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<document id="1878360BD31DAFA1D720F49124C14F2A" ID-DOI="10.11646/zootaxa.3584.1.1" ID-ISSN="1175­5334" ID-ZooBank="072032C4-63FC-499A-A61D-58B428051302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629830673067" checkinUser="felipe" docAuthor="Michat, Mariano C., Alarie, Yves &amp; Watts, Chris H. S." docDate="2012" docId="256287ACFF8A2718FBE8FF05E13086C9" docLanguage="en" docName="zt03584p110.pdf" docOrigin="Zootaxa 3584 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3584.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Limbodessus raeae Watts &amp; Humphreys 2006" docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="D95BFFD4FFC52757FB7FFF92E231831C" masterDocTitle="3584" masterLastPageNumber="110" masterPageNumber="1" pageNumber="80" updateTime="1699118123178" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="39C3839E75DCA5F47BDD95AF8BB8615F">3584</mods:title>
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<mods:namePart id="778DCB97B5D9D2D50200281FADF62710">Michat, Mariano C.</mods:namePart>
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<mods:namePart id="621A8C416F4B4A95E5E417FCC6790FD1">Alarie, Yves</mods:namePart>
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<mods:namePart id="CD7FFDCA862B203EFCAB81759EDBDE21">Watts, Chris H. S.</mods:namePart>
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<treatment id="256287ACFF8A2718FBE8FF05E13086C9" ID-DOI="http://doi.org/10.5281/zenodo.5462394" ID-Zenodo-Dep="5462394" LSID="urn:lsid:plazi:treatment:256287ACFF8A2718FBE8FF05E13086C9" httpUri="http://treatment.plazi.org/id/256287ACFF8A2718FBE8FF05E13086C9" lastPageNumber="80" pageId="79" pageNumber="80">
<subSubSection id="E5D16531FF8A2718FBE8FF05E37383CE" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="AD7436BAFF8A2718FBE8FF05E0E083AD" blockId="79.[151,721,151,210]" box="[151,721,151,177]" pageId="79" pageNumber="80">
<heading id="F63C81D6FF8A2718FBE8FF05E0E083AD" bold="true" box="[151,721,151,177]" fontSize="11" level="1" pageId="79" pageNumber="80" reason="1">
<taxonomicName id="6ACB4D39FF8A2718FBE8FF05E0E083AD" ID-CoL="6QBH6" authority="Watts &amp; Humphreys, 2006" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[151,721,151,177]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="raeae">
<emphasis id="9FBFEAA8FF8A2718FBE8FF05E0E083AD" bold="true" box="[151,721,151,177]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBE8FF05E34C83AD" bold="true" box="[151,381,151,177]" italics="true" pageId="79" pageNumber="80">Limbodessus raeae</emphasis>
<bibRefCitation id="C95A4B4BFF8A2718FAFBFF05E0E083AD" author="Watts &amp; Humphreys" box="[388,721,151,177]" firstAuthor="Watts" pageId="79" pageNumber="80" pagination="123 - 185" refId="ref60425" refString="Watts, C. H. S. &amp; Humphreys, W. F. (2006) Twenty-six new Dytiscidae (Coleoptera) of the genera Limbodessus Guignot and Nirripirti Watts &amp; Humphreys, from underground waters in Australia. Transactions of the Royal Society of South Australia, 130, 123 - 185." type="journal article" year="2006">Watts &amp; Humphreys, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD7436BAFF8A2718FBE8FF28E37383CE" blockId="79.[151,721,151,210]" box="[151,322,186,210]" pageId="79" pageNumber="80">(Figs 208211)</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF8A2718FBE8FF6DE7B38241" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="AD7436BAFF8A2718FBE8FF6DE7B38241" blockId="79.[151,1437,255,1493]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBE8FF6DE34D8204" bold="true" box="[151,380,255,280]" inLineHeading="true" pageId="79" pageNumber="80" reason="1">Source of material.</emphasis>
<materialsCitation id="1DA33CE7FF8A2718FAFBFF6DE7B38241" accessionNumber="BES 14305" collectingDate="2006-09-17" collectorName="W. F. Humphreys &amp; T. Moulds." country="Australia" county="Dawson's Well" latitude="-28.8866" location="One" longLatPrecision="6" longitude="120.1621" municipality="Lake Way Station" pageId="79" pageNumber="80" specimenCount="1">
<location id="A8146061FF8A2718FAFBFF6DE382820B" LSID="urn:lsid:plazi:treatment:256287ACFF8A2718FBE8FF05E13086C9:A8146061FF8A2718FAFBFF6DE382820B" box="[388,435,255,279]" country="Australia" county="Dawson's Well" latitude="-28.8866" longLatPrecision="6" longitude="120.1621" municipality="Lake Way Station" name="One" pageId="79" pageNumber="80">One</location>
specimen of instar III was used for the description (
<tableCitation id="E0490301FF8A2718FF7BFF6DE668820B" box="[1028,1113,255,279]" captionStart="TABLE 1" captionStartId="5.[151,238,608,632]" captionText="TABLE 1. Species and instars described in the present study. The numbers under each instar indicate the specimens studied." pageId="79" pageNumber="80">Table 1</tableCitation>
).
