<documentid="1FC01E43511161370C7CBBFCDEECE35D"ID-DOI="10.5852/ejt.2021.749.1355"ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a"ID-ISSN="2118-9773"ID-Zenodo-Dep="4770293"ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF"IM.bibliography_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="diego"IM.tables_approvedBy="diego"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1621349695117"checkinUser="felipe"docAuthor="Gao, Chuan-bu & Coca-Abia, María Milagro"docDate="2021"docId="03B56743A67C0666FDE3F96BFCF3E656"docLanguage="en"docName="ejt-749_1-94.pdf"docOrigin="European Journal of Taxonomy 749"docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Miridiba scutata Gao & Coca-Abia 2021, comb. nov."docType="treatment"docVersion="12"lastPageNumber="77"masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431"masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data"masterLastPageNumber="94"masterPageNumber="1"pageNumber="74"updateTime="1733755883393"updateUser="felipe"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="994729CD72A14B39BD6EA9DEE8604F22">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
<mods:affiliationid="4E56999F3E4DEC4B16CDF95F6D0174AB">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China.</mods:affiliation>
<mods:affiliationid="19ADA10E8C5FCBC7BFC6FD6F981066A7">Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain.</mods:affiliation>
<taxonomicNameid="4C1CADD6A67C0663FDE3F96BFC70E2EC"authority="(Reitter, 1902)"authorityName="Gao & Coca-Abia"authorityYear="2021"baseAuthorityName="Reitter"baseAuthorityYear="1902"box="[536,914,1730,1757]"class="Insecta"family="Melolonthidae"genus="Miridiba"kingdom="Animalia"order="Coleoptera"pageId="73"pageNumber="74"phylum="Arthropoda"rank="species"species="scutata"status="comb. nov.">
<bibRefCitationid="EF8DABA4A67C0663FD14F96AFC68E2EC"author="Reitter E."box="[751,906,1731,1757]"pageId="73"pageNumber="74"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<figureCitationid="1327CAD0A67C0663FD39F94FFCAAE331"box="[706,840,1766,1792]"captionStart="Figs 20–22"captionStartId="14.[189,241,1251,1277]"captionTargetBox="[291,1324,285,1192]"captionTargetId="figure-142@14.[278,1337,265,1205]"captionTargetPageId="14"captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770322"httpUri="https://zenodo.org/record/4770322/files/figure.png"pageId="73"pageNumber="74">Figs 20–22</figureCitation>
,
<figureCitationid="1327CAD0A67C0663FCAFF94FFC93E331"box="[852,881,1766,1792]"captionStart="Figs 94–97. 94"captionStartId="75.[189,241,1887,1913]"captionTargetBox="[277,1301,581,1832]"captionTargetId="figure-102@75.[264,1324,560,1851]"captionTargetPageId="75"captionText="Figs 94–97. 94. Miridiba scutata (Reitter, 1902), lectotype, ♂ (MFNB). Habitus in drosal view. 95– 97. M. schoolmeesteri Keith, 2010, holotype, ♂ (DKC). 95. Habitus in dorsal view. 96. Parameres in dorsal (A) and lateral (B) views. 97. Endophallus and temones in lateral view. Scale bars: 94–95 = 5.0 mm; 96–97 = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770413"httpUri="https://zenodo.org/record/4770413/files/figure.png"pageId="73"pageNumber="74">94</figureCitation>
<bibRefCitationid="EF8DABA4A67C0663FE52F88EFD9FE373"author="Reitter E."box="[425,637,1831,1858]"pageId="73"pageNumber="74"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902: 175</bibRefCitation>
