treatments-xml/data/CD/4D/87/CD4D87D9FFAB652FFF33BCF6658BF873.xml

<document id="9455631BF600D14670A94E6FD8EECF3B" ID-CLB-Dataset="27710" ID-DOI="10.11646/zootaxa.4543.1.5" ID-GBIF-Dataset="498bf72a-fb02-4c71-adab-0450f4b0432b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617729" ID-ZooBank="92B59468-BDEE-43E7-83B0-24BC0DBE556D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928171120" checkinUser="plazi" docAuthor="Molineri, Carlos &amp; Granados-Martinez, Cristian E." docDate="2019" docId="CD4D87D9FFAB652FFF33BCF6658BF873" docLanguage="en" docName="zootaxa.4543.1.5.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus vichada Molineri &amp; Granados-Martinez 2019, sp. nov." docType="treatment" docVersion="8" masterDocId="3174FFA1FFAA652CFFA4BE236010FF87" masterDocTitle="Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia" updateTime="1698715347377" updateUser="ExternalLinkService">
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<mods:title id="89D85F7309D096AF5B3EB15E58EF2B26">Two new species of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) from Colombia</mods:title>
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<mods:namePart id="3C811A0A940A2B924DDCEE53A65739C0">Molineri, Carlos</mods:namePart>
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<mods:namePart id="3A16D62C5E2EF09DDD0B77D7C9912AF4">Granados-Martinez, Cristian E.</mods:namePart>
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<mods:date id="96656EEA026DCB5F568E9E6E5B4CF286">2019</mods:date>
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<treatment id="CD4D87D9FFAB652FFF33BCF6658BF873" ID-DOI="http://doi.org/10.5281/zenodo.5932816" ID-GBIF-Taxon="154529707" ID-Zenodo-Dep="5932816" LSID="urn:lsid:plazi:treatment:CD4D87D9FFAB652FFF33BCF6658BF873" httpUri="http://treatment.plazi.org/id/CD4D87D9FFAB652FFF33BCF6658BF873" lastPageId="3" pageId="1">
<subSubSection id="0DFE6544FFAB652DFF33BCF661FBFD68" box="[151,491,725,751]" pageId="1" type="nomenclature">
<paragraph id="455B36CFFFAB652DFF33BCF661FBFD68" blockId="1.[151,1126,725,821]" box="[151,491,725,751]" pageId="1">
<heading id="1E1381A3FFAB652DFF33BCF661FBFD68" bold="true" box="[151,491,725,751]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="7790EADDFFAB652DFF33BCF661FBFD68" bold="true" box="[151,491,725,751]" pageId="1">
<taxonomicName id="82E44D4CFFAB652DFF33BCF66197FD68" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[151,391,725,751]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" phylum="Arthropoda" rank="species" species="vichada" status="sp. nov.">
<emphasis id="7790EADDFFAB652DFF33BCF66197FD68" bold="true" box="[151,391,725,751]" italics="true" pageId="1">Campsurus vichada</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CA357A6FFAB652DFE2ABCF661FBFD68" box="[398,491,725,751]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAB652DFF33BCD46476FCB2" pageId="1" type="description">
<paragraph id="455B36CFFFAB652DFF33BCD461ABFC97" blockId="1.[151,1126,725,821]" box="[151,443,759,784]" pageId="1">
(
<figureCitation id="DDDF2A4AFFAB652DFF3BBCD460F7FC97" box="[159,231,759,784]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="1">Figs. 1</figureCitation>
A–C, 
<figureCitation id="DDDF2A4AFFAB652DFE8BBCD46169FC97" box="[303,377,759,784]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Figs. 3</figureCitation>
A–D)
</paragraph>
<paragraph id="455B36CFFFAB652DFF33BD3F6476FCB2" blockId="1.[151,1126,725,821]" box="[151,1126,796,821]" pageId="1">
http://zoobank.org/NomenclaturalActs/ 
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</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAB652DFF33BD4761F0FBD3" pageId="1" type="diagnosis">
<paragraph id="455B36CFFFAB652DFF33BD4761F0FBD3" blockId="1.[151,1437,868,2009]" pageId="1">
