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<document id="20F7249664FFDCAD1A4C6B585FF7CACE" ID-DOI="10.11646/zootaxa.3736.2.5" ID-GBIF-Dataset="8f6fcaa0-5673-4636-a896-ce0ede2c6c42" ID-ISSN="1175-5326" ID-Zenodo-Dep="219269" ID-ZooBank="9E9A6BEF-EBD9-4B06-9C7B-99E2500D7DC6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460697525616" checkinUser="plazi" docAuthor="Enghoff, Henrik &amp; Reboleira, Ana Sofia P. S." docDate="2013" docId="2578E112FFB4FE28FF5B9303FC00FCE1" docLanguage="en" docName="zt03736p186.pdf" docOrigin="Zootaxa 3736 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scutogona minor Enghoff &amp; Reboleira, 2013, n.sp." docType="treatment" docUuid="095A41F2-DD3A-43F6-8A25-07342A1277CC" docUuidSource="ZooBank" docVersion="11" lastPageNumber="184" masterDocId="D941996AFFB0FE21FFCC9251FFFDFFBD" masterDocTitle="A new cave-dwelling millipede of the genus Scutogona from central Portugal (Diplopoda, Chordeumatida, Chamaesomatidae)" masterLastPageNumber="186" masterPageNumber="175" pageNumber="179" updateTime="1698353964119" updateUser="plazi">
<mods:mods id="CCBF0A25EE538B2186CBDDBCD8ABA210" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="779C63AC2E2E04F076B6B9F0DB1DD961">
<mods:title id="668A91B4CD41D38A68B27602065D9439">A new cave-dwelling millipede of the genus Scutogona from central Portugal (Diplopoda, Chordeumatida, Chamaesomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="5AEED15AA640D980CD3B1551D342307E" type="personal">
<mods:role id="BD086D932B6674BADDF7C74CD915B4E8">
<mods:roleTerm id="1793A388E8E28DAC7EA2A74F649DC2F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A8C56E4A550098474E044D791B49423">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:name id="79746955F6D5F92A7965AC9A0C846341" type="personal">
<mods:role id="3D565D9F54757074867C7AC8F435594C">
<mods:roleTerm id="A4B65766F1E726045A7DAA797F947EB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B38C3564D03A0233B9A1BCAE8FE7A955">Reboleira, Ana Sofia P. S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7EBF4ADD1691E0F7BD59411CD582695E">text</mods:typeOfResource>
<mods:relatedItem id="B75E8BA88B16927C4981C21ACA8D12A9" type="host">
<mods:titleInfo id="05096956DEFE5B4DDD0B61C98173E981">
<mods:title id="D74FC9A54137B9912401116125AF9204">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF055224185975A8A363449A15A29A14">
<mods:date id="C60E01CA6BD8A1DF4734A3CB7E629E53">2013</mods:date>
<mods:detail id="86C372C3A08DD73D13AE0B42FDCE0188" type="volume">
<mods:number id="E27168FACC4CB49527FB952385088555">3736</mods:number>
</mods:detail>
<mods:detail id="C7F42E61B0C7E1469709951F585FB194" type="issue">
<mods:number id="F18F344A52119CCAC38E55FFEAAEE962">2</mods:number>
</mods:detail>
<mods:extent id="7F39186E8D6D4F61FB384848CD593612" unit="page">
<mods:start id="45489364D9A75C5E986E1A62C981F01B">175</mods:start>
<mods:end id="40481CB99466CA8D69643E9448434D29">186</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A3A033D81EC08A5B20EC4DE05B5555C9">journal article</mods:classification>
<mods:identifier id="145955E118883FC5053206375B796422" type="DOI">10.11646/zootaxa.3736.2.5</mods:identifier>
<mods:identifier id="D863EF9024889E0FFE997A859BC098C5" type="GBIF-Dataset">8f6fcaa0-5673-4636-a896-ce0ede2c6c42</mods:identifier>
<mods:identifier id="54BA5002F2B8D1F76F6B15F955048BF7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0F493AEF9A3E9F1EB13B3928763EB03D" type="Zenodo-Dep">219269</mods:identifier>
<mods:identifier id="E1E1D66EB5280F8F96E2EEDA97814DCE" type="ZooBank">9E9A6BEF-EBD9-4B06-9C7B-99E2500D7DC6</mods:identifier>
</mods:mods>
<treatment id="2578E112FFB4FE28FF5B9303FC00FCE1" ID-DOI="http://doi.org/10.5281/zenodo.3506131" ID-GBIF-Taxon="119582140" ID-Zenodo-Dep="3506131" LSID="urn:lsid:zoobank.org:act:095A41F2-DD3A-43F6-8A25-07342A1277CC" httpUri="http://treatment.plazi.org/id/2578E112FFB4FE28FF5B9303FC00FCE1" lastPageId="9" lastPageNumber="184" pageId="4" pageNumber="179">
<subSubSection id="E5CB038FFFB4FE25FF5B9303FEF0FE29" pageId="4" pageNumber="179" type="nomenclature">
<paragraph id="AD6E5004FFB4FE25FF5B9303FE57FED1" blockId="4.[151,1436,338,2037]" box="[151,426,338,364]" pageId="4" pageNumber="179">
<heading id="F626E768FFB4FE25FF5B9303FE57FED1" bold="true" box="[151,426,338,364]" fontSize="11" level="1" pageId="4" pageNumber="179" reason="1">
<emphasis id="9FA58C16FFB4FE25FF5B9303FE57FED1" bold="true" box="[151,426,338,364]" pageId="4" pageNumber="179">
<taxonomicName id="6AD12B87FFB4FE25FF5B9303FE95FED1" ID-CoL="84D4F" box="[151,360,338,364]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB4FE25FF5B9303FE95FED1" bold="true" box="[151,360,338,364]" italics="true" pageId="4" pageNumber="179">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB4FE25FEA39303FE57FED1" box="[367,426,338,364]" pageId="4" pageNumber="179" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF5B932AFEF0FE29" blockId="4.[151,1436,338,2037]" box="[151,269,379,404]" pageId="4" pageNumber="179">
<figureCitation id="35EA4C81FFB4FE25FF5B932AFEF4FE29" box="[151,265,379,404]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 10" captionStart-2="FIGURES 11 – 12" captionStart-3="FIGURE 13" captionStartId-0="5.[151,264,1947,1969]" captionStartId-1="6.[151,264,1164,1186]" captionStartId-2="7.[151,264,1944,1966]" captionStartId-3="8.[151,250,1110,1132]" captionTargetBox-0="[239,1356,183,1921]" captionTargetBox-1="[151,1436,193,1142]" captionTargetBox-2="[224,1367,195,1925]" captionTargetBox-3="[340,1218,193,1089]" captionTargetId-0="figure@5.[239,1357,183,1922]" captionTargetId-1="figure@6.[151,1436,193,1142]" captionTargetId-2="figure@7.[224,1367,195,1925]" captionTargetId-3="figure@8.[340,1247,193,1089]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURES 1 – 5. Scutogona minor n. sp., female paratype from Gruta da Arrifana, SEM images. 1. Front end. 2. Head. 3. Lateral hump of midbody pleurotergum, right side. 4. Spinnerets. 5. Close-up of left spinneret, showing exuded thread. — Scales 100 µm (1, 2); 10 µm (3, 4); 1 µm (5)." captionText-1="FIGURES 6 – 10. Scutogona minor n. sp., male paratype from Gruta da Arrifana, anterior gonopods (P 8), SEM images. 