<bibRefCitationid="EF8DABA4A67C0663FE52F8E2FD9AE354"author="Moser J."box="[425,632,1867,1893]"pageId="73"pageNumber="74"pagination="468 - 488"refId="ref45295"refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488."type="journal article"year="1909">Moser, 1909: 471</bibRefCitation>
<bibRefCitationid="EF8DABA4A67C0663FBF0F8E3FB45E354"author="Reitter E."box="[1035,1191,1866,1893]"pageId="73"pageNumber="74"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<treatmentCitationid="0ABDF044A67C0663FF46F8C7FD2BE3B9"author="Dalla Torre K. W. von"box="[189,713,1902,1928]"page="206"pageId="73"pageNumber="74"year="1912">
<bibRefCitationid="EF8DABA4A67C0663FE44F8C7FD2BE3B9"author="Dalla Torre K. W. von"box="[447,713,1902,1928]"pageId="73"pageNumber="74"pagination="135 - 290"refId="ref43620"refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany."type="book chapter"year="1912">Dalla Torre, 1912: 206</bibRefCitation>
<bibRefCitationid="EF8DABA4A67C0663FB98F8C7FB18E3B9"author="Moser J."box="[1123,1274,1902,1928]"pageId="73"pageNumber="74"pagination="468 - 488"refId="ref45295"refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488."type="journal article"year="1909">Moser, 1909</bibRefCitation>
<treatmentCitationid="0ABDF044A67F0660FDEDFEA0FD03E512"author="Chang Y. - W."box="[534,737,265,291]"page="149"pageId="74"pageNumber="75"year="1964">
<bibRefCitationid="EF8DABA4A67F0660FDEDFEA0FD03E512"author="Chang Y. - W."box="[534,737,265,291]"pageId="74"pageNumber="75"pagination="139 - 152"refId="ref43461"refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152."type="journal article"year="1964">Chang 1964: 149</bibRefCitation>
<treatmentCitationid="0ABDF044A67F0660FDE9FE85FCACE577"author="Smetana A. & Kral D."box="[530,846,300,326]"page="220"pageId="74"pageNumber="75"year="2006">
<bibRefCitationid="EF8DABA4A67F0660FDE9FE85FCACE577"author="Smetana A. & Kral D."box="[530,846,300,326]"pageId="74"pageNumber="75"pagination="207 - 228"refId="ref45912"refString="Smetana A. & Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. & Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark."type="book chapter"year="2006">Smetana & Král 2006: 220</bibRefCitation>
<treatmentCitationid="0ABDF044A67F0660FD90FEE6FCDBE55B"author="Chang Y. - W."box="[619,825,335,362]"page="149"pageId="74"pageNumber="75"year="1964">
<bibRefCitationid="EF8DABA4A67F0660FD90FEE6FCDBE55B"author="Chang Y. - W."box="[619,825,335,362]"pageId="74"pageNumber="75"pagination="139 - 152"refId="ref43461"refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152."type="journal article"year="1964">Chang 1964: 149</bibRefCitation>
</treatmentCitation>
(species list; in key). —
<treatmentCitationid="0ABDF044A67F0660FB92FEF9FEFAE5BC"author="Smetana A. & Kral D."page="220"pageId="74"pageNumber="75"year="2006">
<bibRefCitationid="EF8DABA4A67F0660FB92FEF9FEFAE5BC"author="Smetana A. & Kral D."pageId="74"pageNumber="75"pagination="207 - 228"refId="ref45912"refString="Smetana A. & Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. & Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark."type="book chapter"year="2006">Smetana & Král 2006: 220</bibRefCitation>
<figureCitationid="1327CAD0A67F0660FDCEFE4BFD72E5CC"box="[565,656,482,509]"captionStart="Figs 94–97. 94"captionStartId="75.[189,241,1887,1913]"captionTargetBox="[277,1301,581,1832]"captionTargetId="figure-102@75.[264,1324,560,1851]"captionTargetPageId="75"captionText="Figs 94–97. 94. Miridiba scutata (Reitter, 1902), lectotype, ♂ (MFNB). Habitus in drosal view. 95– 97. M. schoolmeesteri Keith, 2010, holotype, ♂ (DKC). 95. Habitus in dorsal view. 96. Parameres in dorsal (A) and lateral (B) views. 97. Endophallus and temones in lateral view. Scale bars: 94–95 = 5.0 mm; 96–97 = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770413"httpUri="https://zenodo.org/record/4770413/files/figure.png"pageId="74"pageNumber="75">Fig. 94</figureCitation>