<emphasis id="7790EADDFFAB652DFF33BD476101FCFA" bold="true" box="[151,273,868,893]" pageId="1">Diagnosis.</emphasis>
<taxonomicName id="82E44D4CFFAB652DFEB3BD4661E7FCFA" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[279,503,868,893]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" phylum="Arthropoda" rank="species" species="vichada" status="sp. nov.">
<emphasis id="7790EADDFFAB652DFEB3BD4661E7FCFA" box="[279,503,868,893]" italics="true" pageId="1">Campsurus vichada</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFAB652DFE59BD476246FCFA" bold="true" box="[509,598,868,893]" pageId="1">
<taxonomicNameLabel id="6CA357A6FFAB652DFE59BD476246FCFA" box="[509,598,868,893]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, here described from male and female imagos, can be separated from other species of the genus by the characters listed for the albifilum species group (
<bibRefCitation id="21754B3EFFAB652DFC55BDA464EEFC27" author="Molineri, C. &amp; Salles, F. F." box="[1009,1278,903,928]" pageId="1" pagination="265 - 267" refId="ref4051" refString="Molineri, C. &amp; Salles, F. F. (2013) Phylogeny and biogeography of the ephemeral Campsurus (Ephemeroptera, Polymitarcyidae). Systematic Entomology, 38 (2), 265 - 267. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00656. x" type="journal article" year="2013">Molineri &amp; Salles 2013</bibRefCitation>
) and from the species of this group, by: 1) FW 10.0–11.0 mm (male), 13.5–16.0 (female); 2) dorsum of head shaded black widely, some specimens with Y–shaped pale area between ocelli and extending through occiput (
<figureCitation id="DDDF2A4AFFAB652DFB7FBDEC653FFC6F" box="[1243,1327,975,1000]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Figs. 3</figureCitation>
A–B); 3) abdominal terga shaded slightly darker posteriorly, with a pale median band and other markings as in 
<figureCitation id="DDDF2A4AFFAB652DFAA8BDD76546FB8B" box="[1292,1366,1012,1037]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Figs. 3</figureCitation>
A–B; 4) pedestals relatively short, subquadrate, main lobes of penes very long and slender, secondary lobe reduced and relatively short (
<figureCitation id="DDDF2A4AFFAB652DFEEBBA1F618AFBD3" box="[335,410,1084,1109]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Figs. 3</figureCitation>
E–D).
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFAB652EFF63BA7C64B4FEC7" lastPageId="2" pageId="1" type="materials_examined">
<paragraph id="455B36CFFFAB652DFF63BA7C6210FB63" blockId="1.[151,1437,868,2009]" pageId="1">
<emphasis id="7790EADDFFAB652DFF63BA7C6165FBFF" bold="true" box="[199,373,1119,1144]" pageId="1">Type material.</emphasis>
<materialsCitation id="F58C3C92FFAB652DFED9BA7C63DEFB1B" ID-GBIF-Occurrence="2147524807" collectingDate="2017-04-04" collectorName="C Granados" country="Colombia" elevation="119" latitude="5.5772223" location="rio Tomo" longLatPrecision="21" longitude="-68.49305" pageId="1" specimenCount="1" specimenCount-male="1" stateProvince="Vichada" typeStatus="holotype">
<typeStatus id="9A5F886DFFAB652DFED9BA7C61F5FBFF" box="[381,485,1119,1144]" pageId="1" type="holotype">Holotype</typeStatus>
<specimenCount id="53E2FD46FFAB652DFE49BA436212FBF0" box="[493,514,1120,1143]" pageId="1" type="male">♂</specimenCount>
imago from 
<collectingCountry id="3DF3765FFFAB652DFD32BA43633EFBFF" box="[662,814,1120,1144]" name="Colombia" pageId="1">COLOMBIA</collectingCountry>
, 
<collectingRegion id="8720F82DFFAB652DFC9EBA7C6389FBFF" box="[826,921,1119,1144]" country="Colombia" name="Vichada" pageId="1">Vichada</collectingRegion>
, 
<location id="403B6014FFAB652DFC00BA7C641FFBF0" LSID="urn:lsid:plazi:treatment:CD4D87D9FFAB652FFF33BCF6658BF873:403B6014FFAB652DFC00BA7C641FFBF0" box="[932,1039,1119,1144]" country="Colombia" latitude="5.5772223" longLatPrecision="21" longitude="-68.49305" name="rio Tomo" pageId="1" stateProvince="Vichada">rio Tomo</location>
