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<document id="390D3424E22FE71C56AFA8C08C560F52" ID-DOI="10.11646/zootaxa.3721.2.3" ID-GBIF-Dataset="f3d01271-9143-4021-b5ec-45791021361d" ID-ISSN="1175-5326" ID-Zenodo-Dep="218326" ID-ZooBank="71B4B001-FB10-4B99-ACF9-720131457534" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460688744557" checkinUser="plazi" docAuthor="Salvador, Rodrigo Brincalepe" docDate="2013" docId="5C371C6D2719FFC8FF208CB0FDF4FD9C" docLanguage="en" docName="zt03721p171.pdf" docOrigin="Zootaxa 3721 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Testacellidae" docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="A00E6415271DFFCEFFB78C28FFCAFF84" masterDocTitle="The fossil pulmonate snails of Sandelzhausen (Early / Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea" masterLastPageNumber="171" masterPageNumber="157" pageNumber="161" updateTime="1698351515074" updateUser="plazi">
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<mods:title id="F2E468F9C73A819AD024322A1A267091">The fossil pulmonate snails of Sandelzhausen (Early / Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea</mods:title>
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<mods:namePart id="4C9F4723B45BABDF6A01D4191B344B81">Salvador, Rodrigo Brincalepe</mods:namePart>
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<mods:date id="D40E8BE7EFD87CB9F6881C0B2F128443">2013</mods:date>
<mods:detail id="D5AC717CD7CCEF02849E8C7115D6231E" type="volume">
<mods:number id="5FB15DCD6E508FD72AAE25ED5DB97F6F">3721</mods:number>
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<mods:identifier id="5A2FBA28636F70F7C2837BEACE102CAC" type="DOI">10.11646/zootaxa.3721.2.3</mods:identifier>
<mods:identifier id="4790C32CB7C54851458F7B3B5E80058E" type="GBIF-Dataset">f3d01271-9143-4021-b5ec-45791021361d</mods:identifier>
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<treatment id="5C371C6D2719FFC8FF208CB0FDF4FD9C" ID-DOI="http://doi.org/10.5281/zenodo.6154543" ID-GBIF-Taxon="119577099" ID-Zenodo-Dep="6154543" LSID="urn:lsid:plazi:treatment:5C371C6D2719FFC8FF208CB0FDF4FD9C" httpUri="http://treatment.plazi.org/id/5C371C6D2719FFC8FF208CB0FDF4FD9C" lastPageId="6" lastPageNumber="163" pageId="4" pageNumber="161">
<subSubSection id="9C84FEF02719FFCAFF208CB0FE3DFF73" pageId="4" pageNumber="161" type="nomenclature">
<paragraph id="D421AD7B2719FFCAFF208CB0FE5CFF36" blockId="4.[151,406,152,178]" box="[151,406,152,178]" pageId="4" pageNumber="161">
<heading id="8F691A172719FFCAFF208CB0FE5CFF36" bold="true" box="[151,406,152,178]" fontSize="11" level="1" pageId="4" pageNumber="161" reason="1">
<emphasis id="E6EA71692719FFCAFF208CB0FE5CFF36" bold="true" box="[151,406,152,178]" pageId="4" pageNumber="161">
Family
<taxonomicName id="139ED6F82719FFCAFF418CB0FE5CFF36" ID-CoL="7NL9L" box="[246,406,152,178]" class="Gastropoda" family="Testacellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="family">Testacellidae</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208CF6FE3DFF73" blockId="4.[151,503,222,247]" box="[151,503,222,247]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF208CF6FE3DFF73" bold="true" box="[151,503,222,247]" pageId="4" pageNumber="161">
Genus
<taxonomicName id="139ED6F82719FFCAFF5D8CF6FE3DFF73" authority="Draparnaud" authorityName="Draparnaud" box="[234,503,222,247]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF5D8CF6FE96FF73" bold="true" box="[234,348,222,247]" italics="true" pageId="4" pageNumber="161">Testacella</emphasis>
Draparnaud
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF208D0EFDEAFD80" pageId="4" pageNumber="161" type="description">
<paragraph id="D421AD7B2719FFCAFF208D0EFF21FEE7" blockId="4.[151,468,294,355]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208D0EFE1EFEBB" authority="Klein, 1853" authorityName="Klein" authorityYear="1853" box="[151,468,294,319]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="zellii">
<emphasis id="E6EA71692719FFCAFF208D0EFE1EFEBB" bold="true" box="[151,468,294,319]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF208D0EFE8FFEBB" bold="true" box="[151,325,294,319]" italics="true" pageId="4" pageNumber="161">Testacella zellii</emphasis>
Klein, 1853
</emphasis>
</taxonomicName>
(
<figureCitation id="4CA5B1FE2719FFCAFF288D62FF28FEE7" box="[159,226,330,355]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="4" pageNumber="161">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208DBAFD9BFE43" blockId="4.[151,729,402,517]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208DBAFE2FFE2C" authority="Klein, 1853: 204" authorityName="Klein" authorityPageNumber="204" authorityYear="1853" box="[151,485,402,424]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="zellii">
<emphasis id="E6EA71692719FFCAFF208DBAFEFEFE2C" box="[151,308,402,424]" italics="true" pageId="4" pageNumber="161">Testacella zellii</emphasis>
Klein, 1853: 204
</taxonomicName>
(pl. 5, fig. 1).
<taxonomicName id="139ED6F82719FFCAFF208D99FE4FFE43" box="[151,389,433,455]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="subSpecies" species="larteti" subSpecies="larteti">
<emphasis id="E6EA71692719FFCAFF208D99FE4FFE43" box="[151,389,433,455]" italics="true" pageId="4" pageNumber="161">Testacella larteti larteti</emphasis>
</taxonomicName>
[sic]: Gall, 1972: 7.