<location id="A8146061FF8A2718FF0DFF6DE6AC820B" LSID="urn:lsid:plazi:treatment:256287ACFF8A2718FBE8FF05E13086C9:A8146061FF8A2718FF0DFF6DE6AC820B" box="[1138,1181,255,279]" country="Australia" county="Dawson's Well" latitude="-28.8866" longLatPrecision="6" longitude="120.1621" municipality="Lake Way Station" name="The" pageId="79" pageNumber="80">The</location>
larva was collected in association with adults at the following locality:
<emphasis id="9FBFEAA8FF8A2718F9C3FEB0E62C8227" bold="true" box="[700,1053,290,315]" pageId="79" pageNumber="80">
<collectingCountry id="D5DC762AFF8A2718F9C3FEB0E11F8227" box="[700,814,290,315]" name="Australia" pageId="79" pageNumber="80">Australia</collectingCountry>
,
<location id="A8146061FF8A2718F844FEB0E1B58227" LSID="urn:lsid:plazi:treatment:256287ACFF8A2718FBE8FF05E13086C9:A8146061FF8A2718F844FEB0E1B58227" box="[827,900,290,315]" country="Australia" county="Dawson's Well" latitude="-28.8866" longLatPrecision="6" longitude="120.1621" municipality="Lake Way Station" name="Carey" pageId="79" pageNumber="80">Carey</location>
palaeovalley
</emphasis>
,
<location id="A8146061FF8A2718FF54FEB0E68D8226" LSID="urn:lsid:plazi:treatment:256287ACFF8A2718FBE8FF05E13086C9:A8146061FF8A2718FF54FEB0E68D8226" box="[1067,1212,290,314]" country="Australia" county="Dawson's Well" latitude="-28.8866" longLatPrecision="6" longitude="120.1621" municipality="Lake Way Station" name="Hinkler Well" pageId="79" pageNumber="80">Hinkler Well</location>
calcrete,
<collectingMunicipality id="4D10ACC0FF8A2718FE56FEB0E2D98241" pageId="79" pageNumber="80">Lake Way Station</collectingMunicipality>
,
<collectingCounty id="44154E36FF8A2718FB8CFED7E3918241" box="[243,416,325,349]" pageId="79" pageNumber="80">Dawsons Well</collectingCounty>
, BES 14305,
<geoCoordinate id="C8FF507DFF8A2718F945FED7E0968241" box="[570,679,325,349]" degrees="28.8866" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="5" value="-28.8866">28.8866S</geoCoordinate>
,
<geoCoordinate id="C8FF507DFF8A2718F9CDFED7E11E8241" box="[690,815,325,349]" degrees="120.1621" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="5" value="120.1621">120.1621E</geoCoordinate>
,
<date id="D975107AFF8A2718F845FED7E1FB8241" box="[826,970,325,349]" pageId="79" pageNumber="80" value="2006-09-17">
<collectingDate id="C931E992FF8A2718F845FED7E1FB8241" box="[826,970,325,349]" pageId="79" pageNumber="80" value="2006-09-17">17IX2006</collectingDate>
</date>
, coll.
<collectorName id="003E536CFF8A2718FF71FED4E6E38241" box="[1038,1234,325,349]" pageId="79" pageNumber="80">W. F. Humphreys</collectorName>
and
<collectorName id="003E536CFF8A2718FE77FED4E7B38241" box="[1288,1410,325,349]" pageId="79" pageNumber="80">T. Moulds.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF8A2718FBB8FEF5E04E8781" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="AD7436BAFF8A2718FBB8FEF5E0608153" blockId="79.[151,1437,255,1493]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBB8FEF5E3E1829C" bold="true" box="[199,464,359,384]" inLineHeading="true" pageId="79" pageNumber="80" reason="1">Diagnosis (instar III).</emphasis>
Medium-sized species (HL
<quantity id="6A339B5FFF8A2718F860FEF5E1FB8263" box="[799,970,359,383]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="11.5" metricValueMin="5.5" pageId="79" pageNumber="80" unit="mm" value="0.85" valueMax="1.15" valueMin="0.55">0.551.15 mm</quantity>
); head (Fig. 208) subpentagonal; nasale subtriangular; half-circle of dense spinulae on lateroventral margins of nasale absent; hole-like structure on ventrodistal surface of nasale absent; lateral margins of nasale not inflated in dorsal view; lateral branches of nasale minute; slender spinulae anterior to seta FR13 scarce (20 or less); occipital foramen well developed (HW/OCW less than 1.90); occipital suture present; lateral margins of parietal straight; secondary spiniform setae on lateral margins of parietal scarce; seta AN2 present; distal half of MN broad; setae LA3, LA4, LA5 and LA8 hair-like; secondary setae on U absent (Fig. 211).