6. Anterior view. 7. Oblique anterior-apical view. 8. Apical view. 9. Posterior view. 10. Lateral view (from right side). — A: right angiocoxite, F: flagellum, KP: two-pronged process, KL: laminar extension, T: left telopodite. — Scales 10 µm." captionText-2="FIGURES 11 – 12. Scutogona minor n. sp., paratypes from Gruta da Arrifana, SEM images. 11. male, P 9 – 11. P 9: left posterior gonopod, P 10: left first post-gonopodal leg, P 11: right second post-gonopodal leg with coxal hook (arrow). — 12. Female, vulvae seen from the left. O: operculum, PV: post-vulvar organ, V: lateral valve. — Scales 10 µm." captionText-3="FIGURE 13. Scutogona minor n. sp., female paratype from Gruta da Arrifana, vulvae in situ behind second pair of legs, posterior view. Insert: semi-diagrammatic drawing of left vulva, showing post-vulvar organ (PV) valves (V) with setae, and the internal separation of the two valves (dotted line). — Scale 50 µm." httpUri-0="https://zenodo.org/record/219270/files/figure.png" httpUri-1="https://zenodo.org/record/219271/files/figure.png" httpUri-2="https://zenodo.org/record/219272/files/figure.png" httpUri-3="https://zenodo.org/record/219273/files/figure.png" pageId="4" pageNumber="179">Figs 1–13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5CB038FFFB4FE25FF5B93FDFD15FDE8" pageId="4" pageNumber="179" type="materials_examined">
<paragraph id="AD6E5004FFB4FE25FF5B93FDFD15FDE8" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF5B93FDFE1CFE78" bold="true" box="[151,481,428,453]" pageId="4" pageNumber="179">
Material studied:
<typeStatus id="726AEEA6FFB4FE25FEB893FDFE1CFE78" box="[372,481,428,453]" pageId="4" pageNumber="179" type="holotype">Holotype</typeStatus>
</emphasis>
♂
<collectingCountry id="D5C61094FFB4FE25FDC793FCFD5FFE78" box="[523,674,429,453]" name="Portugal" pageId="4" pageNumber="179">PORTUGAL</collectingCountry>
, Sicó massif, Gruta da Arrifana (
<geoCoordinate id="C8E536C3FFB4FE25FBE493FDFB25FE78" box="[1064,1240,428,453]" direction="north" orientation="latitude" pageId="4" pageNumber="179" precision="1" value="40.08731">40°05’14.32”N</geoCoordinate>
;
<geoCoordinate id="C8E536C3FFB4FE25FB2B93FDFA72FE78" box="[1255,1423,428,453]" direction="west" orientation="longitude" pageId="4" pageNumber="179" precision="1" value="-8.516389">8°30’59.00”W</geoCoordinate>
),
<date id="D96F76C4FFB4FE25FF5B9381FEF6FE55" box="[151,267,464,489]" pageId="4" pageNumber="179" value="2013-01-13">13.i. 2013</date>
, S. Reboleira leg. (ZMUC).—
<emphasis id="9FA58C16FFB4FE25FDAC9381FD25FE54" bold="true" box="[608,728,464,489]" pageId="4" pageNumber="179">
<typeStatus id="726AEEA6FFB4FE25FDAC9381FD25FE54" box="[608,728,464,489]" pageId="4" pageNumber="179" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="BBD79B8DFFB4FE25FD249381FCE9FE55" box="[744,788,464,488]" pageId="4" pageNumber="179" type="generic">2 ♂</specimenCount>
,
<specimenCount id="BBD79B8DFFB4FE25FCED9380FCB0FE55" box="[801,845,465,488]" pageId="4" pageNumber="179" type="generic">7 ♂</specimenCount>
,
<specimenCount id="BBD79B8DFFB4FE25FC969381FC61FE54" box="[858,924,464,489]" pageId="4" pageNumber="179" type="generic">1 juv.</specimenCount>
, same data (ZMUC,
<specimenCount id="BBD79B8DFFB4FE25FB4B9381FB49FE55" box="[1159,1204,464,488]" pageId="4" pageNumber="179" type="generic">1 ♂</specimenCount>
, 1 ♀ DZUL).
<specimenCount id="BBD79B8DFFB4FE25FA999381FA7CFE55" box="[1365,1409,464,488]" pageId="4" pageNumber="179" type="generic">1 ♂</specimenCount>
, 2 ♀
<collectingCountry id="D5C61094FFB4FE25FF7893A4FEB6FDB0" box="[180,331,501,525]" name="Portugal" pageId="4" pageNumber="179">PORTUGAL</collectingCountry>
, Sicó, Gruta de Santa Maria da Estrela (
<geoCoordinate id="C8E536C3FFB4FE25FCD893A5FC43FDB0" box="[788,958,500,525]" direction="north" orientation="latitude" pageId="4" pageNumber="179" precision="1" value="39.994762">39°59'41.15&quot;N</geoCoordinate>
,
<geoCoordinate id="C8E536C3FFB4FE25FC1F93A5FB8BFDB0" box="[979,1142,500,525]" direction="west" orientation="longitude" pageId="4" pageNumber="179" precision="1" value="-8.549834">8°32'59.40&quot;W</geoCoordinate>
),
<date id="D96F76C4FFB4FE25FB4693A5FAF9FDB0" box="[1162,1284,500,525]" pageId="4" pageNumber="179" value="2009-06-11">11.vi.2009</date>
, S. Reboleira leg. (ZMUC),
<specimenCount id="BBD79B8DFFB4FE25FEF09049FE66FD8C" box="[316,411,536,561]" pageId="4" pageNumber="179" type="generic">1 subad.</specimenCount>
1 ♀ same data but,
<date id="D96F76C4FFB4FE25FD499049FCF1FD8C" box="[645,780,536,561]" pageId="4" pageNumber="179" value="2009-08-29">29.viii.2009</date>
(ZMUC).
<specimenCount id="BBD79B8DFFB4FE25FC449049FC4BFD8D" box="[904,950,536,560]" pageId="4" pageNumber="179" type="generic">1 ♂</specimenCount>
, Sicó, Gruta da Cerâmica (
<geoCoordinate id="C8E536C3FFB4FE25FB219049FA6AFD8C" box="[1261,1431,536,561]" direction="north" orientation="latitude" pageId="4" pageNumber="179" precision="1" value="39.92701">39°55'37.23&quot;N</geoCoordinate>
,
<geoCoordinate id="C8E536C3FFB4FE25FF5B906CFED1FDE8" box="[151,300,573,597]" direction="west" orientation="longitude" pageId="4" pageNumber="179" precision="1" value="-8.517775">8°31'3.99&quot;W</geoCoordinate>
),
<date id="D96F76C4FFB4FE25FEF3906CFE4DFDE8" box="[319,432,573,597]" pageId="4" pageNumber="179" value="2013-05-13">13.v.2013</date>
, S. Reboleira leg. (ZMUC).
</paragraph>
</subSubSection>
<subSubSection id="E5CB038FFFB4FE25FF0B9031FAA5FC25" pageId="4" pageNumber="179" type="diagnosis">
<paragraph id="AD6E5004FFB4FE25FF0B9031FCEAFDC4" blockId="4.[151,1436,338,2037]" box="[199,791,608,633]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF0B9031FEBEFDC4" bold="true" box="[199,323,608,633]" pageId="4" pageNumber="179">Diagnosis:</emphasis>
A species of
<taxonomicName id="6AD12B87FFB4FE25FE109030FDAFFDC5" box="[476,594,609,632]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB4FE25FE109030FDAFFDC5" box="[476,594,609,632]" italics="true" pageId="4" pageNumber="179">Scutogona</emphasis>
</taxonomicName>
, characterized by
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B90D4FC79FD20" blockId="4.[151,1436,338,2037]" box="[199,900,644,669]" pageId="4" pageNumber="179">
- small size (length of male
<quantity id="6A29FDE1FFB4FE25FE3390D5FDA8FD21" box="[511,597,644,669]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="179" unit="mm" value="5.5">5.5 mm</quantity>
vs.