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208DF8FDEAFD80" blockId="4.[151,729,402,517]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208DF8FEE7FE62" box="[151,301,464,486]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="zelli">
<emphasis id="E6EA71692719FFCAFF208DF8FEE7FE62" box="[151,301,464,486]" italics="true" pageId="4" pageNumber="161">Testacella zelli</emphasis>
</taxonomicName>
[sic]: Schlickum, 1976: 15 (pl. 3, fig. 51).
<taxonomicName id="139ED6F82719FFCAFF208DC6FF37FD80" box="[151,253,494,516]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF208DC6FF37FD80" box="[151,253,494,516]" italics="true" pageId="4" pageNumber="161">Testacella</emphasis>
</taxonomicName>
sp.: Moser
<emphasis id="E6EA71692719FFCAFEC38DD8FE6AFD80" box="[372,416,494,517]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
., 2009b: 48.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF208E19FEB6FD16" pageId="4" pageNumber="161" type="materials_examined">
<paragraph id="D421AD7B2719FFCAFF208E19FD21FDCD" blockId="4.[151,1436,561,1234]" box="[151,747,561,586]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF208E19FE4AFDCE" bold="true" box="[151,384,561,586]" pageId="4" pageNumber="161">Material examined.</emphasis>
BSPG
<date id="A0208BBB2719FFCAFE638E19FDECFDCD" box="[468,550,561,586]" pageId="4" pageNumber="161" value="1959-02">1959 II</date>
17689 (1 spcm.).
</paragraph>
<paragraph id="D421AD7B2719FFCAFF708E7DFEB6FD16" blockId="4.[151,1436,561,1234]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708E7DFE32FDEA" bold="true" box="[199,504,597,622]" pageId="4" pageNumber="161">Stratigraphic occurrence.</emphasis>
The single specimen has no stratigraphical data, but Moser
<emphasis id="E6EA71692719FFCAFB158E7EFB1EFDEA" box="[1186,1236,597,622]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
. (2009b) indicate either layer B or C1.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF708EB5FCDDFCA6" pageId="4" pageNumber="161" type="description">
<paragraph id="D421AD7B2719FFCAFF708EB5FD97FD7A" blockId="4.[151,1436,561,1234]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708EB5FE93FD32" bold="true" box="[199,345,669,694]" pageId="4" pageNumber="161">Description.</emphasis>
Shell small, ear-shaped, vestigial. Apex starkly pointed. Shell smooth, except for growth lines. Aperture large and elongated, with a near right angle on the encounter between parietal and palatal regions; parietal and columellar region greatly thickened.
</paragraph>
<paragraph id="D421AD7B2719FFCAFF708F21FCDDFCA6" blockId="4.[151,1436,561,1234]" box="[199,791,777,802]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708F21FE29FCA6" bold="true" box="[199,483,777,802]" pageId="4" pageNumber="161">Measurements (in mm).</emphasis>
¾ whorl; H = 1.7; D = 2.3.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF708F05FE21FCED" pageId="4" pageNumber="161" type="materials_examined">
<paragraph id="D421AD7B2719FFCAFF708F05FE21FCED" blockId="4.[151,1436,561,1234]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708F05FD59FCC2" bold="true" box="[199,659,813,838]" pageId="4" pageNumber="161">Previous identification of the material.</emphasis>
Gall (1972: Nr. 3):
<taxonomicName id="139ED6F82719FFCAFCC48F05FBB6FCC2" box="[883,1148,813,838]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="subSpecies" species="larteti" subSpecies="larteti">
<emphasis id="E6EA71692719FFCAFCC48F05FBB6FCC2" box="[883,1148,813,838]" italics="true" pageId="4" pageNumber="161">Testacella larteti larteti</emphasis>
</taxonomicName>
[sic] Dupuy. Moser
<emphasis id="E6EA71692719FFCAFAD38F06FA5FFCC2" box="[1380,1429,813,838]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
. (2009b: Nr. 42):
<taxonomicName id="139ED6F82719FFCAFEE48F79FE0EFCEE" box="[339,452,849,874]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFEE48F79FE0EFCEE" box="[339,452,849,874]" italics="true" pageId="4" pageNumber="161">Testacella</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF708F5DFD26FB56" pageId="4" pageNumber="161" type="discussion">
<paragraph id="D421AD7B2719FFCAFF708F5DFBF3FBC6" blockId="4.[151,1436,561,1234]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708F5DFE86FC0A" bold="true" box="[199,332,885,910]" pageId="4" pageNumber="161">Discussion.</emphasis>
The shell is vestigial in this monotypic family and, therefore, is of very limited taxonomical value. The single specimen found in Sandelzhausen is fragmentary; nevertheless it presents some distinctive features.