, 
<geoCoordinate id="20D05008FFAB652DFBBEBA4364B5FBFF" box="[1050,1189,1119,1144]" direction="north" orientation="latitude" pageId="1" precision="15" value="5.5772223">N 5° 34' 38&quot;</geoCoordinate>
/ 
<geoCoordinate id="20D05008FFAB652DFB18BA43654DFBFF" box="[1212,1373,1119,1144]" direction="west" orientation="longitude" pageId="1" precision="15" value="-68.49305">W 68° 29' 35&quot;</geoCoordinate>
), 
<quantity id="821C9B2AFFAB652DFAD7BA4360BFFB1B" metricMagnitude="2" metricUnit="m" metricValue="1.19" pageId="1" unit="m" value="119.0">
<elevation id="CEC9D1FCFFAB652DFAD7BA4360BFFB1B" metricMagnitude="2" metricUnit="m" metricValue="1.19" pageId="1" unit="m" value="119.0">119 m</elevation>
</quantity>
, 
<date id="315A100FFFAB652DFF60BAA7612DFB1A" box="[196,317,1156,1181]" pageId="1" value="2017-04-04">
<collectingDate id="211EE9E7FFAB652DFF60BAA7612DFB1A" box="[196,317,1156,1181]" pageId="1" value="2017-04-04">04.iv.2017</collectingDate>
</date>
, 
<collectingMethod id="9CA54ED8FFAB652DFEF5BAA761D0FB1B" box="[337,448,1156,1181]" pageId="1">light trap</collectingMethod>
18–20 hs, 
<collectorName id="E8115319FFAB652DFDF1BAA762F3FB1A" box="[597,739,1156,1181]" pageId="1">C Granados</collectorName>
col. (MUSENUV)
</materialsCitation>
. 
<materialsCitation id="F58C3C92FFAB652DFC45BAA66136FB47" ID-GBIF-Occurrence="2147524570" collectionCode="CM" country="Colombia" pageId="1" specimenCode="IBN800, IBN801" specimenCount="52" specimenCount-male="52" stateProvince="Vichada" typeStatus="paratype">
<typeStatus id="9A5F886DFFAB652DFC45BAA66443FB1A" box="[993,1107,1157,1181]" pageId="1" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="53E2FD46FFAB652DFBC1BAA764B6FB1B" box="[1125,1190,1156,1181]" pageId="1" type="male">52 ♂</specimenCount>
(slides 
<specimenCode id="15429EB4FFAB652DFAABBAA7657BFB1B" box="[1295,1387,1156,1180]" collectionCode="IBN" pageId="1">IBN800</specimenCode>
<collectionCode id="23F5AE0AFFAB652DFACFBAA76588FB1B" box="[1387,1432,1156,1180]" country="China" name="Chongqing Museum" pageId="1">CM</collectionCode>
, 
<specimenCode id="15429EB4FFAB652DFF33BA8B60E0FB47" box="[151,240,1192,1216]" collectionCode="IBN" pageId="1">IBN801</specimenCode>
<collectionCode id="23F5AE0AFFAB652DFF55BA8B610EFB47" box="[241,286,1192,1216]" country="China" name="Chongqing Museum" pageId="1">CM</collectionCode>
)
</materialsCitation>
and 
<materialsCitation id="F58C3C92FFAB652DFEFBBA846269FB47" ID-GBIF-Occurrence="2147524669" box="[351,633,1191,1216]" collectionCode="CM" country="Colombia" pageId="1" specimenCode="IBN802" specimenCount="16" specimenCount-female="16" stateProvince="Vichada" typeStatus="paratype">
<specimenCount id="53E2FD46FFAB652DFEFBBA846188FB38" box="[351,408,1191,1216]" pageId="1" type="female">16 ♀</specimenCount>
(slide 
<specimenCode id="15429EB4FFAB652DFE42BA8B6253FB47" box="[486,579,1192,1216]" collectionCode="IBN" pageId="1">IBN802</specimenCode>
<collectionCode id="23F5AE0AFFAB652DFDE7BA8B6260FB47" box="[579,624,1192,1216]" country="China" name="Chongqing Museum" pageId="1">CM</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="F58C3C92FFAB652DFD21BA8B61EBFB63" ID-GBIF-Occurrence="2147524671" collectingDate="2017-04-04" collectionCode="CEBUC, IBN" collectorName="C Granados" country="Colombia" elevation="119" latitude="5.5772223" location="rio Tomo" longLatPrecision="21" longitude="-68.49305" pageId="1" specimenCount="68" specimenCount-female="16" specimenCount-male="52" stateProvince="Vichada" typeStatus="paratype">
same data as holotype (
<specimenCount id="53E2FD46FFAB652DFC2BBA8B63D9FB38" box="[911,969,1192,1216]" pageId="1" type="male">10 ♂</specimenCount>
and 