<quantity id="6A29FDE1FFB4FE25FD4F90D4FD14FD21" box="[643,745,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="4" pageNumber="179" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7-12 mm</quantity>
in other spp.)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B90F8FE07FD7D" blockId="4.[151,1436,338,2037]" box="[199,506,680,705]" pageId="4" pageNumber="179">
- lack of eyes (≠
<taxonomicName id="6AD12B87FFB4FE25FEB090F9FE0DFD7C" box="[380,496,680,705]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="oculinigra">
<emphasis id="9FA58C16FFB4FE25FEB090F9FE0DFD7C" box="[380,496,680,705]" italics="true" pageId="4" pageNumber="179">oculinigra</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B909CFD8AFD58" blockId="4.[151,1436,338,2037]" box="[199,631,716,741]" pageId="4" pageNumber="179">
- dorsum of body arched (≠
<taxonomicName id="6AD12B87FFB4FE25FE36909DFD92FD58" box="[506,623,716,741]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="oculinigra">
<emphasis id="9FA58C16FFB4FE25FE36909DFD92FD58" box="[506,623,716,741]" italics="true" pageId="4" pageNumber="179">oculinigra</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B90A0FD0AFCB5" blockId="4.[151,1436,338,2037]" box="[199,759,752,777]" pageId="4" pageNumber="179">
- 3+3 macrochaetae on each body ring (≠
<taxonomicName id="6AD12B87FFB4FE25FD5890A1FD13FCB4" box="[660,750,752,777]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="9FA58C16FFB4FE25FD5890A1FD13FCB4" box="[660,750,752,777]" italics="true" pageId="4" pageNumber="179">jeanneli</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9144FD36FC90" blockId="4.[151,1436,338,2037]" box="[199,715,788,813]" pageId="4" pageNumber="179">
- pregonopodal legs unmodified (≠
<taxonomicName id="6AD12B87FFB4FE25FD819145FD3FFC90" box="[589,706,788,813]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="oculinigra">
<emphasis id="9FA58C16FFB4FE25FD819145FD3FFC90" box="[589,706,788,813]" italics="true" pageId="4" pageNumber="179">oculinigra</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9168FC9BFCED" blockId="4.[151,1436,338,2037]" box="[199,870,824,849]" pageId="4" pageNumber="179">
- each angiocoxite with a deep longitudinal furrow (≠
<taxonomicName id="6AD12B87FFB4FE25FCD19168FCA1FCED" box="[797,860,825,848]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="9FA58C16FFB4FE25FCD19168FCA1FCED" box="[797,860,825,848]" italics="true" pageId="4" pageNumber="179">vivesi</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B910CFC8BFCC8" blockId="4.[151,1436,338,2037]" box="[199,886,860,885]" pageId="4" pageNumber="179">
- angiocoxites clearly overreaching syncolpocoxite (≠
<taxonomicName id="6AD12B87FFB4FE25FCED910CFC90FCC9" box="[801,877,861,884]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="9FA58C16FFB4FE25FCED910CFC90FCC9" box="[801,877,861,884]" italics="true" pageId="4" pageNumber="179">mutica</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B91D0FAA5FC25" blockId="4.[151,1436,338,2037]" box="[199,1368,896,921]" pageId="4" pageNumber="179">
- syncolpocoxite mesally with a pair of caudad, broadly rounded lamellar lobes (≠
<taxonomicName id="6AD12B87FFB4FE25FB9191D1FB2FFC24" box="[1117,1234,896,921]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="oculinigra">
<emphasis id="9FA58C16FFB4FE25FB9191D1FB2FFC24" box="[1117,1234,896,921]" italics="true" pageId="4" pageNumber="179">oculinigra</emphasis>
</taxonomicName>
and
<taxonomicName id="6AD12B87FFB4FE25FAC491D0FAB5FC25" box="[1288,1352,897,920]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="9FA58C16FFB4FE25FAC491D0FAB5FC25" box="[1288,1352,897,920]" italics="true" pageId="4" pageNumber="179">vivesi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E5CB038FFFB4FE25FF0B91F5FBBFFC5C" pageId="4" pageNumber="179" type="etymology">
<paragraph id="AD6E5004FFB4FE25FF0B91F5FBBFFC5C" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF0B91F5FEACFC00" bold="true" box="[199,337,932,957]" pageId="4" pageNumber="179">Etymology:</emphasis>
<taxonomicName id="6AD12B87FFB4FE25FE9591F4FE61FC01" box="[345,412,933,956]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9FA58C16FFB4FE25FE9591F4FE61FC01" box="[345,412,933,956]" italics="true" pageId="4" pageNumber="179">minor</emphasis>
</taxonomicName>
is a Latin comparative adjective. All
<taxonomicName id="6AD12B87FFB4FE25FC8891F4FC47FC01" box="[836,954,933,956]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB4FE25FC8891F4FC47FC01" box="[836,954,933,956]" italics="true" pageId="4" pageNumber="179">Scutogona</emphasis>
</taxonomicName>
species are small, and whereas
<taxonomicName id="6AD12B87FFB4FE25FAEE91F4FA7FFC01" box="[1314,1410,933,956]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9FA58C16FFB4FE25FAEE91F4FA7FFC01" box="[1314,1410,933,956]" italics="true" pageId="4" pageNumber="179">S. minor</emphasis>
</taxonomicName>
is smaller than the other known species, a still smaller one may be found in the future.
</paragraph>
</subSubSection>
<subSubSection id="E5CB038FFFB4FE25FF0B91BDFDFEF849" pageId="4" pageNumber="179" type="description">
<paragraph id="AD6E5004FFB4FE25FF0B91BDFD5DFB95" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF0B91BDFEA6FBB8" bold="true" box="[199,347,1004,1029]" pageId="4" pageNumber="179">Description:</emphasis>
Males and females completely unpigmented, with 29 pleurotergites (“30 segments”), body length
<quantity id="6A29FDE1FFB4FE25FF5B9641FF0BFB95" box="[151,246,1040,1065]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="179" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
, vertical body diameter
<quantity id="6A29FDE1FFB4FE25FDC89641FD61FB95" box="[516,668,1040,1065]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="4" pageNumber="179" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.45–0.5 mm</quantity>
.
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9665FE0FFB28" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
Head (
<figureCitation id="35EA4C81FFB4FE25FED09665FE97FBF0" box="[284,362,1076,1101]" captionStart="FIGURES 1 – 5" captionStartId="5.[151,264,1947,1969]" captionTargetBox="[239,1356,183,1921]" captionTargetId="figure@5.[239,1357,183,1922]" captionTargetPageId="5" captionText="FIGURES 1 – 5. Scutogona minor n. sp., female paratype from Gruta da Arrifana, SEM images. 1. Front end. 2. Head. 3. Lateral hump of midbody pleurotergum, right side. 4. Spinnerets. 5. Close-up of left spinneret, showing exuded thread. — Scales 100 µm (1, 2); 10 µm (3, 4); 1 µm (5)." httpUri="https://zenodo.org/record/219270/files/figure.png" pageId="4" pageNumber="179">Fig. 2</figureCitation>
) densely pilose, labrum tridentate, mandibular stipites (“cheeks”) subglobular, strongly protruding. Antennae stouter than in other species, ”antennal club” (“massue antennaire” = antennomeres 5+6+7– 8) ca. 4 times as long as broad.