<collectingCountry id="AC89EDEB2719FFCAFACA8FB2FA56FC36" box="[1405,1436,922,946]" name="American Samoa" pageId="4" pageNumber="161">As</collectingCountry>
already stated by Moser
<emphasis id="E6EA71692719FFCAFE198F96FE2BFC52" box="[430,481,957,982]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
. (2009b), the present specimen differs from
<taxonomicName id="139ED6F82719FFCAFC6B8F96FB52FC52" authority="Dupuy" authorityName="Dupuy" box="[988,1176,957,982]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="lartetii">
<emphasis id="E6EA71692719FFCAFC6B8F96FB8EFC52" box="[988,1092,957,982]" italics="true" pageId="4" pageNumber="161">T. lartetii</emphasis>
Dupuy
</taxonomicName>
by being much deeper and by having an almost right angle on its aperture and thus cannot be classified as this species. These features, alongside the starkly pointed apex, compare fittingly to
<taxonomicName id="139ED6F82719FFCAFCBF882EFC93FB9A" box="[776,857,1029,1054]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="zellii">
<emphasis id="E6EA71692719FFCAFCBF882EFC93FB9A" box="[776,857,1029,1054]" italics="true" pageId="4" pageNumber="161">T. zellii</emphasis>
</taxonomicName>
, a species known only from a few fossil sites of the
<collectingCountry id="AC89EDEB2719FFCAFF208802FF3BFBC6" box="[151,241,1066,1090]" name="Germany" pageId="4" pageNumber="161">German</collectingCountry>
Early/Middle Miocene Silvana-beds (
<emphasis id="E6EA71692719FFCAFD298801FC96FBC6" box="[670,860,1065,1090]" italics="true" pageId="4" pageNumber="161">Silvanaschichten</emphasis>
; Schlickum, 1976).
</paragraph>
<paragraph id="D421AD7B2719FFCAFF708866FD26FB56" blockId="4.[151,1436,561,1234]" pageId="4" pageNumber="161">
Recent
<taxonomicName id="139ED6F82719FFCAFE938865FE5FFBE2" box="[292,405,1101,1126]" class="Gastropoda" family="Testacellidae" genus="Testacella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFE938865FE5FFBE2" box="[292,405,1101,1126]" italics="true" pageId="4" pageNumber="161">Testacella</emphasis>
</taxonomicName>
species are voracious predators, preying on slugs, snails, centipedes and, especially, earthworms; living mainly underground and requiring, at least, a moderately moist soil cover (Barker &amp; Efford, 2004). This burrowing habit should make it difficult for the shells to be carried away post-mortem and preserved in the lake sediments, hence its rarity in Sandelzhausen.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF20890DFCECF97E" pageId="4" pageNumber="161" type="description">
<paragraph id="D421AD7B2719FFCAFF20890DFE7AFABA" blockId="4.[151,432,1317,1342]" box="[151,432,1317,1342]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF20890DFE7AFABA" bold="true" box="[151,432,1317,1342]" pageId="4" pageNumber="161">Superfamily Helicoidea</emphasis>
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208945FE96FA02" blockId="4.[151,348,1389,1414]" box="[151,348,1389,1414]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF208945FE96FA02" bold="true" box="[151,348,1389,1414]" pageId="4" pageNumber="161">
Family
<taxonomicName id="139ED6F82719FFCAFF458945FE96FA02" box="[242,348,1389,1414]" class="Gastropoda" family="Elonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="family">Elonidae</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="D421AD7B2719FFCAFF20899DFE47FA4A" blockId="4.[151,397,1461,1486]" box="[151,397,1461,1486]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF20899DFE47FA4A" bold="true" box="[151,397,1461,1486]" pageId="4" pageNumber="161">
Genus
<taxonomicName id="139ED6F82719FFCAFF5D899DFE47FA4A" authority="Pilsbry" authorityName="Pilsbry" box="[234,397,1461,1486]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF5D899DFEFAFA4A" bold="true" box="[234,304,1461,1486]" italics="true" pageId="4" pageNumber="161">Klikia</emphasis>
Pilsbry
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="D421AD7B2719FFCAFF2089D5FEDBF9BE" blockId="4.[151,531,1533,1594]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF2089D5FDD9F992" bold="true" box="[151,531,1533,1558]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF2089D5FF17F992" box="[151,221,1533,1558]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF2089D5FF17F992" bold="true" box="[151,221,1533,1558]" italics="true" pageId="4" pageNumber="161">Klikia</emphasis>
</taxonomicName>
cf. c
<emphasis id="E6EA71692719FFCAFEA389D6FEB8F991" bold="true" box="[276,370,1534,1557]" italics="true" pageId="4" pageNumber="161">oarctata</emphasis>
(Klein, 1853)
</emphasis>
(
<figureCitation id="4CA5B1FE2719FFCAFF288A09FEC0F9BE" box="[159,266,1569,1594]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="4" pageNumber="161">Figs. 6–9</figureCitation>
)
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208A41FDBBF9FB" blockId="4.[151,957,1641,1786]" box="[151,625,1641,1663]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208A41FF04F9FB" box="[151,206,1641,1663]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF208A41FF04F9FB" box="[151,206,1641,1663]" italics="true" pageId="4" pageNumber="161">Helix</emphasis>
</taxonomicName>
c
<emphasis id="E6EA71692719FFCAFF688A42FEFEF9FB" box="[223,308,1642,1663]" italics="true" pageId="4" pageNumber="161">oarctata</emphasis>
Klein, 1853: 206 (pl. 5, fig. 6).
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208AA0FD0AF91A" blockId="4.[151,957,1641,1786]" box="[151,704,1672,1694]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208AA0FD76F91A" authority="Wenz, 1923a: 534" authorityName="Wenz" authorityPageNumber="534" authorityYear="1923" box="[151,700,1672,1694]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="subSpecies" species="coarctata" subGenus="Apula" subSpecies="coarctata">
<emphasis id="E6EA71692719FFCAFF208AA0FE3FF91A" box="[151,501,1672,1694]" italics="true" pageId="4" pageNumber="161">Klikia (Apula) coarctata coarctata</emphasis>
: Wenz, 1923a: 534
</taxonomicName>
.
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208A8EFD78F938" blockId="4.[151,957,1641,1786]" box="[151,690,1702,1725]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208A8EFD64F938" authority="Gall, 1972: 10" authorityName="Gall" authorityPageNumber="10" authorityYear="1972" box="[151,686,1702,1725]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="subSpecies" species="giengensis" subGenus="Klikia" subSpecies="giengensis">
<emphasis id="E6EA71692719FFCAFF208A8EFDC6F939" box="[151,524,1702,1725]" italics="true" pageId="4" pageNumber="161">Klikia (Klikia) giengensis giengensis</emphasis>
: Gall, 1972: 10
</taxonomicName>
.