<specimenCount id="53E2FD46FFAB652DFBA6BA8B643DFB38" box="[1026,1069,1192,1216]" pageId="1" type="female">4 ♀</specimenCount>
in MUSENUV, 
<specimenCount id="53E2FD46FFAB652DFB4DBA8B6532FB38" box="[1257,1314,1192,1216]" pageId="1" type="male">10 ♂</specimenCount>
and 
<specimenCount id="53E2FD46FFAB652DFAFFBA8B656EFB47" box="[1371,1406,1192,1216]" pageId="1" type="female">4♀</specimenCount>
in 
<collectionCode id="23F5AE0AFFAB652DFF33BAEF60E7FB63" box="[151,247,1228,1252]" pageId="1">CEBUC</collectionCode>
, 
<specimenCount id="53E2FD46FFAB652DFEA6BAEF612AFB63" box="[258,314,1228,1252]" pageId="1" type="male">32 ♂</specimenCount>
and 
<specimenCount id="53E2FD46FFAB652DFED5BAEF618BFB63" box="[369,411,1228,1252]" pageId="1" type="female">8 ♀</specimenCount>
in 
<collectionCode id="23F5AE0AFFAB652DFE1ABAEF61E3FB63" box="[446,499,1228,1252]" pageId="1">IBN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="455B36CFFFAB652DFF63BACC6109F8CF" blockId="1.[151,1437,868,2009]" pageId="1">
Male imago. Length (mm): body, 10.5–11.0; FW, 10.0–11.0; HW, 4.5–4.7; forelegs, 4.9–5.1; cerci, 27.5–28.5. General coloration yellowish white, shaded with grey dorsally. Head (
<figureCitation id="DDDF2A4AFFAB652DFC1ABB376408FAAB" box="[958,1048,1300,1325]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3A</figureCitation>
) shaded dorsally with grey, some males with lighter coloration on medial line and around median ocellus; vertex with grey shading; some specimens with Y–shaped pale area between ocelli and extending through occiput. Antennae whitish shaded with grey, stronger at margins of scape and pedicel. Thorax (
<figureCitation id="DDDF2A4AFFAB652DFD62BB5C6331FA1F" box="[710,801,1407,1432]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3A</figureCitation>
): pronotum hyaline translucent shaded very slightly with grey on anteromedian zone; mesonotum shaded gray on carinae and on posterior scutal protuberances (PSP); metanotum yellowish, shaded slightly with grey medially. Forelegs whitish shaded completely with purplish grey, lighter on tarsi; legs II–III yellowish translucent with apical black dash (
<figureCitation id="DDDF2A4AFFAB652DFC18BBCF6405F983" box="[956,1045,1516,1541]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3D</figureCitation>
). Wings: veins whitish translucent, shaded with purplish grey at the base of longitudinal veins, shading more extensive on veins (and membrane around them) Sc and R1. Abdomen (
<figureCitation id="DDDF2A4AFFAB652DFD9FB8176288F9CB" box="[571,664,1588,1613]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3A</figureCitation>
): terga shaded with grey along a median band; tergum I lightest, IX darkest; tergum I with a median mark (not broken by a pale area medially; terga II–VI shaded with gray except on medial pale band and pale subcircular mark at each side of medial band (
<figureCitation id="DDDF2A4AFFAB652DFC6DB85F6433F913" box="[969,1059,1660,1685]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3A</figureCitation>
); terga VII–VIII similar but more widely pigmented with gray, and with two pairs of pale circular marks; tergum IX widely pigmented, darker medially, a pair of pale circular marks on anteromedian margin. Abdominal sterna pale not shaded. Genitalia (
<figureCitation id="DDDF2A4AFFAB652DFAC0B8E760ABF887" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="1">Figs. 1A</figureCitation>
, 
<figureCitation id="DDDF2A4AFFAB652DFF63B8CB60C4F887" box="[199,212,1768,1792]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">3</figureCitation>
E–D): yellowish white; pedestals relatively short, with outer corner indented so apical half is directed slightly towards the outer side; main lobes of penes very long and slender, secondary lobe relatively short. Cerci whitish translucent.