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B96F1FE26FAD0" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
Body subcylindrical, slightly tapering in both ends, lateral humps (
<figureCitation id="35EA4C81FFB4FE25FC0B96F1FBF6FB05" box="[967,1035,1184,1209]" captionStart="FIGURES 1 – 5" captionStartId="5.[151,264,1947,1969]" captionTargetBox="[239,1356,183,1921]" captionTargetId="figure@5.[239,1357,183,1922]" captionTargetPageId="5" captionText="FIGURES 1 – 5. Scutogona minor n. sp., female paratype from Gruta da Arrifana, SEM images. 1. Front end. 2. Head. 3. Lateral hump of midbody pleurotergum, right side. 4. Spinnerets. 5. Close-up of left spinneret, showing exuded thread. — Scales 100 µm (1, 2); 10 µm (3, 4); 1 µm (5)." httpUri="https://zenodo.org/record/219270/files/figure.png" pageId="4" pageNumber="179">Fig. 3</figureCitation>
) situated high up on the flank, but leaving dorsum of body arched. Surface with polygonal microsculpture (
<figureCitation id="35EA4C81FFB4FE25FC209695FBCEFB60" box="[1004,1075,1220,1245]" captionStart="FIGURES 1 – 5" captionStartId="5.[151,264,1947,1969]" captionTargetBox="[239,1356,183,1921]" captionTargetId="figure@5.[239,1357,183,1922]" captionTargetPageId="5" captionText="FIGURES 1 – 5. Scutogona minor n. sp., female paratype from Gruta da Arrifana, SEM images. 1. Front end. 2. Head. 3. Lateral hump of midbody pleurotergum, right side. 4. Spinnerets. 5. Close-up of left spinneret, showing exuded thread. — Scales 100 µm (1, 2); 10 µm (3, 4); 1 µm (5)." httpUri="https://zenodo.org/record/219270/files/figure.png" pageId="4" pageNumber="179">Fig. 3</figureCitation>
). Each pleurotergum with 3+3 macrochaetae situated on lateral humps, angle between them (MA) 120–130° on midbody rings, 160° on posterior rings. Distance between external and middle macrochaetae = distance between middle and internal ones (i.e., CIX (midbody) = 1), = ~ 1/3 of distance between middorsal suture and internal macrochaeta (i.e., MIX (midbody) = 3), = 1/3-of macrochaeta length.
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9728FCA3FA2C" blockId="4.[151,1436,338,2037]" box="[199,862,1400,1425]" pageId="4" pageNumber="179">Legs slightly (10–15%) longer than vertical body diameter.</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B97CDFE27FA08" blockId="4.[151,1436,338,2037]" box="[199,474,1436,1461]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF0B97CDFE27FA08" bold="true" box="[199,474,1436,1461]" pageId="4" pageNumber="179">Male sexual characters</emphasis>
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9791FDDBFA64" blockId="4.[151,1436,338,2037]" box="[199,550,1472,1497]" pageId="4" pageNumber="179">Pregonopodal legs unmodified.</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B97B5FC9BF90C" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
Anterior gonopods (P8) (
<figureCitation id="35EA4C81FFB4FE25FE2697B5FD9DFA40" box="[490,608,1508,1533]" captionStart="FIGURES 6 – 10" captionStartId="6.[151,264,1164,1186]" captionTargetBox="[151,1436,193,1142]" captionTargetId="figure@6.[151,1436,193,1142]" captionTargetPageId="6" captionText="FIGURES 6 – 10. Scutogona minor n. sp., male paratype from Gruta da Arrifana, anterior gonopods (P 8), SEM images. 6. Anterior view. 7. Oblique anterior-apical view. 8. Apical view. 9. Posterior view. 10. Lateral view (from right side). — A: right angiocoxite, F: flagellum, KP: two-pronged process, KL: laminar extension, T: left telopodite. — Scales 10 µm." httpUri="https://zenodo.org/record/219271/files/figure.png" pageId="4" pageNumber="179">Figs 6–10</figureCitation>
): Angiocoxites (A) fused only at their base, curving backwards, covering tip of syncolpocoxite, each angiocoxite with a deep longitudinal furrow. Syncolpocoxite with a pair of stout, twopronged processes (KP) originating laterally and pointing disto-mesad, mesally with a pair of posterior laminar extensions (KL) (probably homologues of the ‘sac’ described by Ribaut (1913) for
<taxonomicName id="6AD12B87FFB4FE25FB829400FB4AF9D5" box="[1102,1207,1617,1640]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="9FA58C16FFB4FE25FB829400FB4AF9D5" box="[1102,1207,1617,1640]" italics="true" pageId="4" pageNumber="179">S. mutica</emphasis>
</taxonomicName>
and the ‘lame paire pectiné’ described by Mauriès &amp; Vicente (1977) for
<taxonomicName id="6AD12B87FFB4FE25FD2E9424FCC0F931" box="[738,829,1653,1676]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="9FA58C16FFB4FE25FD2E9424FCC0F931" box="[738,829,1653,1676]" italics="true" pageId="4" pageNumber="179">S. vivesi</emphasis>
</taxonomicName>
. Telopodites (T) long, slender (but not as long as in
<taxonomicName id="6AD12B87FFB4FE25FA4B9424FF2AF90D" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="9FA58C16FFB4FE25FA4B9424FF2AF90D" italics="true" pageId="4" pageNumber="179">S. vivesi</emphasis>
</taxonomicName>
), with a terminal pore. Flagella (F) as in the other species.
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B94EDFDF1F944" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
Posterior gonopods (P9, ‘paragonopodes’) (
<figureCitation id="35EA4C81FFB4FE25FD2F94EDFCC2F968" box="[739,831,1724,1749]" captionStart="FIGURES 11 – 12" captionStartId="7.[151,264,1944,1966]" captionTargetBox="[224,1367,195,1925]" captionTargetId="figure@7.[224,1367,195,1925]" captionTargetPageId="7" captionText="FIGURES 11 – 12. Scutogona minor n. sp., paratypes from Gruta da Arrifana, SEM images. 11. male, P 9 – 11. P 9: left posterior gonopod, P 10: left first post-gonopodal leg, P 11: right second post-gonopodal leg with coxal hook (arrow). — 12. Female, vulvae seen from the left. O: operculum, PV: post-vulvar organ, V: lateral valve. — Scales 10 µm." httpUri="https://zenodo.org/record/219272/files/figure.png" pageId="4" pageNumber="179">Fig. 11</figureCitation>
) with both telopodomeres of ca. equal length, telopodomeres without tubercles.
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9555FF02F8FD" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
First and second pairs of post-gonopodal legs (P10 and P11) with coxal pores, P11 with a hook next to the pore (
<figureCitation id="35EA4C81FFB4FE25FF539579FF0CF8FD" box="[159,241,1832,1857]" captionStart="FIGURES 11 – 12" captionStartId="7.[151,264,1944,1966]" captionTargetBox="[224,1367,195,1925]" captionTargetId="figure@7.[224,1367,195,1925]" captionTargetPageId="7" captionText="FIGURES 11 – 12. Scutogona minor n. sp., paratypes from Gruta da Arrifana, SEM images. 11. male, P 9 – 11. P 9: left posterior gonopod, P 10: left first post-gonopodal leg, P 11: right second post-gonopodal leg with coxal hook (arrow). — 12. Female, vulvae seen from the left. O: operculum, PV: post-vulvar organ, V: lateral valve. — Scales 10 µm." httpUri="https://zenodo.org/record/219272/files/figure.png" pageId="4" pageNumber="179">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B951DFE09F8D8" blockId="4.[151,1436,338,2037]" box="[199,500,1868,1893]" pageId="4" pageNumber="179">
<emphasis id="9FA58C16FFB4FE25FF0B951DFE09F8D8" bold="true" box="[199,500,1868,1893]" pageId="4" pageNumber="179">Female sexual characters</emphasis>
</paragraph>
<paragraph id="AD6E5004FFB4FE25FF0B9521FDFEF849" blockId="4.[151,1436,338,2037]" pageId="4" pageNumber="179">
Vulvae (
<figureCitation id="35EA4C81FFB4FE25FEE29521FE4AF834" box="[302,439,1904,1929]" captionStart-0="FIGURES 11 – 12" captionStart-1="FIGURE 13" captionStartId-0="7.[151,264,1944,1966]" captionStartId-1="8.[151,250,1110,1132]" captionTargetBox-0="[224,1367,195,1925]" captionTargetBox-1="[340,1218,193,1089]" captionTargetId-0="figure@7.[224,1367,195,1925]" captionTargetId-1="figure@8.[340,1247,193,1089]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 11 – 12. Scutogona minor n. sp., paratypes from Gruta da Arrifana, SEM images. 11. male, P 9 – 11. P 9: left posterior gonopod, P 10: left first post-gonopodal leg, P 11: right second post-gonopodal leg with coxal hook (arrow). — 12. Female, vulvae seen from the left. O: operculum, PV: post-vulvar organ, V: lateral valve. — Scales 10 µm." captionText-1="FIGURE 13. Scutogona minor n. sp., female paratype from Gruta da Arrifana, vulvae in situ behind second pair of legs, posterior view. Insert: semi-diagrammatic drawing of left vulva, showing post-vulvar organ (PV) valves (V) with setae, and the internal separation of the two valves (dotted line). — Scale 50 µm." httpUri-0="https://zenodo.org/record/219272/files/figure.png" httpUri-1="https://zenodo.org/record/219273/files/figure.png" pageId="4" pageNumber="179">Figs 12–13</figureCitation>
) juxtaposed, operculum (O) ca. same length as valves, its distal margin slightly emarginated and with 2+2 setae; mesal and lateral valves (V) of same length, lateral valve with 5 setae, mesal valve with 4–7 setae; internal limit between valves strongly sclerotized, with a deep laterad notch. Post-vulvar organ (PV) with a short mesal process.