). Body size 14.0–16.0 mm. Dorsal surface glabrous. Clypeus rounded laterally, with conspicuous punctures, anterior edge emarginate at middle. Frons with carina moderately developed and blunt. Pronotal surface shiny, regularly punctate, with punctures closer together just behind anterior margin, glabrous, at most with a tiny seta on each puncture, anterior pronotal margin flanged, glabrous; posterior margin glabrous; lateral margins serrated and pubescent; anterior angles almost right not projected forward, posterior angles obtuse and marked. Scutellum without punctures and glabrous. Elytral surface glabrous, at most with a tiny seta on each puncture, with striations weakly defined, punctures distributed between striae; epipleuron glabrous, at most with few setae at base. Foretibia with dorsal carina moderately developed; insertion of inner spur closer third outer tooth than second one. Meso- and metatibia with transverse carina complete, inner margin of dorsal surface without spines. Pygidium with deep punctuation, surface with inconspicuous pubescence, margin pubescent. Ventrites with conspicuous punctures; ventrites 2, 3 and 4 glabrous, at most with a tiny seta in each puncture; ventrite 5 with long pubescence irregularly distributed at posterior half; ventrite 6 moderately bulging at anterior margin, with long pubescence on posterior margin. Male genitalia: parameres (
<figureCitationid="1327CAD0A67F0660FE46FC66FDF6E7D8"box="[445,532,975,1001]"captionStart="Figs 20–22"captionStartId="14.[189,241,1251,1277]"captionTargetBox="[291,1324,285,1192]"captionTargetId="figure-142@14.[278,1337,265,1205]"captionTargetPageId="14"captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770322"httpUri="https://zenodo.org/record/4770322/files/figure.png"pageId="74"pageNumber="75">Fig. 20</figureCitation>
) with dorsal and ventral branches joined laterally. Dorsal branches fused with apex asymmetrical, elevated and twisted. Ventral branches reduced, lightly sclerotized and fused by a ventral membrane. Endophallus (
<figureCitationid="1327CAD0A67F0660FDA3FBBFFD4CE001"box="[600,686,1046,1072]"captionStart="Figs 20–22"captionStartId="14.[189,241,1251,1277]"captionTargetBox="[291,1324,285,1192]"captionTargetId="figure-142@14.[278,1337,265,1205]"captionTargetPageId="14"captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770322"httpUri="https://zenodo.org/record/4770322/files/figure.png"pageId="74"pageNumber="75">Fig. 21</figureCitation>
) with a raspula with soft setae and spines at distal end; with spines isolated at middle. Apophysis of temones (
<figureCitationid="1327CAD0A67F0660FCF2FB90FCBDE062"box="[777,863,1081,1107]"captionStart="Figs 20–22"captionStartId="14.[189,241,1251,1277]"captionTargetBox="[291,1324,285,1192]"captionTargetId="figure-142@14.[278,1337,265,1205]"captionTargetPageId="14"captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770322"httpUri="https://zenodo.org/record/4770322/files/figure.png"pageId="74"pageNumber="75">Fig. 21</figureCitation>
) long and separated from each other. Female genitalia (