</paragraph>
<paragraph id="455B36CFFFAB652EFF63B9776229FF7F" blockId="1.[151,1437,868,2009]" lastBlockId="2.[151,1437,151,392]" lastPageId="2" pageId="1">
Female imago. Length (mm): body, 13.2–19.3; FW, 13.5– 16.0; HW, 6.1–7.7; forelegs broken; cerci, 5.5–6.0. General coloration yellowish (
<figureCitation id="DDDF2A4AFFAB652DFE48B9546254F817" box="[492,580,1911,1936]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3B</figureCitation>
). Head with dorsal grey shading except on median zone of vertex (
<figureCitation id="DDDF2A4AFFAB652DFA92B954659EF817" box="[1334,1422,1911,1936]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3B</figureCitation>
). Thorax: pronotum shaded grey widely, except some pale areas as in 
<figureCitation id="DDDF2A4AFFAB652DFC38B9BF63E3F833" box="[924,1011,1948,1973]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="1">Fig. 3B</figureCitation>
. Mesonotum as male. Foreleg broken off in all specimens, similar to middle and hind legs in shape. Middle and hind legs with apical black dash as in male (
<figureCitation id="DDDF2A4AFFA8652EFF79BEB46125FF37" box="[221,309,151,176]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="2">Fig. 3C</figureCitation>
). Abdomen similar to male except broader pale medial band on terga II–VII (
<figureCitation id="DDDF2A4AFFA8652EFB3EBEB464E2FF37" box="[1178,1266,151,176]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="2">Fig. 3B</figureCitation>
). Sternum VIII with a pair of relatively large anteromedian sockets (
<figureCitation id="DDDF2A4AFFA8652EFD41BE9F6333FF53" box="[741,803,188,213]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="2">Fig. 1</figureCitation>
B–C), sockets partially fused forming a single structure. Caudal filaments whitish translucent.
</paragraph>
<paragraph id="455B36CFFFA8652EFF63BF2664B4FEC7" blockId="2.[151,1437,151,392]" pageId="2">Eggs. Length, 245–275 µm; width, 200–210 µm. Hemispheric, yellowish, with one whitish polar cap formed by a compound thread (ca. 5.0 mm long) formed by numerous thin filaments tightly braided.</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFA8652EFF63BF6F62A5FEE2" box="[199,693,332,357]" pageId="2" type="etymology">
<paragraph id="455B36CFFFA8652EFF63BF6F62A5FEE2" blockId="2.[151,1437,151,392]" box="[199,693,332,357]" pageId="2">
<emphasis id="7790EADDFFA8652EFF63BF6F615EFEE2" bold="true" box="[199,334,332,357]" pageId="2">Etymology.</emphasis>
<collectingRegion id="8720F82DFFA8652EFEF0BF6F61A3FEE2" box="[340,435,332,357]" country="Colombia" name="Vichada" pageId="2">Vichada</collectingRegion>
, from the 
<typeStatus id="9A5F886DFFA8652EFD80BF6E6244FEE2" box="[548,596,333,357]" pageId="2">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFA8652EFF63BF4C62BEFE0F" box="[199,686,367,392]" pageId="2" type="distribution">
<paragraph id="455B36CFFFA8652EFF63BF4C62BEFE0F" blockId="2.[151,1437,151,392]" box="[199,686,367,392]" pageId="2">
<emphasis id="7790EADDFFA8652EFF63BF4C6149FE0F" bold="true" box="[199,345,367,392]" pageId="2">Distribution</emphasis>
(
<figureCitation id="DDDF2A4AFFA8652EFECCBF4C61BDFE0F" box="[360,429,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1026,1048]" captionTargetBox="[152,1435,196,1002]" captionTargetId="figure@7.[151,1436,193,1005]" captionTargetPageId="7" captionText="FIGURE 5. Map indicating the distributional records of the new species." httpUri="https://zenodo.org/record/2617739/files/figure.png" pageId="2">Fig. 5</figureCitation>
). 