</paragraph>
<paragraph id="D421AD7B2719FFCAFF208AEDFCECF97E" blockId="4.[151,957,1641,1786]" pageId="4" pageNumber="161">
<taxonomicName id="139ED6F82719FFCAFF208AEDFDA4F95F" authority="Schlickum, 1976: 17" authorityName="Schlickum" authorityPageNumber="17" authorityYear="1976" box="[151,622,1733,1755]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="species" species="coarctata" subGenus="Apula">
<emphasis id="E6EA71692719FFCAFF208AEDFE45F95F" box="[151,399,1733,1755]" italics="true" pageId="4" pageNumber="161">Klikia (Apula) coarctata</emphasis>
: Schlickum, 1976: 17
</taxonomicName>
(pl. 4, fig. 60); Kókay, 2006: 92.
<taxonomicName id="139ED6F82719FFCAFF208ACCFE9CF97D" authority="Prosoosia" authorityName="Prosoosia" box="[151,342,1764,1785]" class="Gastropoda" family="Helicodontidae" genus="Soosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692719FFCAFF208ACCFE9CF97D" box="[151,342,1764,1785]" italics="true" pageId="4" pageNumber="161">Soosia (Prosoosia)</emphasis>
</taxonomicName>
cf.
<emphasis id="E6EA71692719FFCAFEC98ACCFE03F97E" box="[382,457,1764,1786]" italics="true" pageId="4" pageNumber="161">godarti</emphasis>
: Moser
<emphasis id="E6EA71692719FFCAFDAA8ACDFD83F97E" box="[541,585,1764,1786]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
., 2009b: 49 (fig. 6B).
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCAFF208B0EFAD2F803" pageId="4" pageNumber="161" type="materials_examined">
<paragraph id="D421AD7B2719FFCAFF208B0EFC65F8BB" blockId="4.[151,1436,1830,2035]" box="[151,943,1830,1855]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF208B0EFE4AF8BB" bold="true" box="[151,384,1830,1855]" pageId="4" pageNumber="161">Material examined.</emphasis>
BSPG
<date id="A0208BBB2719FFCAFE638B0EFDECF8BA" box="[468,550,1830,1855]" pageId="4" pageNumber="161" value="1959-02">1959 II</date>
16144 (1 spcm.), 17301 (1 spcm.).
</paragraph>
<paragraph id="D421AD7B2719FFCAFF708B62FAD2F803" blockId="4.[151,1436,1830,2035]" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708B62FE3DF8E7" bold="true" box="[199,503,1866,1891]" pageId="4" pageNumber="161">Stratigraphic occurrence.</emphasis>
Layers C3 (probably) and D1. Specimen BSPG
<date id="A0208BBB2719FFCAFBA48B62FBAEF8E6" box="[1043,1124,1866,1891]" pageId="4" pageNumber="161" value="1959-02">1959 II</date>
– 16144 comes from a sand lens in the western part of the fossil site, which is probably layer C3 according to Moser
<emphasis id="E6EA71692719FFCAFBCF8B47FB63F803" box="[1144,1193,1902,1927]" italics="true" pageId="4" pageNumber="161">et al</emphasis>
. (2009b).
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02719FFCBFF708BBAFAAAFEC4" lastPageId="5" lastPageNumber="162" pageId="4" pageNumber="161" type="description">
<paragraph id="D421AD7B2719FFCBFF708BBAFD9CFE99" blockId="4.[151,1436,1830,2035]" lastBlockId="5.[151,1436,151,608]" lastPageId="5" lastPageNumber="162" pageId="4" pageNumber="161">
<emphasis id="E6EA71692719FFCAFF708BBAFE93F82F" bold="true" box="[199,345,1938,1963]" pageId="4" pageNumber="161">Description.</emphasis>
Shell small, depressed; shell height ~1/2 its width. Protoconch (~1 whorl) blunt, wide, apparently smooth; transition to teleoconch unclear. Teleoconch sculptured by regularly distributed fine papillae, giving the impression of axial striae; papillae apparently stronger closer to suture and to umbilicus. Whorls’ profile slightly convex. Suture well-marked, deep. Whorls regularly increasing in size, but with a decrease in size in the last quarter of body whorl. Body whorl with conspicuous constriction right before the aperture (“extralabial depression”
<emphasis id="E6EA71692718FFCBFE928CC9FEA9FF7C" box="[293,355,225,248]" italics="true" pageId="5" pageNumber="162">sensu</emphasis>
Binder, 2008). Aperture crescent-shaped; aperture height ~4/5 its width; aperture width ~2/5 shell width. Peristome greatly reflected.