</paragraph>
<paragraph id="AD7436BAFF8A2718FBB8FDC9E3FC816E" blockId="79.[151,1437,255,1493]" box="[199,461,602,627]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBB8FDC9E318816F" bold="true" box="[199,297,603,627]" pageId="79" pageNumber="80">Instar I.</emphasis>
Not available.
</paragraph>
<paragraph id="AD7436BAFF8A2718FBB8FDEFE3E98189" blockId="79.[151,1437,255,1493]" box="[199,472,637,661]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBB8FDEFE3028189" bold="true" box="[199,307,637,661]" pageId="79" pageNumber="80">Instar II.</emphasis>
Not available.
</paragraph>
<paragraph id="AD7436BAFF8A2718FBB8FD32E04E8781" blockId="79.[151,1437,255,1493]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBB8FD32E30881A4" bold="true" box="[199,313,672,696]" pageId="79" pageNumber="80">Instar III</emphasis>
(Figs 208211). Head (Fig. 208). A3 less than 2.25 times longer than A1; A3 less than 1.45 times longer than A2; MN less than 4.85 times longer than broad; MP less than 1.75 times longer than labial palpus; MP2 2.103.10 times longer than MP3; LP2 more than 0.65 times as long as LP1. Legs (Figs 209210). L3 2.252.85 times longer than HW. Abdomen (Fig. 211). Chaetotaxy. Anteroventral margin of nasale with 59 lamellae clypeales distributed in 2 rows; proCO with 14 anterior secondary setae; mesoCO with 15 anterior secondary setae; pro-, meso- and metaCO without posterior secondary setae; proCO with less than 11 secondary setae; metaCO with less than 20 secondary setae; anterodorsal secondary setae on pro- and metaFE present; metaFE with less than 12 anteroventral secondary setae; posterodorsal secondary setae on meso- and metaFE absent; metaFE with 927 secondary setae; anterodorsal and anteroventral secondary setae on proTI present; metaTI with less than 9 anteroventral secondary setae; posterodorsal secondary setae on proTI absent; metaTI with less than 16 secondary setae; anterodorsal and posteroventral secondary setae on pro-, meso- and metaTA absent; anteroventral secondary setae on pro- and metaTA present; posterodorsal secondary setae on proTA absent; posterodorsal secondary setae on meso- and metaTA present; proTA with 17 secondary setae; metaTA with less than 16 secondary setae; secondary setae on U absent. Measurements and ratios that characterize the body shape are shown in
<tableCitation id="E0490301FF8A2718FE4DFBF0E7A98766" box="[1330,1432,1122,1146]" captionStart="TABLE 11" captionStartId="41.[151,239,155,179]" captionText="TABLE 11. Measurements and ratios for the third-instar larvae of species of Limbodessus." pageId="79" pageNumber="80">Table 11</tableCitation>
. Secondary leg setation detailed in
<tableCitation id="E0490301FF8A2718F967FB17E04A8781" box="[536,635,1157,1181]" captionStart="TABLE 19" captionStartId="82.[151,239,871,895]" captionText="TABLE 19. Number and position of secondary setae on the legs of third-instar larvae of species of Limbodessus. Numbers between slash marks refer to pro-, meso- and metathoracic leg, respectively. A = anterior, D = dorsal, P = posterior, Pr = proximal, V = ventral, Total = total number of secondary setae on the segment (excluding primary setae)." pageId="79" pageNumber="80">Table 19</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF8A2718FBB8FB3AE13086C9" pageId="79" pageNumber="80" type="discussion">
<paragraph id="AD7436BAFF8A2718FBB8FB3AE13086C9" blockId="79.[151,1437,255,1493]" pageId="79" pageNumber="80">
<emphasis id="9FBFEAA8FF8A2718FBB8FB3AE30A87DC" bold="true" box="[199,315,1192,1216]" inLineHeading="true" pageId="79" pageNumber="80" reason="1">Remarks.</emphasis>
The description of
<taxonomicName id="6ACB4D39FF8A2718F961FB3BE04C87DC" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[542,637,1193,1216]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="raeae">
<emphasis id="9FBFEAA8FF8A2718F961FB3BE04C87DC" box="[542,637,1193,1216]" italics="true" pageId="79" pageNumber="80">L. raeae</emphasis>
</taxonomicName>
is based on a single instar III in which the mesothoracic legs and the second urogomphomere were broken, which prevented the evaluation of some morphometric and chaetotaxic characters. This species belongs to the group of stygobitic species characterized by the absence of secondary setae on the urogomphus (
<taxonomicName id="6ACB4D39FF8A2718FAE6FA83E00C8634" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[409,573,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="challaensis">