</paragraph>
</subSubSection>
<caption id="F9AE008CFFB5FE24FF5B95CAFE12F84C" httpUri="https://zenodo.org/record/219270/files/figure.png" pageId="5" pageNumber="180" targetBox="[239,1356,183,1921]" targetPageId="5">
<paragraph id="AD6E5004FFB5FE24FF5B95CAFE12F84C" blockId="5.[151,1436,1947,2033]" pageId="5" pageNumber="180">
<emphasis id="9FA58C16FFB5FE24FF5B95CAFEC7F80C" bold="true" box="[151,314,1947,1969]" pageId="5" pageNumber="180">FIGURES 1–5.</emphasis>
<taxonomicName id="6AD12B87FFB5FE24FEF395CDFE11F80C" box="[319,492,1948,1969]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB5FE24FEF395CDFE11F80C" box="[319,492,1948,1969]" italics="true" pageId="5" pageNumber="180">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB5FE24FE3F95CDFDD9F80C" box="[499,548,1948,1969]" pageId="5" pageNumber="180" rank="species">n.sp.</taxonomicNameLabel>
, female paratype from Gruta da Arrifana, SEM images. 1. Front end. 2. Head. 3. Lateral hump of midbody pleurotergum, right side. 4. Spinnerets. 5. Close-up of left spinneret, showing exuded thread.—Scales 100 µm (1, 2); 10 µm (3, 4); 1 µm (5).
</paragraph>
</caption>
<caption id="F9AE008CFFB6FE27FF5B96DDFB01FB5E" httpUri="https://zenodo.org/record/219271/files/figure.png" pageId="6" pageNumber="181" targetBox="[151,1436,193,1142]" targetPageId="6">
<paragraph id="AD6E5004FFB6FE27FF5B96DDFB01FB5E" blockId="6.[151,1436,1164,1252]" pageId="6" pageNumber="181">
<emphasis id="9FA58C16FFB6FE27FF5B96DDFEB7FB1F" bold="true" box="[151,330,1164,1186]" pageId="6" pageNumber="181">FIGURES 6–10.</emphasis>
<taxonomicName id="6AD12B87FFB6FE27FE9896DCFDF8FB1F" box="[340,517,1165,1186]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB6FE27FE9896DCFDF8FB1F" box="[340,517,1165,1186]" italics="true" pageId="6" pageNumber="181">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB6FE27FDC396DCFDBCFB1F" box="[527,577,1165,1186]" pageId="6" pageNumber="181" rank="species">n.sp.</taxonomicNameLabel>
, male paratype from Gruta da Arrifana, anterior gonopods (P8), SEM images. 6. Anterior view. 7. Oblique anterior-apical view. 8. Apical view. 9. Posterior view. 10. Lateral view (from right side).—A: right angiocoxite, F: flagellum, KP: two-pronged process, KL: laminar extension, T: left telopodite.—Scales 10 µm.
</paragraph>
</caption>
<subSubSection id="E5CB038FFFB6FE28FF5B9761FBF6FEFD" lastPageId="9" lastPageNumber="184" pageId="6" pageNumber="181" type="distribution">
<paragraph id="AD6E5004FFB6FE27FF5B9761FE44FAF4" blockId="6.[151,441,1328,1353]" box="[151,441,1328,1353]" pageId="6" pageNumber="181">
<emphasis id="9FA58C16FFB6FE27FF5B9761FE44FAF4" bold="true" box="[151,441,1328,1353]" pageId="6" pageNumber="181">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="AD6E5004FFB6FE27FF5B9728FB83F99D" blockId="6.[151,1436,1400,2037]" pageId="6" pageNumber="181">
<taxonomicName id="6AD12B87FFB6FE27FF5B9728FEAAFA2D" box="[151,343,1401,1424]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB6FE27FF5B9728FEAAFA2D" box="[151,343,1401,1424]" italics="true" pageId="6" pageNumber="181">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB6FE27FE929728FE64FA2D" box="[350,409,1401,1424]" pageId="6" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
has been discovered in three caves of Sicó karst: Gruta da Arrifana, Gruta da Cerâmica and Gruta de Santa Maria da Estrela, all shallow caves distributed along a highly dissected Jurassic karst (
<figureCitation id="35EA4C81FFB6FE27FAC597CDFAA1FA09" box="[1289,1372,1436,1461]" captionStart="FIGURE 14" captionStartId="8.[151,250,1958,1980]" captionTargetBox="[156,1430,1250,1932]" captionTargetId="figure@8.[151,1436,1246,1937]" captionTargetPageId="8" captionText="FIGURE 14. Location of the caves within the Sicó karst in central Portugal in which Scutogona minor n. sp. has been found. 1 — Gruta da Arrifana (type locality); 2 — Gruta de Santa Maria da Estrela and 3 — Gruta da Cerâmica." httpUri="https://zenodo.org/record/219274/files/figure.png" pageId="6" pageNumber="181">Fig. 14</figureCitation>
). The distance between the extremes of the known distribution of the new species is
<quantity id="6A29FDE1FFB6FE27FBC89791FBB3FA64" box="[1028,1102,1472,1497]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="181" unit="km" value="18.0">18 km</quantity>
. The specimens of
<taxonomicName id="6AD12B87FFB6FE27FAEA9790FF27FA41" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9FA58C16FFB6FE27FAEA9790FF27FA41" italics="true" pageId="6" pageNumber="181">Scutogona minor</emphasis>
</taxonomicName>
were only collected in the deepest and most thermally insulated parts of those caves. Humidity was 100% throughout the year, and temperatures ranged from around 16.4º C to 15.3ºC at soil level.
