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<document id="0A721406C95BD87C708AC12690D9B3CE" ID-CLB-Dataset="172795" ID-DOI="10.5281/zenodo.5039032" ID-GBIF-Dataset="7eb79be8-aa10-4707-a05e-e856090182de" ID-ISSN="0253-116X" ID-Zenodo-Dep="5039032" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624912051484" checkinUser="carolina" docAuthor="Assing, Volker" docDate="2021" docId="6B178795FF8A7B6EFF76FD5FFD6B467A" docLanguage="en" docName="LinzerbiolBeitr.52.2.891-931.pdf" docOrigin="Linzer biologische Beiträge 52 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Cephennium (Macroderus) cazorlae Assing 2021, nov.sp." docType="treatment" docVersion="14" lastPageNumber="928" masterDocId="972EFFEDFFAF7B4BFFE1FFD4FFB54350" masterDocTitle="On the Cephennium fauna of the Iberian Peninsula and the Atlantic Islands (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="931" masterPageNumber="891" pageNumber="928" updateTime="1698978031781" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5FC2446FA347B34E3DA553A808EED2BC">On the Cephennium fauna of the Iberian Peninsula and the Atlantic Islands (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="6B178795FF8A7B6EFF76FD5FFD6B467A" ID-DOI="http://doi.org/10.5281/zenodo.7254906" ID-GBIF-Taxon="203398224" ID-Zenodo-Dep="7254906" LSID="urn:lsid:plazi:treatment:6B178795FF8A7B6EFF76FD5FFD6B467A" httpUri="http://treatment.plazi.org/id/6B178795FF8A7B6EFF76FD5FFD6B467A" lastPageNumber="928" pageId="37" pageNumber="928">
<subSubSection id="ABA46508FF8A7B6EFF76FD5FFCE941F2" box="[151,860,651,674]" pageId="37" pageNumber="928" type="nomenclature">
<paragraph id="E3013683FF8A7B6EFF76FD5FFCE941F2" blockId="37.[151,860,651,674]" box="[151,860,651,674]" pageId="37" pageNumber="928">
<heading id="B84981EFFF8A7B6EFF76FD5FFCE941F2" box="[151,860,651,674]" fontSize="8" level="2" pageId="37" pageNumber="928" reason="2">
<emphasis id="D1CAEA91FF8A7B6EFF76FD5FFDD141F1" bold="true" box="[151,612,651,673]" pageId="37" pageNumber="928">
<taxonomicName id="24BE4D00FF8A7B6EFF76FD5FFDA741F1" ID-CoL="8XWW8" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[151,530,651,673]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="928" phylum="Arthropoda" rank="species" species="cazorlae" status="sp. nov." subGenus="Macroderus">
<emphasis id="D1CAEA91FF8A7B6EFF76FD5FFEAB41F1" bold="true" box="[151,286,651,673]" italics="true" pageId="37" pageNumber="928">Cephennium</emphasis>
(
<emphasis id="D1CAEA91FF8A7B6EFECCFD5FFE1941F1" bold="true" box="[301,428,651,673]" italics="true" pageId="37" pageNumber="928">Macroderus</emphasis>
)
<emphasis id="D1CAEA91FF8A7B6EFE5AFD5FFDA741F1" bold="true" box="[443,530,651,673]" italics="true" pageId="37" pageNumber="928">cazorlae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAF957EAFF8A7B6EFDF9FD58FDD141F1" box="[536,612,652,673]" pageId="37" pageNumber="928" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7B852A06FF8A7B6EFD93FD58FD7541F1" box="[626,704,652,674]" captionStart="Figs 17-20" captionStartId="15.[151,191,1499,1521]" captionTargetBox="[160,1024,206,1472]" captionTargetId="figure-43@15.[152,1059,202,1487]" captionTargetPageId="15" captionText="Figs 17-20: Male habitus of species of the subgenus Macroderus in dorsal view: (17) Cephennium segurae; (18) C. nevadae; (19) C. cazorlae; (20) C. malagae. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5039046" httpUri="https://zenodo.org/record/5039046/files/figure.png" pageId="37" pageNumber="928">Figs 19</figureCitation>