<emphasis id="9FBFEAA8FF8A2718FAE6FA83E00C8634" box="[409,573,1296,1320]" italics="true" pageId="79" pageNumber="80">L. challaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F92BFA83E0878634" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[596,694,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="9FBFEAA8FF8A2718F92BFA83E0878634" box="[596,694,1296,1320]" italics="true" pageId="79" pageNumber="80">L. exilis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F9B3FA83E1AC8634" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[716,925,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="fridaywellensis">
<emphasis id="9FBFEAA8FF8A2718F9B3FA83E1AC8634" box="[716,925,1296,1320]" italics="true" pageId="79" pageNumber="80">L. fridaywellensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F8CCFA83E65B8634" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2004" box="[947,1130,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hillviewensis">
<emphasis id="9FBFEAA8FF8A2718F8CCFA83E65B8634" box="[947,1130,1296,1320]" italics="true" pageId="79" pageNumber="80">L. hillviewensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FFFEFA83E6CF8634" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[1153,1278,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hinkleri">
<emphasis id="9FBFEAA8FF8A2718FFFEFA83E6CF8634" box="[1153,1278,1296,1320]" italics="true" pageId="79" pageNumber="80">L. hinkleri</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FE6AFA83E75F8634" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[1301,1390,1296,1320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="leysi">
<emphasis id="9FBFEAA8FF8A2718FE6AFA83E75F8634" box="[1301,1390,1296,1320]" italics="true" pageId="79" pageNumber="80">L. leysi</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FEFAFA83E3298656" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="masonensis">
<emphasis id="9FBFEAA8FF8A2718FEFAFA83E3298656" italics="true" pageId="79" pageNumber="80">L. masonensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FA58FAA1E3E68657" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[295,471,1331,1355]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="millbilliensis">
<emphasis id="9FBFEAA8FF8A2718FA58FAA1E3E68657" box="[295,471,1331,1355]" italics="true" pageId="79" pageNumber="80">L. millbilliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FA99FAA1E0528656" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[486,611,1331,1354]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF8A2718FA99FAA1E0528656" box="[486,611,1331,1354]" italics="true" pageId="79" pageNumber="80">L. morgani</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F90CFAA1E1368657" authorityName="Watts &amp; Humphreys" authorityYear="2009" box="[627,775,1331,1355]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="ordinarius">
<emphasis id="9FBFEAA8FF8A2718F90CFAA1E1368657" box="[627,775,1331,1355]" italics="true" pageId="79" pageNumber="80">L. ordinarius</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F869FAA1E1448657" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[790,885,1331,1355]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="pulpa">
<emphasis id="9FBFEAA8FF8A2718F869FAA1E1448657" box="[790,885,1331,1355]" italics="true" pageId="79" pageNumber="80">L. pulpa</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718F8FAFAA1E6758657" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[901,1092,1331,1355]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="windarraensis">
<emphasis id="9FBFEAA8FF8A2718F8FAFAA1E6758657" box="[901,1092,1331,1355]" italics="true" pageId="79" pageNumber="80">L. windarraensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF8A2718FF2BFAA1E7238656" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2003" box="[1108,1298,1331,1354]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="yuinmeryensis">
<emphasis id="9FBFEAA8FF8A2718FF2BFAA1E7238656" box="[1108,1298,1331,1354]" italics="true" pageId="79" pageNumber="80">L. yuinmeryensis</emphasis>
</taxonomicName>
) (Fig. 211). Compared to these species, larvae of
<taxonomicName id="6ACB4D39FF8A2718F945FAC4E0A98671" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[570,664,1366,1389]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="raeae">
<emphasis id="9FBFEAA8FF8A2718F945FAC4E0A98671" box="[570,664,1366,1389]" italics="true" pageId="79" pageNumber="80">L. raeae</emphasis>
</taxonomicName>
can be distinguished by the lateral margins of the nasale not inflated in dorsal view (Fig. 208), the presence of anterior secondary setae on the pro- and mesocoxa (Fig. 209), the presence of anteroventral secondary setae on the protibia and the protarsus (Fig. 209), and the absence of posteroventral secondary setae on the tarsus (Fig. 210).
</paragraph>
</subSubSection>
</treatment>
</document>