<collectingCountry id="3DF3765FFFA8652EFE66BF4C6220FE0F" box="[450,560,367,392]" name="Colombia" pageId="2">Colombia</collectingCountry>
(
<collectingRegion id="8720F82DFFA8652EFD9BBF4C62B2FE0F" box="[575,674,367,392]" country="Colombia" name="Vichada" pageId="2">Vichada</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="119B6647FFA8652EFF33B9A7652AF83D" httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="2" startId="2.[151,264,1924,1947]" targetBox="[163,1436,455,1898]" targetPageId="2">
<paragraph id="455B36CFFFA8652EFF33B9A7652AF83D" blockId="2.[151,1436,1924,1978]" pageId="2">
<emphasis id="7790EADDFFA8652EFF33B9A76142F81C" bold="true" box="[151,338,1924,1947]" pageId="2">FIGURES 1A–G.</emphasis>
<taxonomicName id="82E44D4CFFA8652EFEFFB9A56237F81C" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[347,551,1925,1947]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" phylum="Arthropoda" rank="species" species="vichada" status="sp. nov.">
<emphasis id="7790EADDFFA8652EFEFFB9A56237F81C" box="[347,551,1925,1947]" italics="true" pageId="2">Campsurus vichada</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFA8652EFD94B9A56293F81C" bold="true" box="[560,643,1926,1947]" pageId="2">
<taxonomicNameLabel id="6CA357A6FFA8652EFD94B9A56293F81C" box="[560,643,1926,1947]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. 
<taxonomicName id="82E44D4CFFA8652EFF33B9866176F83D" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[151,358,1956,1978]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" phylum="Arthropoda" rank="species" species="cristales" status="sp. nov.">
<emphasis id="7790EADDFFA8652EFF33B9866176F83D" box="[151,358,1956,1978]" italics="true" pageId="2">Campsurus cristales</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFA8652EFEC8B98761ADF83E" bold="true" box="[364,445,1956,1977]" pageId="2">
<taxonomicNameLabel id="6CA357A6FFA8652EFEC8B98761ADF83E" box="[364,445,1956,1977]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets.
</paragraph>
</caption>
<caption id="119B6647FFA9652FFF33BDE26383FB93" httpUri="https://zenodo.org/record/2617733/files/figure.png" pageId="3" startId="3.[151,250,961,983]" targetBox="[170,1410,193,946]" targetPageId="3">
<paragraph id="455B36CFFFA9652FFF33BDE26383FB93" blockId="3.[151,1436,961,1044]" pageId="3">
<emphasis id="7790EADDFFA9652FFF33BDE26107FC51" bold="true" box="[151,279,961,983]" pageId="3">FIGURE 2.</emphasis>
Shortest parsimony tree obtained with matrix on Appendix 1, using implied weights. Numbers above nodes indicate character numbers that have changed in this clade or terminal (i.e., apomorphies). Larger numbers below nodes indicate group support (GC differences from 300 replicates of jackknifing).