<figureCitationid="1327CAD0A67F0660FEC8FBF5FE6BE047"box="[307,393,1116,1142]"captionStart="Figs 20–22"captionStartId="14.[189,241,1251,1277]"captionTargetBox="[291,1324,285,1192]"captionTargetId="figure-142@14.[278,1337,265,1205]"captionTargetPageId="14"captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.4770322"httpUri="https://zenodo.org/record/4770322/files/figure.png"pageId="74"pageNumber="75">Fig. 22</figureCitation>
<bibRefCitationid="EF8DABA4A67F0660FBAFFB46FB0FE138"author="Reitter E."box="[1108,1261,1263,1289]"pageId="74"pageNumber="75"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<taxonomicNameid="4C1CADD6A67F0660FDF9FABBFC60E11C"authority="(Reitter, 1902)"authorityName="Gao & Coca-Abia"authorityYear="2021"baseAuthorityName="Reitter"baseAuthorityYear="1902"box="[514,898,1298,1325]"class="Insecta"family="Melolonthidae"genus="Miridiba"kingdom="Animalia"order="Coleoptera"pageId="74"pageNumber="75"phylum="Arthropoda"rank="species"species="scutata">
<bibRefCitationid="EF8DABA4A67F0660FD20FABBFC98E11C"author="Reitter E."box="[731,890,1298,1325]"pageId="74"pageNumber="75"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<collectionCodeid="ED0D4E90A67F0660FF46FA9FFEF6E161"box="[189,276,1334,1360]"country="Italy"httpUri="http://grbio.org/cool/irpn-c0nq"name="Museo Friulano di Storia Naturale"pageId="74"pageNumber="75">MFNB</collectionCode>
<materialsCitationid="3B74DC08A67F0660FF11FADEFB20E1E9"ID-GBIF-Occurrence="3114121385"collectionCode="MFNB"collectorName="Brenske"country="China"determinerName="Chuan-bu Gao & Coca-Abia"location="CHINA"pageId="74"pageNumber="75"specimenCount="1"specimenCount-male="1"typeStatus="lectotype">
<bibRefCitationid="EF8DABA4A67F0660FE53FA14FD9BE1E9"author="Dalla Torre K. W. von"box="[424,633,1469,1496]"pageId="74"pageNumber="75"pagination="135 - 290"refId="ref43620"refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany."type="book chapter"year="1912">Dalla Torre, 1912</bibRefCitation>
<collectionCodeid="ED0D4E90A67F0660FB90FA17FB20E1E9"box="[1131,1218,1470,1496]"collectionName="Italy, Udine, Museo Friulano di Storia Naturale"country="Italy"httpUri="http://grbio.org/cool/irpn-c0nq"name="Museo Friulano di Storia Naturale"pageId="74"pageNumber="75">MFNB</collectionCode>
<materialsCitationid="3B74DC08A67F0660FF46F98BFD64E251"ID-GBIF-Occurrence="3114121351"collectingDate="2004-05"collectionCode="DKC"collectorName="Lassalle & Vietnam"country="Vietnam"determinerName="D. Keith"location="VIETNAM"pageId="74"pageNumber="75"specimenCount="1"specimenCount-male="1"stateProvince="Yen Bai">
<locationDeviationid="9A11CF77A67F0660FE6BF98BFD35E20D"box="[400,727,1570,1597]"pageId="74"pageNumber="75">Nghia Lo ouest de Yen Bai</locationDeviation>
” [
<locationDeviationid="9A11CF77A67F0660FD0AF98BFBD5E20C"box="[753,1079,1570,1597]"pageId="74"pageNumber="75">Nghia Lo, west of Yen Bai</locationDeviation>
<materialsCitationid="3B74DC08A67F0660FF46F903FCD4E2F4"ID-GBIF-Occurrence="3114121373"box="[189,822,1706,1733]"collectionCode="MFNB"collectorName="Pipitz & Coll. Brenske"country="China"location="CHINA"pageId="74"pageNumber="75"specimenCount="2"specimenCount-female="1"specimenCount-male="1">
<collectionCodeid="ED0D4E90A67F0660FD24F902FCD4E2F4"box="[735,822,1707,1733]"collectionName="Italy, Udine, Museo Friulano di Storia Naturale"country="Italy"httpUri="http://grbio.org/cool/irpn-c0nq"name="Museo Friulano di Storia Naturale"pageId="74"pageNumber="75">MFNB</collectionCode>
), mentioning a publication, which, supposedly, contained the original species description. Unfortunately, we have not found this paper.
