pageId="41" pageNumber="42" value="2002-06-01">1.VI.2002</date>
, Jianwen Liu (
<collectionCode id="ED77AE36FFFDFFE9FD00AFACFCF8F922" LSID="urn:lsid:biocol.org:col:33074" box="[721,789,1761,1787]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); ɗ,
<collectingCountry id="F3717663FFFDFFE9FC9DAFACFC43F922" box="[844,942,1761,1787]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guangxi, Linaoshan,
<quantity id="4C9E9B16FFFDFFE9FB64AFACFAFCF922" box="[1205,1297,1761,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FFD81033FFFDFFE9FACDAFACFA7AF922" box="[1308,1431,1761,1787]" pageId="41" pageNumber="42" value="2002-05-28">28.V.2002</date>
, Jianwen Liu (
<collectionCode id="ED77AE36FFFDFFE9FEE7AE44FE97F8FA" LSID="urn:lsid:biocol.org:col:33074" box="[310,378,1801,1827]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); ɗ,
<collectingCountry id="F3717663FFFDFFE9FE7EAE44FDFDF8FA" box="[431,528,1801,1827]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guangxi, Linaoshan,
<quantity id="4C9E9B16FFFDFFE9FCC7AE44FC9CF8FA" box="[790,881,1801,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FFD81033FFFDFFE9FCACAE44FC1AF8FA" box="[893,1015,1801,1827]" pageId="41" pageNumber="42" value="2002-05-28">28.V.2002</date>
, Guofang Jiang (
<collectionCode id="ED77AE36FFFDFFE9FB6CAE44FAECF8FA" LSID="urn:lsid:biocol.org:col:33074" box="[1213,1281,1801,1827]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); ɗ,
<collectingCountry id="F3717663FFFDFFE9FAE6AE44FA75F8FA" box="[1335,1432,1801,1827]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guangxi, Tianlin, Laoshan Forestry Center,
<quantity id="4C9E9B16FFFDFFE9FD11AE7CFCCEF892" box="[704,803,1841,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FFD81033FFFDFFE9FCE4AE7CFC42F892" box="[821,943,1841,1867]" pageId="41" pageNumber="42" value="2002-06-05">5.VI.2002</date>
, Guofang Jiang (
<collectionCode id="ED77AE36FFFDFFE9FB5DAE7CFB3DF892" LSID="urn:lsid:biocol.org:col:33074" box="[1164,1232,1841,1867]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); ɗ Ψ,
<collectingCountry id="F3717663FFFDFFE9FAE6AE7CFA74F892" box="[1335,1433,1841,1867]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Gaungxi, Tianlin, Yaojiawan,
<quantity id="4C9E9B16FFFDFFE9FE2AAE14FD4FF8AA" box="[507,674,1881,1907]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="41" pageNumber="42" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200–1400 m</quantity>
,
<date id="FFD81033FFFDFFE9FD7EAE14FCC5F8AA" box="[687,808,1881,1907]" pageId="41" pageNumber="42" value="2002-06-01">1.VI.2002</date>
, Xiujuan Yang (HBU); Ψ,
<collectingCountry id="F3717663FFFDFFE9FBB6AE14FB2AF8AA" box="[1127,1223,1881,1907]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
, Guangxi, Tian-e, Buliuhe,
<date id="FFD81033FFFDFFE9FED3AECCFE62F842" box="[258,399,1921,1947]" pageId="41" pageNumber="42" value="2002-08-06">6.VIII.2002</date>
, Ding Yang (
<collectionCode id="ED77AE36FFFDFFE9FDF8AECCFD80F842" LSID="urn:lsid:biocol.org:col:33074" box="[553,621,1921,1947]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
); Ψ,
<collectingCountry id="F3717663FFFDFFE9FD75AECCFCEBF842" box="[676,774,1921,1947]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: Guizhou, Pingtang, Yenanzhou (
<collectionCode id="ED77AE36FFFDFFE9FB5CAECCFB3CF842" LSID="urn:lsid:biocol.org:col:33074" box="[1165,1233,1921,1947]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="41" pageNumber="42">CAU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6578FFFDFFE9FF16AEE4FB73F832" pageId="41" pageNumber="42" type="etymology">
<paragraph id="8BD936F3FFFDFFE9FF16AEE4FB73F832" blockId="41.[151,1437,1201,2027]" pageId="41" pageNumber="42">
<emphasis id="B912EAE1FFFDFFE9FF16AEE4FEAEF81B" box="[199,323,1961,1986]" italics="true" pageId="41" pageNumber="42">Etymology</emphasis>
. The specific epithet ‘
<taxonomicName id="4C664D70FFFDFFE9FD99AEE4FD64F81B" box="[584,649,1961,1986]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="B912EAE1FFFDFFE9FD99AEE4FD64F81B" box="[584,649,1961,1986]" italics="true" pageId="41" pageNumber="42">yangi</emphasis>
</taxonomicName>
’ refers to the famous entomologist Prof. Chikun Yang, who passed away in 2006, for his great contributions to the systematics of the Chinese
<taxonomicName id="4C664D70FFFDFFE9FBD3AE9CFB77F832" box="[1026,1178,2001,2027]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="order">Megaloptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C6578FFFEFFEAFF16A9D5FC2DFF6B" box="[199,960,152,178]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BD936F3FFFEFFEAFF16A9D5FC2DFF6B" blockId="42.[151,1437,152,458]" box="[199,960,152,178]" pageId="42" pageNumber="43">
<emphasis id="B912EAE1FFFEFFEAFF16A9D5FEBBFF68" box="[199,342,152,177]" italics="true" pageId="42" pageNumber="43">Distribution</emphasis>
.