</paragraph>
<paragraph id="D421AD7B2718FFCBFF708D0FFAAAFEC4" blockId="5.[151,1436,151,608]" box="[199,1376,295,320]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF708D0FFE29FEC4" bold="true" box="[199,483,295,320]" pageId="5" pageNumber="162">Measurements (in mm).</emphasis>
BSPG
<date id="A0208BBB2718FFCBFD8F8D0FFD40FEBB" box="[568,650,295,320]" pageId="5" pageNumber="162" value="1959-02">1959 II</date>
16144 (
<figureCitation id="4CA5B1FE2718FFCBFD518D0FFC87FEC4" box="[742,845,295,320]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="5" pageNumber="162">Figs.6–9</figureCitation>
): 4¾ whorls; H = 5.0; D = 9.3; h = 3.3; d = 4.0.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02718FFCBFF708D64FB00FE0C" pageId="5" pageNumber="162" type="materials_examined">
<paragraph id="D421AD7B2718FFCBFF708D64FB00FE0C" blockId="5.[151,1436,151,608]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF708D64FD56FEE1" bold="true" box="[199,668,332,357]" pageId="5" pageNumber="162">Previous identification of the material.</emphasis>
Gall (1972: Nr. 18):
<taxonomicName id="139ED6F82718FFCBFC218D64FA52FEE1" authority="Klein" authorityName="Klein" box="[918,1432,332,357]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="subSpecies" species="giengensis" subGenus="Klikia" subSpecies="giengensis">
<emphasis id="E6EA71692718FFCBFC218D64FAF0FEE0" box="[918,1338,332,357]" italics="true" pageId="5" pageNumber="162">Klikia (Klikia) giengensis giengensis</emphasis>
(Klein)
</taxonomicName>
. Moser
<emphasis id="E6EA71692718FFCBFF528D59FEDFFE0C" box="[229,277,367,392]" italics="true" pageId="5" pageNumber="162">et al</emphasis>
. (2009b: Nr. 54, fig. 6B):
<taxonomicName id="139ED6F82718FFCBFD8B8D58FCDBFE03" authority="Prosoosia" authorityName="Prosoosia" box="[572,785,368,391]" class="Gastropoda" family="Helicodontidae" genus="Soosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692718FFCBFD8B8D58FCDBFE03" box="[572,785,368,391]" italics="true" pageId="5" pageNumber="162">Soosia (Prosoosia)</emphasis>
</taxonomicName>
cf.
<emphasis id="E6EA71692718FFCBFC8B8D47FC44FE0C" box="[828,910,367,392]" italics="true" pageId="5" pageNumber="162">godarti</emphasis>
Michaud
<emphasis id="E6EA71692718FFCBFBB78D59FBF4FE0C" box="[1024,1086,369,392]" italics="true" pageId="5" pageNumber="162">sensu</emphasis>
Sandberger.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02718FFCBFF708DBCFEA8FD49" pageId="5" pageNumber="162" type="discussion">
<paragraph id="D421AD7B2718FFCBFF708DBCFC27FDE4" blockId="5.[151,1436,151,608]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF708DBCFE86FE29" bold="true" box="[199,332,404,429]" pageId="5" pageNumber="162">Discussion.</emphasis>
The specimens from Sandelzhausen compares fittingly with
<taxonomicName id="139ED6F82718FFCBFBA88DBDFB65FE28" box="[1055,1199,405,428]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="species" species="coarctata">
<emphasis id="E6EA71692718FFCBFBA88DBDFB65FE28" box="[1055,1199,405,428]" italics="true" pageId="5" pageNumber="162">K. coarctata</emphasis>
</taxonomicName>
, a species from the Silvanaschichten of southwestern
<collectingCountry id="AC89EDEB2718FFCBFDA88D90FD4DFE54" box="[543,647,440,464]" name="Germany" pageId="5" pageNumber="162">Germany</collectingCountry>
(MN5–6; Klein 1853; Schlickum 1976). However, one of the main features of this species, a covered umbilicus (Klein 1853; Harzhauser &amp; Binder 2004), cannot be seen in the material from Sandelzhausen, since the single complete shell preserved shows a collapsed umbilical region. In any case, the degree in which the umbilicus is covered seems to be variable in some
<taxonomicName id="139ED6F82718FFCBFBFE8E0CFB44FDB9" box="[1097,1166,548,573]" class="Gastropoda" family="Elonidae" genus="Klikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692718FFCBFBFE8E0CFB44FDB9" box="[1097,1166,548,573]" italics="true" pageId="5" pageNumber="162">Klikia</emphasis>
</taxonomicName>
species (Harzhauser &amp; Binder 2004) and as such, it might not be such a useful diagnostic character.
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208E9CFEA8FD49" blockId="5.[151,354,692,717]" box="[151,354,692,717]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF208E9CFEA8FD49" bold="true" box="[151,354,692,717]" pageId="5" pageNumber="162">
Family
<taxonomicName id="139ED6F82718FFCBFF458E9CFEA8FD49" box="[242,354,692,717]" class="Gastropoda" family="Helicidae" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="family">Helicidae</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02718FFCBFF208ED4FDF7FB18" pageId="5" pageNumber="162" type="description">
<paragraph id="D421AD7B2718FFCBFF208ED4FE4AFC91" blockId="5.[151,384,764,789]" box="[151,384,764,789]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF208ED4FE4AFC91" bold="true" box="[151,384,764,789]" pageId="5" pageNumber="162">
Genus
<taxonomicName id="139ED6F82718FFCBFF5D8ED4FE4AFC91" authority="Held" authorityName="Held" box="[234,384,764,789]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="E6EA71692718FFCBFF5D8ED4FE8AFC91" bold="true" box="[234,320,764,789]" italics="true" pageId="5" pageNumber="162">Cepaea</emphasis>
Held
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208F6CFEE7FC04" blockId="5.[151,540,836,896]" pageId="5" pageNumber="162">
<taxonomicName id="139ED6F82718FFCBFF208F6CFDD6FCD9" authority="Deshayes, 1851" authorityName="Deshayes" authorityYear="1851" box="[151,540,836,861]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA71692718FFCBFF208F6CFDD6FCD9" bold="true" box="[151,540,836,861]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF208F6CFF26FCD9" bold="true" box="[151,236,836,861]" italics="true" pageId="5" pageNumber="162">Cepaea</emphasis>
cf.