,
<figureCitation id="7B852A06FF8A7B6EFD2AFD58FCBD41F2" box="[715,776,652,674]" captionStart="Figs 72-78" captionStartId="23.[151,191,1499,1521]" captionTargetBox="[156,1059,219,1480]" captionTargetId="figure-47@23.[152,1059,200,1485]" captionTargetPageId="23" captionText="Figs 72-78: Cephennium nevadae (72-74), C. carzolae (75-77), and C. malagae (78): (72-73, 75- 76) aedeagus in lateral and in ventral view; (74, 77-78) internal structures of aedeagus in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7250379" httpUri="https://zenodo.org/record/7250379/files/figure.png" pageId="37" pageNumber="928">75-77</figureCitation>
,
<figureCitation id="7B852A06FF8A7B6EFCF2FD58FCE041F2" box="[787,853,652,674]" captionStart="Map 3" captionStartId="33.[151,196,985,1006]" captionTargetBox="[164,1060,200,972]" captionTargetId="figure-236@33.[154,1060,200,972]" captionTargetPageId="33" captionText="Map 3: Distributions of Cephennium asturicum (black circles; based on Besuchet (unpubl.) and examined material) and of species of the subgenus Macroderus in the Iberian Peninsula: C. alacantae (black triangles); C. cujonicum (white diamond); C. segurae (black diamond); C. verrens (white circles); C. cazorlae (black square); C. nevadae (black star); C. malagae (white triangles)." figureDoi="http://doi.org/10.5281/zenodo.5039072" httpUri="https://zenodo.org/record/5039072/files/figure.png" pageId="37" pageNumber="928">Map 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABA46508FF8A7B6EFF76FD63FE1F4039" pageId="37" pageNumber="928" type="materials_examined">
<paragraph id="E3013683FF8A7B6EFF76FD63FE4E419D" blockId="37.[151,1059,695,873]" box="[151,507,695,717]" pageId="37" pageNumber="928">
<taxonomicName id="24BE4D00FF8A7B6EFF76FD63FE68419C" authorityName="MULLER &amp; KUNZE" authorityYear="1822" box="[151,477,695,716]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="928" phylum="Arthropoda" rank="genus">
<emphasis id="D1CAEA91FF8A7B6EFF76FD63FEC0419C" box="[151,373,695,716]" italics="true" pageId="37" pageNumber="928">Cephennium foveolatum</emphasis>
BESUCHET
</taxonomicName>
i.l.
</paragraph>
<paragraph id="E3013683FF8A7B6EFF76FD02FD924049" blockId="37.[151,1059,695,873]" pageId="37" pageNumber="928">
Type material:
<materialsCitation id="53D63CDEFF8A7B6EFE46FD02FD924049" ID-GBIF-Occurrence="3946761443" collectingDate="1993-10-06" collectionCode="cAss" collectorName="Wunderle &amp; Cazorla" country="Spain" determinerName="V. Assing" elevation="1200" location="S. da Cazorla" pageId="37" pageNumber="928" specimenCount="1" specimenCount-male="1" stateProvince="Andalusia" typeStatus="holotype">
<typeStatus id="3C058821FF8A7B6EFE46FD02FE4941BC" box="[423,508,726,748]" pageId="37" pageNumber="928" type="holotype">Holotype</typeStatus>
<specimenCount id="F5B8FD0AFF8A7B6EFDE9FD02FDA941BB" box="[520,540,726,747]" pageId="37" pageNumber="928" type="male">♂</specimenCount>
: &quot;E–
<collectingRegion id="217AF861FF8A7B6EFDACFD02FD1F41BC" box="[589,682,726,748]" country="Spain" name="Andalucia" pageId="37" pageNumber="928">Andalusia</collectingRegion>
(J),