<collectingCountry id="F3717663FFFEFFEAFEB4A9D5FE46FF6B" box="[357,427,152,178]" name="China" pageId="42" pageNumber="43">China</collectingCountry>
(Guangxi, Guizhou);
<collectingCountry id="F3717663FFFEFFEAFD7CA9D5FCFDFF6B" box="[685,784,152,178]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
(Mt. Pia Oac).
</paragraph>
</subSubSection>
<subSubSection id="C37C6578FFFEFFEAFF16A98DFCDDFE13" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BD936F3FFFEFFEAFF16A98DFCDDFE13" blockId="42.[151,1437,152,458]" pageId="42" pageNumber="43">
<emphasis id="B912EAE1FFFEFFEAFF16A98DFEC0FF00" box="[199,301,192,217]" italics="true" pageId="42" pageNumber="43">Remarks</emphasis>
. This new species might have a close affinity with
<taxonomicName id="4C664D70FFFEFFEAFC4DA98DFBD2FF00" box="[924,1087,192,217]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hunanensis">
<emphasis id="B912EAE1FFFEFFEAFC4DA98DFBD2FF00" box="[924,1087,192,217]" italics="true" pageId="42" pageNumber="43">P. hunanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D70FFFEFFEAFBADA98DFAF6FF00" box="[1148,1307,192,217]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="B912EAE1FFFEFFEAFBADA98DFAF6FF00" box="[1148,1307,192,217]" italics="true" pageId="42" pageNumber="43">P. fujianensis</emphasis>
</taxonomicName>
due to the similar wing pattern with extremely enlarged round whitish marking on the apical 1/3 of both wings. However, the male ninth sternum of the new species bears a deep V-shaped posterior incision. Alternatively, the posterior incision of the male ninth sternum is trapezoidal in both
<taxonomicName id="4C664D70FFFEFFEAFC7AA875FBA0FE88" box="[939,1101,312,337]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hunanensis">
<emphasis id="B912EAE1FFFEFFEAFC7AA875FBA0FE88" box="[939,1101,312,337]" italics="true" pageId="42" pageNumber="43">P. hunanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D70FFFEFFEAFB59A875FAC8FE88" box="[1160,1317,312,337]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="B912EAE1FFFEFFEAFB59A875FAC8FE88" box="[1160,1317,312,337]" italics="true" pageId="42" pageNumber="43">P. fujianensis</emphasis>
</taxonomicName>
. The new species also somewhat resembles
<taxonomicName id="4C664D70FFFEFFEAFDF5A82DFD22FEA0" box="[548,719,352,377]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="triangulatus">
<emphasis id="B912EAE1FFFEFFEAFDF5A82DFD22FEA0" box="[548,719,352,377]" italics="true" pageId="42" pageNumber="43">P. triangulatus</emphasis>
</taxonomicName>
in having the similar V-shaped posterior incision of the male ninth sternum, but it can be distinguished by the male tenth tergum ventrally with feebly developed tubercle. In
<taxonomicName id="4C664D70FFFEFFEAFF67A8FDFE8FFE10" box="[182,354,432,457]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="triangulatus">
<emphasis id="B912EAE1FFFEFFEAFF67A8FDFE8FFE10" box="[182,354,432,457]" italics="true" pageId="42" pageNumber="43">P. triangulatus</emphasis>
</taxonomicName>
, the ventral tubercle is well developed.
</paragraph>
</subSubSection>
</treatment>
</document>
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