<emphasis id="E6EA71692718FFCBFEA08F6CFEAAFCDF" bold="true" box="[279,352,836,859]" italics="true" pageId="5" pageNumber="162">eversa</emphasis>
Deshayes, 1851
</emphasis>
</taxonomicName>
(
<figureCitation id="4CA5B1FE2718FFCBFF288F4FFEECFC04" box="[159,294,871,896]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="5" pageNumber="162">Figs. 10–13</figureCitation>
)
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208F98FD8AFC60" blockId="5.[151,945,944,1181]" pageId="5" pageNumber="162">
<taxonomicName id="139ED6F82718FFCBFF208F98FE3AFC42" authority="Deshayes, 1851: 395" authorityName="Deshayes" authorityPageNumber="395" authorityYear="1851" box="[151,496,944,966]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA71692718FFCBFF208F98FEDCFC42" box="[151,278,944,966]" italics="true" pageId="5" pageNumber="162">Helix eversa</emphasis>
Deshayes, 1851: 395
</taxonomicName>
(pl. 1, figs. 5–7).
<taxonomicName id="139ED6F82718FFCBFF208FE7FDF6FC60" authority="Wenz, 1923a: 617" authorityName="Wenz" authorityPageNumber="617" authorityYear="1923" box="[151,572,974,997]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="subSpecies" species="eversa" subSpecies="eversa">
<emphasis id="E6EA71692718FFCBFF208FE7FEBFFC61" box="[151,373,975,997]" italics="true" pageId="5" pageNumber="162">Cepaea eversa eversa</emphasis>
: Wenz, 1923a: 617
</taxonomicName>
.
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208FC5FD4AFBC4" blockId="5.[151,945,944,1181]" pageId="5" pageNumber="162">
<taxonomicName id="139ED6F82718FFCBFF208FC5FEA8FB87" box="[151,354,1005,1027]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="species" species="asperula">
<emphasis id="E6EA71692718FFCBFF208FC5FEA8FB87" box="[151,354,1005,1027]" italics="true" pageId="5" pageNumber="162">Hemicycla asperula</emphasis>
</taxonomicName>
cf.
<emphasis id="E6EA71692718FFCBFE3E8FC5FDCCFB87" box="[393,518,1005,1027]" italics="true" pageId="5" pageNumber="162">leymerieana</emphasis>
: Gall, 1972: 10.
<taxonomicName id="139ED6F82718FFCBFF208824FEBBFBA6" box="[151,369,1036,1058]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="subSpecies" species="eversa" subSpecies="larteti">
<emphasis id="E6EA71692718FFCBFF208824FEBBFBA6" box="[151,369,1036,1058]" italics="true" pageId="5" pageNumber="162">Cepaea eversa larteti</emphasis>
</taxonomicName>
(in part): Gall, 1972: 10; Gall, 1973: 10 (pl.1, figs. 4–6).
<taxonomicName id="139ED6F82718FFCBFF208803FE49FBC4" box="[151,387,1066,1088]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="subSpecies" species="silvana" subSpecies="silvana">
<emphasis id="E6EA71692718FFCBFF208803FE49FBC4" box="[151,387,1066,1088]" italics="true" pageId="5" pageNumber="162">Cepaea silvana silvana</emphasis>
</taxonomicName>
(in part): Gall, 1972: 11.
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208862FDE0FBDB" blockId="5.[151,945,944,1181]" box="[151,554,1097,1119]" pageId="5" pageNumber="162">
<taxonomicName id="139ED6F82718FFCBFF208862FDECFBDB" authority="Hir &amp; Kokay, 2004: 89" authorityName="Hir &amp; Kokay" authorityPageNumber="89" authorityYear="2004" box="[151,550,1097,1119]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA71692718FFCBFF208862FEE7FBDB" box="[151,301,1098,1119]" italics="true" pageId="5" pageNumber="162">Cepaea eversa</emphasis>
: Hír &amp; Kókay, 2004: 89
</taxonomicName>
.
</paragraph>
<paragraph id="D421AD7B2718FFCBFF208841FDF7FB18" blockId="5.[151,945,944,1181]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF208841FECAFBFA" box="[151,256,1129,1150]" italics="true" pageId="5" pageNumber="162">
“
<taxonomicName id="139ED6F82718FFCBFF288841FF33FBFA" box="[159,249,1129,1150]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
”
</emphasis>
sp. 1. (in part): Moser
<emphasis id="E6EA71692718FFCBFE5D8841FDDCFBFA" box="[490,534,1128,1150]" italics="true" pageId="5" pageNumber="162">et al</emphasis>
., 2009b: 50.