<location id="E6616058FF8A7B6EFD00FD02FCDC41BC" LSID="urn:lsid:plazi:treatment:6B178795FF8A7B6EFF76FD5FFD6B467A:E6616058FF8A7B6EFD00FD02FCDC41BC" box="[737,873,726,748]" country="Spain" name="S. da Cazorla" pageId="37" pageNumber="928" stateProvince="Andalusia">S. da Cazorla</location>
<quantity id="24469B66FF8A7B6EFC94FD02FC7641BB" box="[885,963,726,747]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="928" unit="m" value="1200.0">
<elevation id="6893D1B0FF8A7B6EFC94FD02FC7641BB" box="[885,963,726,747]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="928" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, 06.10.93
<collectorName id="4E4B5355FF8A7B6EFF76FD39FF444053" box="[151,241,749,771]" pageId="37" pageNumber="928">Wunderle</collectorName>
/
<collectorName id="4E4B5355FF8A7B6EFEE8FD39FEE74053" box="[265,338,749,771]" pageId="37" pageNumber="928">Cazorla</collectorName>
, Umg. Gualaquivirquelle [sic], Macchia, Kiefer / Holotypus ♂
<originalDetermination id="C0A08E7FFF8A7B6EFC4EFD39FEA94049" pageId="37" pageNumber="928">
<taxonomicName id="24BE4D00FF8A7B6EFC4EFD39FF504048" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="928" phylum="Arthropoda" rank="species" species="cazorlae" status="sp. nov.">
<emphasis id="D1CAEA91FF8A7B6EFC4EFD39FF504048" italics="true" pageId="37" pageNumber="928">Cephennium cazorlae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAF957EAFF8A7B6EFF0AFCD0FEA94049" box="[235,284,772,793]" pageId="37" pageNumber="928" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
det.
<determinerName id="16DDDA47FF8A7B6EFEA9FCD0FE164049" box="[328,419,771,793]" pageId="37" pageNumber="928">V. Assing</determinerName>
2020&quot; (
<collectionCode id="85AFAE46FF8A7B6EFE0AFCD7FDAE4049" box="[491,539,771,793]" pageId="37" pageNumber="928">cAss</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E3013683FF8A7B6EFF76FCF7FE1F4039" blockId="37.[151,1059,695,873]" pageId="37" pageNumber="928">
Additional type material examined by
<collectorName id="4E4B5355FF8A7B6EFDFBFCF7FD3B4069" box="[538,654,803,825]" pageId="37" pageNumber="928">C. Besuchet</collectorName>
(unpubl.; with type labels &quot;
<emphasis id="D1CAEA91FF8A7B6EFC55FCF7FE904000" italics="true" pageId="37" pageNumber="928">
<taxonomicName id="24BE4D00FF8A7B6EFC55FCF7FB964068" authorityName="MULLER &amp; KUNZE" authorityYear="1822" box="[948,1059,803,824]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="928" phylum="Arthropoda" rank="genus">Cephennium</taxonomicName>
foveolatum
</emphasis>
&quot;):
<materialsCitation id="53D63CDEFF8A7B6EFEA0FCEFFCA74001" ID-GBIF-Occurrence="3946761402" box="[321,786,827,849]" collectionCode="MHNG" collectorName="Besuchet" country="Spain" location="Sierra de Cazorla" pageId="37" pageNumber="928" specimenCount="13" specimenCount-female="6" specimenCount-male="7">
<specimenCount id="F5B8FD0AFF8A7B6EFEA0FCEFFEC54000" box="[321,368,827,848]" pageId="37" pageNumber="928" type="male">7♂♂</specimenCount>
,
<specimenCount id="F5B8FD0AFF8A7B6EFE98FCEFFE1D4001" box="[377,424,827,849]" pageId="37" pageNumber="928" type="female">6♀♀</specimenCount>
: &quot;
<location id="E6616058FF8A7B6EFE58FCEFFDE04001" LSID="urn:lsid:plazi:treatment:6B178795FF8A7B6EFF76FD5FFD6B467A:E6616058FF8A7B6EFE58FCEFFDE04001" box="[441,597,827,849]" country="Spain" municipality="Besuchet" name="Sierra de Cazorla" pageId="37" pageNumber="928">Sierra de Cazorla</location>