<bibRefCitationid="EF8DABA4A67F0660FD1EF8C9FC6FE34A"author="Reitter E."box="[741,909,1888,1915]"pageId="74"pageNumber="75"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter (1902)</bibRefCitation>
without specifying the origin or number of specimens described. Later,
<bibRefCitationid="EF8DABA4A67F0660FBD6F82DFB36E3AF"author="Moser J."box="[1069,1236,1924,1950]"pageId="74"pageNumber="75"pagination="468 - 488"refId="ref45295"refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488."type="journal article"year="1909">Moser (1909)</bibRefCitation>
, ignoring the existence of Brenske’s species and Reitter’s description, published another taxon named
<bibRefCitationid="EF8DABA4A67F0660FEEEF862FE45E3D5"author="Moser J."box="[277,423,1994,2021]"pageId="74"pageNumber="75"pagination="468 - 488"refId="ref45295"refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488."type="journal article"year="1909">Moser, 1909</bibRefCitation>
<bibRefCitationid="EF8DABA4A67F0660FCCAF863FBE6E3D5"author="Dalla Torre K. W. von"box="[817,1028,1994,2021]"pageId="74"pageNumber="75"pagination="135 - 290"refId="ref43620"refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany."type="book chapter"year="1912">Dalla Torre (1912)</bibRefCitation>
realized the primary homonymy and proposed the name
<bibRefCitationid="EF8DABA4A67E0661FC66FEA3FBA3E512"author="Moser J."box="[925,1089,265,292]"pageId="75"pageNumber="76"pagination="468 - 488"refId="ref45295"refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488."type="journal article"year="1909">Moser (1909)</bibRefCitation>
<bibRefCitationid="EF8DABA4A67E0661FE36FE85FD86E577"author="Reitter E."box="[461,612,300,326]"pageId="75"pageNumber="76"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
(MFNB), which shows external features that characterize the genus
<bibRefCitationid="EF8DABA4A67E0661FD62FEE6FCD3E55B"author="Reitter E."box="[665,817,335,362]"pageId="75"pageNumber="76"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<bibRefCitationid="EF8DABA4A67E0661FB5FFEDBFA95E5BC"author="Dalla Torre K. W. von"box="[1188,1399,370,397]"pageId="75"pageNumber="76"pagination="135 - 290"refId="ref43620"refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany."type="book chapter"year="1912">Dalla Torre, 1912</bibRefCitation>
</taxonomicName>
(MFNB), verifying the similarities of external morphology and male genitalia (parameres shape, sclerotized structures and sensillae in the endophallus) between both taxa, which confirm the synonymy of
based on a male specimen. According to the original description and illustrations (Keith 2010: figs 9–11), the characteristic shape of aedeagus, its asymmetry, the fusion and twist of dorsal branches, and fusion of ventral ones, distinguish it from other known
<bibRefCitationid="EF8DABA4A6790666FDF7FE10FD44E5E2"author="Reitter E."box="[524,678,441,467]"pageId="76"pageNumber="77"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>
<taxonomicNameid="4C1CADD6A67E0661FE7EF8F6FCE5E348"authority="(Reitter, 1902)"authorityName="Gao & Coca-Abia"authorityYear="2021"baseAuthorityName="Reitter"baseAuthorityYear="1902"box="[389,775,1887,1913]"class="Insecta"family="Melolonthidae"genus="Miridiba"kingdom="Animalia"order="Coleoptera"pageId="75"pageNumber="76"phylum="Arthropoda"rank="species"species="scutata">
<bibRefCitationid="EF8DABA4A67E0661FDA6F8F6FD1CE348"author="Reitter E."box="[605,766,1887,1913]"pageId="75"pageNumber="76"pagination="93 - 303"refId="ref45541"refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303."type="journal article"year="1902">Reitter, 1902</bibRefCitation>