<emphasis id="E6EA71692718FFCBFF2088AFFECAFB18" box="[151,256,1159,1180]" italics="true" pageId="5" pageNumber="162">
“
<taxonomicName id="139ED6F82718FFCBFF2888AFFF33FB18" box="[159,249,1159,1180]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="162" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
”
</emphasis>
sp. 2.: Moser
<emphasis id="E6EA71692718FFCBFE2688A0FE74FB18" box="[401,446,1158,1181]" italics="true" pageId="5" pageNumber="162">et al</emphasis>
., 2009b: 50.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02718FFCBFF2088E1FC63F931" pageId="5" pageNumber="162" type="materials_examined">
<paragraph id="D421AD7B2718FFCBFF2088E1FCF2F9A2" blockId="5.[151,1436,1225,2006]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF2088E1FE49FB66" bold="true" box="[151,387,1225,1250]" pageId="5" pageNumber="162">Material examined.</emphasis>
BSPG
<date id="A0208BBB2718FFCBFE6B88E1FDFBFB65" box="[476,561,1225,1250]" pageId="5" pageNumber="162" value="1959-02">1959 II</date>
460 (1 spcm.), 461 (1 spcm.), 16157 (1 spcm.), 16158 (1 spcm.), 16160 (1 spcm.), 16162 (1 spcm.), 16165 (1 spcm.), 16166 (1 spcm.), 16167 (1 spcm.), 16168 (1 spcm.), 17336 (1 spcm.), 17337 (1 spcm.), 17338 (1 spcm.), 17339 (2 spcm.), 17340 (1 spcm.), 17341 (1 spcm.), 17342 (1 spcm.), 17343 (1 spcm.), 17344 (1 spcm.), 17345 (1 spcm.), 17346 (1 spcm.), 17347 (1 spcm.), 17376 (1 spcm.), 17377 (1 spcm.), 17378 (1 spcm.), 17379 (1 spcm.), 17380 (1 spcm.), 17381 (1 spcm.), 17382 (1 spcm.), 17383 (1 spcm.), 17384 (1 spcm.), 17385 (1 spcm.), 17386 (1 spcm.), 17387 (1 spcm.), 17388 (1 spcm.), 17389 (1 spcm.), 17782 (1 spcm.), 17783 (1 spcm.), 17784 (1 spcm.), 17785 (1 spcm.), 17786 (1 spcm.), 17787 (2 spcm.), 17788 (1 spcm.), 17789 (1 spcm.), 17790 (1 spcm.), 17791 (1 spcm.), 17792 (1 spcm.), 17793 (1 spcm.), 17794 (1 spcm.), 17795 (1 spcm.), 17796 (1 spcm.), 17797 (1 spcm.), 17798 (1 spcm.), 17799 (1 spcm.), 17800 (1 spcm.), 17801 (1 spcm.), 17802 (1 spcm.), 17803 (1 spcm.), 17804 (2 spcm.), 17805 (1 spcm.).
</paragraph>
<paragraph id="D421AD7B2718FFCBFF708A19FC63F931" blockId="5.[151,1436,1225,2006]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF708A19FE30F9CE" bold="true" box="[199,506,1585,1610]" pageId="5" pageNumber="162">Stratigraphic occurrence.</emphasis>
Layers B1 (5 spcm.), B2 (5 spcm.), C1 (3 spcm.), C2 (1 spcm.), D1 (7 spcm.). Moreover,
<specimenCount id="C29866F22718FFCBFEA58A7EFE63F9EA" box="[274,425,1621,1646]" pageId="5" pageNumber="162" type="generic">37 specimens</specimenCount>
are from excavation sites for which no profile is available, but, based on their height in the sediment and their preservation, it is possible to infer the layer of origin for some: 1 is likely from layer C1, 1 from C3, 12 from D, 3 from either B2 or C1, and 7 are from either C3 or D.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF02718FFC8FF708AE9FF31FF50" lastPageId="6" lastPageNumber="163" pageId="5" pageNumber="162" type="description">
<paragraph id="D421AD7B2718FFCBFF708AE9FCACF852" blockId="5.[151,1436,1225,2006]" pageId="5" pageNumber="162">
<emphasis id="E6EA71692718FFCBFF708AE9FE93F95E" bold="true" box="[199,345,1729,1754]" pageId="5" pageNumber="162">Description.</emphasis>
Shell large, 4½ to 4¾ whorls, with depressed spire; shell length ~2/3 its width. Protoconch flattened, smooth; transition to teleoconch unclear. Teleoconch smooth, except for growth lines. Suture wellmarked. Body whorl slightly bent downwards near aperture; apertural region enlarged. Aperture prosocline, crescent-shaped, with strong callus; aperture length ~1/2 its width and ~1/2 shell length; aperture width ~1/2 shell width. Peristome markedly reflexed; thickened on basal and columellar regions. Umbilicus imperforate, with a marked callus formed by the peristome. On two specimens (BSPG
<date id="A0208BBB2718FFCBFC1C8B5DFBCBF809" box="[939,1025,1909,1934]" pageId="5" pageNumber="162" value="1959-02">1959 II</date>
16160 and 17339) it is possible to observe, under UV light, vestiges of three colored broad parallel spiral bands, the topmost one right below the carina, the other two regularly spaced on basal portion of whorl.
</paragraph>
<paragraph id="D421AD7B271BFFC8FF708CBFFF31FF50" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="163">
<emphasis id="E6EA7169271BFFC8FF708CBFFE28FF34" bold="true" box="[199,482,151,176]" pageId="6" pageNumber="163">Measurements (in mm).</emphasis>
BSPG
<date id="A0208BBB271BFFC8FD828CBFFD4DFF2B" box="[565,647,151,176]" pageId="6" pageNumber="163" value="1959-02">1959 II</date>
16162 (deformed specimen;
<figureCitation id="4CA5B1FE271BFFC8FC7B8CBFFB9EFF34" box="[972,1108,151,176]" captionStart="FIGURES 1 – 21. 1 – 2" captionStartId="3.[151,264,1790,1813]" captionTargetBox="[234,1349,193,1769]" captionTargetId="figure@3.[234,1350,193,1771]" captionTargetPageId="3" captionText="FIGURES 1 – 21. 1 – 2. Succinea minima (BSPG 1959 II 17471; H = 2.6 mm). 3 – 4. Palaeoglandina sp. (BSPG 1959 II 16145; H = 49.7 mm). 5. Testacella zelli (BSPG 1959 II 17689; H = 2.3 mm). 6 – 9. Klikia cf. c oarctata (BSPG 1959 II 16144; D = 9.3 mm). 10 – 13. Cepaea cf. eversa (BSPG 1959 II 16162; D = 26.0 mm). 14 – 17. Cepaea cf. sylvestrina (BSPG 1959 II 16156; D = 19.9 mm). 18 – 19. Tropidomphalus cf. incrassatus (BSPG 1959 II 456; D = 28.8 mm). 20 – 21. Tropidomphalus cf. incrassatus (BSPG 1959 II 457; D = 35.1 mm)." httpUri="https://zenodo.org/record/218327/files/figure.png" pageId="6" pageNumber="163">Figs. 10–13</figureCitation>
): H = 18.4; D = 26.0; h = 8.1; d = 14.3.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF0271BFFC8FF708CF7FBF6FEC4" pageId="6" pageNumber="163" type="materials_examined">
<paragraph id="D421AD7B271BFFC8FF708CF7FBF6FEC4" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="163">
<emphasis id="E6EA7169271BFFC8FF708CF7FD68FF7C" bold="true" box="[199,674,223,248]" pageId="6" pageNumber="163">Previous identification of the material.</emphasis>
Gall (1972: Nr. 19 and, in part, 20–21): respectively,
<taxonomicName id="139ED6F8271BFFC8FA938CF7FF33FE99" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="asperula">
<emphasis id="E6EA7169271BFFC8FA938CF7FF33FE99" italics="true" pageId="6" pageNumber="163">Hemicycla asperula</emphasis>
</taxonomicName>
cf.