,
<collectorName id="4E4B5355FF8A7B6EFDBEFCEFFD044001" box="[607,689,827,849]" pageId="37" pageNumber="928">Besuchet</collectorName>
&quot; (
<collectionCode id="85AFAE46FF8A7B6EFD25FCEFFCB94001" box="[708,780,827,849]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="37" pageNumber="928" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="53D63CDEFF8A7B6EFCFDFCEFFE1F4039" ID-GBIF-Occurrence="3946761576" collectorName="de Cazorla" location="Sierra de Cazorla" pageId="37" pageNumber="928" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="F5B8FD0AFF8A7B6EFCFDFCEFFC8C4000" box="[796,825,827,848]" pageId="37" pageNumber="928" type="male">1♂</specimenCount>
,
<specimenCount id="F5B8FD0AFF8A7B6EFCA2FCEFFCC74000" box="[835,882,827,848]" pageId="37" pageNumber="928" type="female">3♀♀</specimenCount>
: &quot;
<location id="E6616058FF8A7B6EFC63FCEFFBAA4001" LSID="urn:lsid:plazi:treatment:6B178795FF8A7B6EFF76FD5FFD6B467A:E6616058FF8A7B6EFC63FCEFFBAA4001" box="[898,1055,827,849]" name="Sierra de Cazorla" pageId="37" pageNumber="928">
Sierra
<collectorName id="4E4B5355FF8A7B6EFC5FFCEFFBAA4001" box="[958,1055,827,849]" pageId="37" pageNumber="928">de Cazorla</collectorName>
</location>
,
<quantity id="24469B66FF8A7B6EFF25FC87FEB64038" box="[196,259,851,872]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="928" unit="m" value="1000.0">1000m</quantity>
, Franz&quot; (
<collectionCode id="85AFAE46FF8A7B6EFEB3FC87FE154039" box="[338,416,851,873]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="37" pageNumber="928" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ABA46508FF8A7B6EFF76FCA3FDED40FB" pageId="37" pageNumber="928" type="etymology">
<paragraph id="E3013683FF8A7B6EFF76FCA3FDED40FB" blockId="37.[151,1057,887,939]" pageId="37" pageNumber="928">
Etymology: The specific epithet is the genitive of Cazorla, the name of the mountain where the
<typeStatus id="3C058821FF8A7B6EFE86FC41FE2640FA" box="[359,403,917,938]" pageId="37" pageNumber="928">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="ABA46508FF8A7B6EFF76FC6EFEAE47D8" pageId="37" pageNumber="928" type="description">
<paragraph id="E3013683FF8A7B6EFF76FC6EFE414778" blockId="37.[151,1059,954,1064]" pageId="37" pageNumber="928">
Description: Body length
<quantity id="24469B66FF8A7B6EFDE4FC6EFDED4080" box="[517,600,954,976]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="928" unit="mm" value="1.4">1.4 mm</quantity>
. Habitus as in
<figureCitation id="7B852A06FF8A7B6EFD17FC6EFCF6409F" box="[758,835,954,976]" captionStart="Figs 17-20" captionStartId="15.[151,191,1499,1521]" captionTargetBox="[160,1024,206,1472]" captionTargetId="figure-43@15.[152,1059,202,1487]" captionTargetPageId="15" captionText="Figs 17-20: Male habitus of species of the subgenus Macroderus in dorsal view: (17) Cephennium segurae; (18) C. nevadae; (19) C. cazorlae; (20) C. malagae. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5039046" httpUri="https://zenodo.org/record/5039046/files/figure.png" pageId="37" pageNumber="928">Fig. 19</figureCitation>
. Colouration reddish. Eyes composed of approximately five ommatidia. Elytra with anterior sulci approximately as long as the combined length of the basal three antennomeres. Other external characters not distinctive.