</paragraph>
<paragraph id="AD6E5004FFB6FE27FF0B947DFE6EF944" blockId="6.[151,1436,1400,2037]" pageId="6" pageNumber="181">
The new species shares its habitat with other detritivores hypogean species, such as the isopods
<taxonomicName id="6AD12B87FFB6FE27FF5B9401FDB2F9D4" authority="Vandel, 1946" authorityName="Vandel" authorityYear="1946" box="[151,591,1616,1641]" class="Malacostraca" family="Trichoniscidae" genus="Trichoniscoides" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="broteroi">
<emphasis id="9FA58C16FFB6FE27FF5B9401FE51F9D4" box="[151,428,1616,1641]" italics="true" pageId="6" pageNumber="181">Trichoniscoides broteroi</emphasis>
Vandel, 1946
</taxonomicName>
, a new species of
<emphasis id="9FA58C16FFB6FE27FCE59401FC4AF9D4" box="[809,951,1616,1641]" italics="true" pageId="6" pageNumber="181">
<taxonomicName id="6AD12B87FFB6FE27FCE59401FC4EF9D4" box="[809,947,1616,1641]" class="Malacostraca" family="Trichoniscidae" genus="Miktoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">Miktoniscus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="6AD12B87FFB6FE27FC389401FA65F9D4" authority="Vandel, 1946" authorityName="Vandel" authorityYear="1946" box="[1012,1432,1616,1641]" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="9FA58C16FFB6FE27FC389401FB09F9D4" box="[1012,1268,1616,1641]" italics="true" pageId="6" pageNumber="181">Porcellio cavernicolus</emphasis>
Vandel, 1946
</taxonomicName>
, together with the campodeid dipluran
<emphasis id="9FA58C16FFB6FE27FD9C9425FC80F930" box="[592,893,1652,1677]" italics="true" pageId="6" pageNumber="181">
<taxonomicName id="6AD12B87FFB6FE27FD9C9425FD2BF930" box="[592,726,1652,1677]" class="Entognatha" family="Campodeidae" genus="Podocampa" kingdom="Animalia" order="Diplura" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">Podocampa</taxonomicName>
cf. fragiloides
</emphasis>
Silvestri, 1932. This biocoenosis also includes hypogean predators: the pseudoscorpions
<taxonomicName id="6AD12B87FFB6FE27FDB394C8FBACF90C" authority="Beier, 1962" authorityName="Beier" authorityYear="1962" box="[639,1105,1688,1713]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="9FA58C16FFB6FE27FDB394C8FC51F90C" box="[639,940,1688,1713]" italics="true" pageId="6" pageNumber="181">Roncocreagris blothroides</emphasis>
(Beier, 1962)
</taxonomicName>
and
<taxonomicName id="6AD12B87FFB6FE27FB5C94C8FEB2F969" authority="Zaragoza &amp; Reboleira, 2013" authorityName="Zaragoza &amp; Reboleira" authorityYear="2013" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="borgesi">
<emphasis id="9FA58C16FFB6FE27FB5C94C8FAFAF90C" box="[1168,1287,1688,1713]" italics="true" pageId="6" pageNumber="181">R. borgesi</emphasis>
Zaragoza &amp; Reboleira, 2013
</taxonomicName>
, and the rove beetle
<taxonomicName id="6AD12B87FFB6FE27FDF294ECFBC2F969" authority="Reboleira &amp; Oromi, 2011" authorityName="Reboleira &amp; Oromi" authorityYear="2011" box="[574,1087,1724,1749]" class="Insecta" family="Staphylinidae" genus="Domene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="9FA58C16FFB6FE27FDF294ECFCEFF968" box="[574,786,1724,1749]" italics="true" pageId="6" pageNumber="181">Domene lusitanica</emphasis>
Reboleira &amp; Oromí, 2011
</taxonomicName>
(Reboleira
<emphasis id="9FA58C16FFB6FE27FB0B94ECFAFCF968" box="[1223,1281,1724,1749]" italics="true" pageId="6" pageNumber="181">et al.</emphasis>
2010, 2011a, 2011b, 2013a, 2013b).
</paragraph>
<paragraph id="AD6E5004FFB6FE27FF0B9555FC94F848" blockId="6.[151,1436,1400,2037]" pageId="6" pageNumber="181">
Some interesting troglophile species are common in this
<typeStatus id="726AEEA6FFB6FE27FCF29554FC93F8A0" box="[830,878,1797,1821]" pageId="6" pageNumber="181">type</typeStatus>
of caves, such as the snail
<taxonomicName id="6AD12B87FFB6FE27FB549555FED1F8FD" authority="Beck, 1837" authorityName="Beck" authorityYear="1837" class="Gastropoda" family="Oxychilidae" genus="Oxychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="181" phylum="Mollusca" rank="species" species="draparnaudi">
<emphasis id="9FA58C16FFB6FE27FB549555FA61F8A0" box="[1176,1436,1796,1821]" italics="true" pageId="6" pageNumber="181">Oxychilus draparnaudi</emphasis>
(Beck, 1837)
</taxonomicName>
, the centipede
<taxonomicName id="6AD12B87FFB6FE27FE189579FC99F8FD" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" box="[468,868,1832,1857]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="pilicornis">
<emphasis id="9FA58C16FFB6FE27FE189579FD4CF8FC" box="[468,689,1832,1857]" italics="true" pageId="6" pageNumber="181">Lithobius pilicornis</emphasis>
Newport, 1844
</taxonomicName>
, the springtails
<taxonomicName id="6AD12B87FFB6FE27FBDA9579FA6AF8FD" authority="Willem, 1902" authorityName="Willem" authorityYear="1902" box="[1046,1431,1832,1857]" class="Entognatha" family="Isotomidae" genus="Folsomia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="candida">
<emphasis id="9FA58C16FFB6FE27FBDA9579FB1CF8FC" box="[1046,1249,1832,1857]" italics="true" pageId="6" pageNumber="181">Folsomia candida</emphasis>
(Willem, 1902)
</taxonomicName>
,
<taxonomicName id="6AD12B87FFB6FE27FF5B951CFDFDF8D8" authority="Gama, 1964" authorityName="Gama" authorityYear="1964" box="[151,512,1868,1893]" class="Entognatha" family="Isotomidae" genus="Proisotoma" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="gisini">
<emphasis id="9FA58C16FFB6FE27FF5B951CFE9EF8D9" box="[151,355,1869,1892]" italics="true" pageId="6" pageNumber="181">Proisotoma gisini</emphasis>
Gama, 1964
</taxonomicName>
,
<taxonomicName id="6AD12B87FFB6FE27FDDC951DFBD8F8D9" authority="Schoebotham, 1911" authorityName="Schoebotham" authorityYear="1911" box="[528,1061,1868,1893]" class="Entognatha" family="Oncopoduridae" genus="Oncopodura" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="9FA58C16FFB6FE27FDDC951DFCCEF8D9" box="[528,819,1868,1893]" italics="true" pageId="6" pageNumber="181">Oncopodura crassicornis</emphasis>
Schoebotham, 1911
</taxonomicName>
,
<taxonomicName id="6AD12B87FFB6FE27FBF9951DFA65F8D8" authority="Folsom, 1896" authorityName="Folsom" authorityYear="1896" box="[1077,1432,1868,1893]" class="Entognatha" family="Neelidae" genus="Neelus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="murinus">
<emphasis id="9FA58C16FFB6FE27FBF9951DFB17F8D9" box="[1077,1258,1868,1893]" italics="true" pageId="6" pageNumber="181">Neelus murinus</emphasis>
Folsom, 1896
</taxonomicName>
,