<emphasis id="E6EA7169271BFFC8FE948D2CFE64FE99" box="[291,430,260,285]" italics="true" pageId="6" pageNumber="163">leymerieana</emphasis>
(Noulet),
<taxonomicName id="139ED6F8271BFFC8FD978D2DFCB7FE99" authority="Klein" authorityName="Klein" box="[544,893,260,285]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="silvana" subSpecies="silvana">
<emphasis id="E6EA7169271BFFC8FD978D2DFCEFFE99" box="[544,805,260,285]" italics="true" pageId="6" pageNumber="163">Cepaea silvana silvana</emphasis>
(Klein)
</taxonomicName>
and
<taxonomicName id="139ED6F8271BFFC8FC058D2DFAC6FE99" authority="Boissy" authorityName="Boissy" box="[946,1292,260,285]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="subSpecies" species="eversa" subSpecies="larteti">
<emphasis id="E6EA7169271BFFC8FC058D2DFB68FE99" box="[946,1186,260,285]" italics="true" pageId="6" pageNumber="163">Cepaea eversa larteti</emphasis>
(Boissy)
</taxonomicName>
. Moser
<emphasis id="E6EA7169271BFFC8FAD38D2DFA5FFE99" box="[1380,1429,260,285]" italics="true" pageId="6" pageNumber="163">et al</emphasis>
. (2009b: Nr. 64, in part, and 65): respectively,
<emphasis id="E6EA7169271BFFC8FD218D00FCC0FEBB" box="[662,778,296,319]" italics="true" pageId="6" pageNumber="163">
“
<taxonomicName id="139ED6F8271BFFC8FD288D00FCC9FEBB" box="[671,771,296,319]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
”
</emphasis>
sp. 1. and
<emphasis id="E6EA7169271BFFC8FC338D00FC32FEBB" box="[900,1016,296,319]" italics="true" pageId="6" pageNumber="163">
“
<taxonomicName id="139ED6F8271BFFC8FC3A8D00FC3BFEBB" box="[909,1009,296,319]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="genus">Cepaea</taxonomicName>
”
</emphasis>
sp. 2.
</paragraph>
</subSubSection>
<subSubSection id="9C84FEF0271BFFC8FF708D64FDF4FD9C" pageId="6" pageNumber="163" type="discussion">
<paragraph id="D421AD7B271BFFC8FF708D64FDF4FD9C" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="163">
<emphasis id="E6EA7169271BFFC8FF708D64FE86FEE1" bold="true" box="[199,332,332,357]" pageId="6" pageNumber="163">Discussion.</emphasis>
The specimens from Sandelzhausen compare well with
<taxonomicName id="139ED6F8271BFFC8FC718D65FBFBFEE0" box="[966,1073,333,356]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA7169271BFFC8FC718D65FBFBFEE0" box="[966,1073,333,356]" italics="true" pageId="6" pageNumber="163">C. eversa</emphasis>
</taxonomicName>
, but are always larger, with half whorl more than typical
<taxonomicName id="139ED6F8271BFFC8FE1A8D58FDD2FE0C" box="[429,536,368,392]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA7169271BFFC8FE1A8D58FDD2FE0C" box="[429,536,368,392]" italics="true" pageId="6" pageNumber="163">C. eversa</emphasis>
</taxonomicName>
specimens. Moreover, the peristome of the present specimens is more reflected and only thickened on the basal and columellar regions, while typical
<taxonomicName id="139ED6F8271BFFC8FC1C8DBDFBDFFE28" box="[939,1045,405,428]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA7169271BFFC8FC1C8DBDFBDFFE28" box="[939,1045,405,428]" italics="true" pageId="6" pageNumber="163">C. eversa</emphasis>
</taxonomicName>
has the entire peristome thickened and not much reflected. Nevertheless,
<taxonomicName id="139ED6F8271BFFC8FDF88D90FD70FE54" box="[591,698,440,464]" class="Gastropoda" family="Helicidae" genus="Cepaea" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="163" phylum="Mollusca" rank="species" species="eversa">
<emphasis id="E6EA7169271BFFC8FDF88D90FD70FE54" box="[591,698,440,464]" italics="true" pageId="6" pageNumber="163">C. eversa</emphasis>
</taxonomicName>
seems to be a very variable species regarding shell morphology (Hír &amp; Kókay 2004). It is also a very frequent species in Middle Miocene (MN 5–7) deposits of Central and Western Europe (Hír &amp; Kókay 2004).
</paragraph>
</subSubSection>
</treatment>
</document>
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