</paragraph>
<paragraph id="E3013683FF8A7B6EFF76FBECFEAE47D8" blockId="37.[151,1058,1079,1160]" pageId="37" pageNumber="928">
<specimenCount id="F5B8FD0AFF8A7B6EFF76FBECFF18471D" box="[151,173,1080,1101]" pageId="37" pageNumber="928" type="male">♂</specimenCount>
: aedeagus (
<figureCitation id="7B852A06FF8A7B6EFEC7FBE3FE2D471D" box="[294,408,1079,1101]" captionStart="Figs 72-78" captionStartId="23.[151,191,1499,1521]" captionTargetBox="[156,1059,219,1480]" captionTargetId="figure-47@23.[152,1059,200,1485]" captionTargetPageId="23" captionText="Figs 72-78: Cephennium nevadae (72-74), C. carzolae (75-77), and C. malagae (78): (72-73, 75- 76) aedeagus in lateral and in ventral view; (74, 77-78) internal structures of aedeagus in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7250379" httpUri="https://zenodo.org/record/7250379/files/figure.png" pageId="37" pageNumber="928">Figs 75-77</figureCitation>
)
<quantity id="24469B66FF8A7B6EFE49FBE3FDB1471D" box="[424,516,1079,1101]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="928" unit="mm" value="0.4">0.40 mm</quantity>
long; apex of ventral process of triangular shape in ventral view; internal structures large; parameres apically nearly reaching apex of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="ABA46508FF8A7B6EFF76FB43FC18479B" pageId="37" pageNumber="928" type="discussion">
<paragraph id="E3013683FF8A7B6EFF76FB43FC18479B" blockId="37.[151,1058,1175,1227]" pageId="37" pageNumber="928">
Comparative notes:
<taxonomicName id="24BE4D00FF8A7B6EFDE5FB43FD5F47FC" authorityName="Assing" authorityYear="2021" box="[516,746,1175,1196]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="928" phylum="Arthropoda" rank="species" species="cazorlae">
<emphasis id="D1CAEA91FF8A7B6EFDE5FB43FD5F47FC" box="[516,746,1175,1196]" italics="true" pageId="37" pageNumber="928">Cephennium cazorlae</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName id="24BE4D00FF8A7B6EFF76FB61FEA6479A" authorityName="CROISSANDEAU" authorityYear="1894" box="[151,275,1205,1226]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="924" phylum="Arthropoda" rank="subGenus" subGenus="Macroderus">
<emphasis id="D1CAEA91FF8A7B6EFF76FB61FEA6479A" box="[151,275,1205,1226]" italics="true" pageId="37" pageNumber="928">Macroderus</emphasis>
</taxonomicName>
species only by the shape and internal structures of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="ABA46508FF8A7B6EFF76FB0EFD6B467A" pageId="37" pageNumber="928" type="distribution">
<paragraph id="E3013683FF8A7B6EFF76FB0EFD6B467A" blockId="37.[151,1058,1242,1323]" pageId="37" pageNumber="928">
Distribution and natural history: The type locality is situated near the source of Guadalquivir river in Sierra de Cazorla (
<figureCitation id="7B852A06FF8A7B6EFCEBFB23FCE4465D" box="[778,849,1271,1293]" captionStart="Map 3" captionStartId="33.[151,196,985,1006]" captionTargetBox="[164,1060,200,972]" captionTargetId="figure-236@33.[154,1060,200,972]" captionTargetPageId="33" captionText="Map 3: Distributions of Cephennium asturicum (black circles; based on Besuchet (unpubl.) and examined material) and of species of the subgenus Macroderus in the Iberian Peninsula: C. alacantae (black triangles); C. cujonicum (white diamond); C. segurae (black diamond); C. verrens (white circles); C. cazorlae (black square); C. nevadae (black star); C. malagae (white triangles)." figureDoi="http://doi.org/10.5281/zenodo.5039072" httpUri="https://zenodo.org/record/5039072/files/figure.png" pageId="37" pageNumber="928">Map 3</figureCitation>
). The
<typeStatus id="3C058821FF8A7B6EFC78FB23FC47465D" box="[921,1010,1271,1293]" pageId="37" pageNumber="928" type="holotype">holotype</typeStatus>
was sifted in macchia with pine trees at an altitude of
<quantity id="24469B66FF8A7B6EFD6BFAC1FD6F467A" box="[650,730,1301,1322]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="928" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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