<taxonomicName id="6AD12B87FFB6FE27FF5B9521FD11F834" authority="Cassagnau &amp; Delamare, 1953" authorityName="Cassagnau &amp; Delamare" authorityYear="1953" box="[151,748,1904,1929]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="9FA58C16FFB6FE27FF5B9521FE79F834" box="[151,388,1904,1929]" italics="true" pageId="6" pageNumber="181">Arrhopalites elegans</emphasis>
Cassagnau &amp; Delamare, 1953
</taxonomicName>
,
<taxonomicName id="6AD12B87FFB6FE27FD379521FB3CF834" authority="Gisin, 1951" authorityName="Gisin" authorityYear="1951" box="[763,1217,1904,1929]" class="Entognatha" family="Hypogastruridae" genus="Acherontiella" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="xenylliformis">
<emphasis id="9FA58C16FFB6FE27FD379521FBD2F834" box="[763,1071,1904,1929]" italics="true" pageId="6" pageNumber="181">Acherontiella xenylliformis</emphasis>
Gisin, 1951
</taxonomicName>
and
<taxonomicName id="6AD12B87FFB6FE27FB339521FE62F811" authority="Caroli, 1914" authorityName="Caroli" authorityYear="1914" class="Entognatha" family="Hypogastruridae" genus="Mesachorutes" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="libycus">
<emphasis id="9FA58C16FFB6FE27FB339521FF1BF810" italics="true" pageId="6" pageNumber="181">Mesachorutes libycus</emphasis>
(Caroli, 1914)
</taxonomicName>
, the ground beetle
<taxonomicName id="6AD12B87FFB6FE27FD5895C5FC10F811" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" box="[660,1005,1940,1965]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="fulvus">
<emphasis id="9FA58C16FFB6FE27FD5895C5FCC3F810" box="[660,830,1940,1965]" italics="true" pageId="6" pageNumber="181">Trechus fulvus</emphasis>
Dejean, 1831
</taxonomicName>
, the guanobiont rove beetle
<taxonomicName id="6AD12B87FFB6FE27FA9F95C5FE37F86C" authority="Brisout, 1863" authorityName="Brisout" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="subcavicola">
<emphasis id="9FA58C16FFB6FE27FA9F95C5FEE0F86C" italics="true" pageId="6" pageNumber="181">Atheta subcavicola</emphasis>
Brisout, 1863
</taxonomicName>
, and the spiders
<taxonomicName id="6AD12B87FFB6FE27FD5795E8FB06F86C" authority="Machado &amp; Ribera, 1986" authorityName="Machado &amp; Ribera" authorityYear="1986" box="[667,1275,1976,2001]" class="Arachnida" family="Leptonetidae" genus="Leptoneta" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="conimbricensis">
<emphasis id="9FA58C16FFB6FE27FD5795E8FC3CF86C" box="[667,961,1976,2001]" italics="true" pageId="6" pageNumber="181">Leptoneta conimbricensis</emphasis>
Machado &amp; Ribera, 1986
</taxonomicName>
and
<taxonomicName id="6AD12B87FFB6FE27FAF195E9FE3FF848" authority="Ferrandez, 1996" authorityName="Ferrandez" authorityYear="1996" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="9FA58C16FFB6FE27FAF195E9FEF8F848" italics="true" pageId="6" pageNumber="181">Dysdera machadoi</emphasis>
Ferrández, 1996
</taxonomicName>
(Gama 1963, Reboleira et al. 2011b).
</paragraph>
<caption id="F9AE008CFFB7FE26FF5B95C9FB92F84D" httpUri="https://zenodo.org/record/219272/files/figure.png" pageId="7" pageNumber="182" targetBox="[224,1367,195,1925]" targetPageId="7">
<paragraph id="AD6E5004FFB7FE26FF5B95C9FB92F84D" blockId="7.[151,1436,1944,2032]" pageId="7" pageNumber="182">
<emphasis id="9FA58C16FFB7FE26FF5B95C9FEAFF810" bold="true" box="[151,338,1944,1966]" pageId="7" pageNumber="182">FIGURES 11–12.</emphasis>
<taxonomicName id="6AD12B87FFB7FE26FE9595C8FDFBF813" box="[345,518,1945,1966]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB7FE26FE9595C8FDFBF813" box="[345,518,1945,1966]" italics="true" pageId="7" pageNumber="182">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB7FE26FDC195C8FDC2F813" box="[525,575,1945,1966]" pageId="7" pageNumber="182" rank="species">n.sp.</taxonomicNameLabel>
, paratypes from Gruta da Arrifana, SEM images. 11. male, P 9–11. P9: left posterior gonopod, P10: left first post-gonopodal leg, P11: right second post-gonopodal leg with coxal hook (arrow).—12. Female, vulvae seen from the left. O: operculum, PV: post-vulvar organ, V: lateral valve.—Scales 10 µm.
</paragraph>
</caption>
<caption id="F9AE008CFFB8FE29FF5B9607FCCFFB16" httpUri="https://zenodo.org/record/219273/files/figure.png" pageId="8" pageNumber="183" targetBox="[340,1218,193,1089]" targetPageId="8">
<paragraph id="AD6E5004FFB8FE29FF5B9607FCCFFB16" blockId="8.[151,1436,1110,1196]" pageId="8" pageNumber="183">
<emphasis id="9FA58C16FFB8FE29FF5B9607FEDEFBD1" bold="true" box="[151,291,1110,1132]" pageId="8" pageNumber="183">FIGURE 13.</emphasis>
<taxonomicName id="6AD12B87FFB8FE29FEE19606FE23FBD0" box="[301,478,1111,1133]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB8FE29FEE19606FE23FBD0" box="[301,478,1111,1133]" italics="true" pageId="8" pageNumber="183">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB8FE29FE259606FDE6FBD1" box="[489,539,1111,1132]" pageId="8" pageNumber="183" rank="species">n.sp.</taxonomicNameLabel>
, female paratype from Gruta da Arrifana, vulvae
<emphasis id="9FA58C16FFB8FE29FBE39609FB8FFBD0" box="[1071,1138,1112,1133]" italics="true" pageId="8" pageNumber="183">in situ</emphasis>
behind second pair of legs, posterior view. Insert: semi-diagrammatic drawing of left vulva, showing post-vulvar organ (PV) valves (V) with setae, and the internal separation of the two valves (dotted line).—Scale 50 µm.
</paragraph>
</caption>
<caption id="F9AE008CFFB8FE29FF5B95F7FB5EF860" httpUri="https://zenodo.org/record/219274/files/figure.png" pageId="8" pageNumber="183" targetBox="[156,1430,1250,1932]" targetPageId="8">
<paragraph id="AD6E5004FFB8FE29FF5B95F7FB5EF860" blockId="8.[151,1437,1958,2013]" pageId="8" pageNumber="183">
<emphasis id="9FA58C16FFB8FE29FF5B95F7FEDDF801" bold="true" box="[151,288,1958,1980]" pageId="8" pageNumber="183">FIGURE 14.</emphasis>
Location of the caves within the Sicó karst in central Portugal in which
<taxonomicName id="6AD12B87FFB8FE29FBCB95F6FB48F801" box="[1031,1205,1959,1980]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="9FA58C16FFB8FE29FBCB95F6FB48F801" box="[1031,1205,1959,1980]" italics="true" pageId="8" pageNumber="183">Scutogona minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8496316DFFB8FE29FB7195F6FB09F801" box="[1213,1268,1959,1980]" pageId="8" pageNumber="183" rank="species">n. sp.</taxonomicNameLabel>
has been found. 1—Gruta da Arrifana (type locality); 2—Gruta de Santa Maria da Estrela and 3—Gruta da Cerâmica.
</paragraph>
</caption>
<paragraph id="AD6E5004FFB9FE28FF0B92C9FBF6FEFD" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="184">
In the twilight area and near the entrances of these caves three species of non-hypogean millipedes are frequently observed, the chordeumatidan
<taxonomicName id="6AD12B87FFB9FE28FDA692EDFBAFFF69" authority="Verhoeff, 1900" authorityName="Verhoeff" authorityYear="1900" box="[618,1106,188,213]" class="Diplopoda" family="Haplobainosomatidae" genus="Haplobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="lusitanum">
<emphasis id="9FA58C16FFB9FE28FDA692EDFC5DFF68" box="[618,928,188,213]" italics="true" pageId="9" pageNumber="184">Haplobainosoma lusitanum</emphasis>
Verhoeff, 1900
</taxonomicName>
, the callipodidan
<taxonomicName id="6AD12B87FFB9FE28FAD592ECFE3CFF45" authority="Verhoeff, 1893" authorityName="Verhoeff" authorityYear="1893" class="Diplopoda" family="Dorypetalidae" genus="Lusitanipus" kingdom="Animalia" order="Callipodida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="9FA58C16FFB9FE28FAD592ECFEFCFF45" italics="true" pageId="9" pageNumber="184">Lusitanipus alternans</emphasis>
(Verhoeff, 1893)
</taxonomicName>
and the julid
<taxonomicName id="6AD12B87FFB9FE28FD90928EFBE8FF45" authority="Porat, 1870" authorityName="Porat" authorityYear="1870" box="[604,1045,223,248]" class="Diplopoda" family="Julidae" genus="Cylindroiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="9FA58C16FFB9FE28FD90928EFC87FF45" box="[604,890,223,248]" italics="true" pageId="9" pageNumber="184">Cylindroiulus propinquus</emphasis>
(Porat, 1870)
</taxonomicName>
(Reboleira &amp; Enghoff in press). The discovery of
<taxonomicName id="6AD12B87FFB9FE28FE419354FDB3FEA1" box="[397,590,261,284]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9FA58C16FFB9FE28FE419354FDB3FEA1" box="[397,590,261,284]" italics="true" pageId="9" pageNumber="184">Scutogona minor</emphasis>
</taxonomicName>
increases the troglobiont diversity in the richest cave of central Portugal— Gruta da Cerâmica—which now counts 11 species of cave-adapted arthropods.
</paragraph>
</subSubSection>
<subSubSection id="E5CB038FFFB9FE28FF5B93CBFC00FCE1" pageId="9" pageNumber="184" type="discussion">
<paragraph id="AD6E5004FFB9FE28FF5B93CBFEE8FE0E" blockId="9.[151,277,410,435]" box="[151,277,410,435]" pageId="9" pageNumber="184">
<emphasis id="9FA58C16FFB9FE28FF5B93CBFEE8FE0E" bold="true" box="[151,277,410,435]" pageId="9" pageNumber="184">Discussion</emphasis>
</paragraph>
<paragraph id="AD6E5004FFB9FE28FF5B938DFC00FCE1" blockId="9.[151,1436,476,861]" pageId="9" pageNumber="184">
The gonopods of
<taxonomicName id="6AD12B87FFB9FE28FEAD938CFDDFFE49" box="[353,546,477,500]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9FA58C16FFB9FE28FEAD938CFDDFFE49" box="[353,546,477,500]" italics="true" pageId="9" pageNumber="184">Scutogona minor</emphasis>
</taxonomicName>
are particularly similar to those of
<taxonomicName id="6AD12B87FFB9FE28FC74938CFBEBFE49" box="[952,1046,477,500]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="vivesi">
<emphasis id="9FA58C16FFB9FE28FC74938CFBEBFE49" box="[952,1046,477,500]" italics="true" pageId="9" pageNumber="184">S. vivesi</emphasis>
</taxonomicName>
, but also, surprisingly, to those of
<taxonomicName id="6AD12B87FFB9FE28FF5B9050FDCFFDA4" authority="Silvestri, 1896" authorityName="Silvestri" authorityYear="1896" box="[151,562,512,537]" class="Diplopoda" family="Chamaesomatidae" genus="Meinerteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="lucasi">
<emphasis id="9FA58C16FFB9FE28FF5B9050FE8BFDA4" box="[151,374,512,537]" italics="true" pageId="9" pageNumber="184">Meinerteuma lucasi</emphasis>
(Silvestri, 1896)
</taxonomicName>
(=
<taxonomicName id="6AD12B87FFB9FE28FD959050FC53FDA4" authority="Mauries, 1982" authorityName="Mauries" authorityYear="1982" box="[601,942,512,537]" class="Diplopoda" family="Chamaesomatidae" genus="Meinerteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="edoughense">
<emphasis id="9FA58C16FFB9FE28FD959050FCFFFDA4" box="[601,770,512,537]" italics="true" pageId="9" pageNumber="184">M. edoughense</emphasis>
Mauriès, 1982
</taxonomicName>
). Mauriès (1982) compared
<taxonomicName id="6AD12B87FFB9FE28FB269050FA82FDA5" box="[1258,1407,513,536]" class="Diplopoda" family="Chamaesomatidae" genus="Meinerteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB9FE28FB269050FA82FDA5" box="[1258,1407,513,536]" italics="true" pageId="9" pageNumber="184">Meinerteuma</emphasis>
</taxonomicName>
to
<taxonomicName id="6AD12B87FFB9FE28FF5B9074FEC1FD81" box="[151,316,549,572]" class="Diplopoda" family="Origmatogonidae" genus="Origmatogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB9FE28FF5B9074FEC1FD81" box="[151,316,549,572]" italics="true" pageId="9" pageNumber="184">Origmatogona</emphasis>
</taxonomicName>
and
<taxonomicName id="6AD12B87FFB9FE28FEB79074FDF0FD80" box="[379,525,549,573]" class="Diplopoda" family="Chamaesomatidae" genus="Coiffaiteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB9FE28FEB79074FDF0FD80" box="[379,525,549,573]" italics="true" pageId="9" pageNumber="184">Coiffaiteuma</emphasis>
</taxonomicName>
, but also mentioned similarities with
<taxonomicName id="6AD12B87FFB9FE28FC029074FBB9FD81" box="[974,1092,549,572]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB9FE28FC029074FBB9FD81" box="[974,1092,549,572]" italics="true" pageId="9" pageNumber="184">Scutogona</emphasis>
</taxonomicName>
. It is a question whether the distinction between these genera, and maybe all of the scutogonine genera, can be upheld. In his description of
<taxonomicName id="6AD12B87FFB9FE28FAB19018FEE6FD38" class="Diplopoda" family="Chamaesomatidae" genus="Meinerteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="edoughense">
<emphasis id="9FA58C16FFB9FE28FAB19018FEE6FD38" italics="true" pageId="9" pageNumber="184">M. edoughense</emphasis>
</taxonomicName>
,
<typeStatus id="726AEEA6FFB9FE28FEE3903CFEA2FD38" box="[303,351,621,645]" pageId="9" pageNumber="184">type</typeStatus>
species of
<emphasis id="9FA58C16FFB9FE28FE3C903CFD78FD39" box="[496,645,621,644]" italics="true" pageId="9" pageNumber="184">Meienrteuma</emphasis>
, Mauriès stated that none of the walking legs had any particular structures. Re-examination of the
<typeStatus id="726AEEA6FFB9FE28FDDB90C1FD84FD14" box="[535,633,656,681]" pageId="9" pageNumber="184" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6AD12B87FFB9FE28FD6C90C0FCB6FD14" box="[672,843,656,681]" class="Diplopoda" family="Chamaesomatidae" genus="Meinerteuma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="edoughense">
<emphasis id="9FA58C16FFB9FE28FD6C90C0FCB6FD14" box="[672,843,656,681]" italics="true" pageId="9" pageNumber="184">M. edoughense</emphasis>
</taxonomicName>
(ZMUC) revealed, however, that although both P10 and both P11 are broken, enough remains of P11 to show that coxal hooks, similar to those seen in
<taxonomicName id="6AD12B87FFB9FE28FB3A90E4FA91FD71" box="[1270,1388,693,716]" class="Diplopoda" family="Chamaesomatidae" genus="Scutogona" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="9FA58C16FFB9FE28FB3A90E4FA91FD71" box="[1270,1388,693,716]" italics="true" pageId="9" pageNumber="184">Scutogona</emphasis>
</taxonomicName>
and several other scutogonine genera, are actually present. Coxal hooks on P11 were also found by re-examination of the Tunisian male reported by Akkari et al. (2010)—these authors likewise didn’t notice this character, which contributes to the confusion about genus delimitations in Scutogoninae, clarification of which will require a much more comprehensive approach than lies within the scope of the present paper.
</paragraph>
</subSubSection>
</treatment>
</document>
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