</paragraph>
</caption>
<subSubSection id="0DFE6544FFA9652FFF63BA18611CFA7A" pageId="3" type="discussion">
<paragraph id="455B36CFFFA9652FFF63BA18611BFB39" blockId="3.[151,1437,1083,1534]" pageId="3">
<emphasis id="7790EADDFFA9652FFF63BA18615CFBD3" bold="true" box="[199,332,1083,1108]" pageId="3">Discussion.</emphasis>
<taxonomicName id="82E44D4CFFA9652FFEF1BA1F6228FBD3" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[341,568,1083,1108]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="vichada" status="sp. nov.">
<emphasis id="7790EADDFFA9652FFEF1BA1F6228FBD3" box="[341,568,1083,1108]" italics="true" pageId="3">Campsurus vichada</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFA9652FFDE6BA18628EFBD3" bold="true" box="[578,670,1083,1108]" pageId="3">
<taxonomicNameLabel id="6CA357A6FFA9652FFDE6BA18628EFBD3" box="[578,670,1083,1108]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows characters in common with the two clades in the albifilum group found by 
<bibRefCitation id="21754B3EFFA9652FFEEABA7C6260FBF0" author="Molineri, C. &amp; Salles, F. F." box="[334,624,1119,1144]" pageId="3" pagination="265 - 267" refId="ref4051" refString="Molineri, C. &amp; Salles, F. F. (2013) Phylogeny and biogeography of the ephemeral Campsurus (Ephemeroptera, Polymitarcyidae). Systematic Entomology, 38 (2), 265 - 267. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00656. x" type="journal article" year="2013">Molineri &amp; Salles (2013)</bibRefCitation>
. The large size of the body and the slightly curved pedestals are shared with 
<taxonomicName id="82E44D4CFFA9652FFF70BAA0614EFB1C" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[212,350,1154,1179]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="albifilum">
<emphasis id="7790EADDFFA9652FFF70BAA0614EFB1C" box="[212,350,1154,1179]" italics="true" pageId="3">C. albifilum</emphasis>
</taxonomicName>
and 
<taxonomicName id="82E44D4CFFA9652FFE39BAA0622AFB1C" box="[413,570,1154,1179]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="fuliginatus">
<emphasis id="7790EADDFFA9652FFE39BAA0622AFB1C" box="[413,570,1154,1179]" italics="true" pageId="3">C. fuliginatus</emphasis>
</taxonomicName>
, but the extremely long penes are more similar to 
<taxonomicName id="82E44D4CFFA9652FFB3CBAA06553FB1C" box="[1176,1347,1154,1179]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="gracilipenis">
<emphasis id="7790EADDFFA9652FFB3CBAA06553FB1C" box="[1176,1347,1154,1179]" italics="true" pageId="3">C. gracilipenis</emphasis>
</taxonomicName>
and 
<taxonomicName id="82E44D4CFFA9652FFA26BAA06114FB38" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="homaulos">
<emphasis id="7790EADDFFA9652FFA26BAA06114FB38" italics="true" pageId="3">C. homaulos</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="455B36CFFFA9652FFF63BAE9653FFA13" blockId="3.[151,1437,1083,1534]" pageId="3">
We added the new species to the matrix used in 
<bibRefCitation id="21754B3EFFA9652FFD7FBAEA63E6FB65" author="Molineri, C. &amp; Salles, F. F." box="[731,1014,1225,1250]" pageId="3" pagination="265 - 267" refId="ref4051" refString="Molineri, C. &amp; Salles, F. F. (2013) Phylogeny and biogeography of the ephemeral Campsurus (Ephemeroptera, Polymitarcyidae). Systematic Entomology, 38 (2), 265 - 267. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00656. x" type="journal article" year="2013">Molineri &amp; Salles (2013)</bibRefCitation>
, and the searches resulted in the same topology shown by these authors, with the new species resulting sister to the pair 
<taxonomicName id="82E44D4CFFA9652FFBE7BACD64C2FA81" box="[1091,1234,1261,1286]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="homaulus">
<emphasis id="7790EADDFFA9652FFBE7BACD64C2FA81" box="[1091,1234,1261,1286]" italics="true" pageId="3">C. homaulus</emphasis>
</taxonomicName>
+ 
<taxonomicName id="82E44D4CFFA9652FFB50BACD658CFA81" box="[1268,1436,1261,1286]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="gracilipenis">
<emphasis id="7790EADDFFA9652FFB50BACD658CFA81" box="[1268,1436,1261,1286]" italics="true" pageId="3">C. gracilipenis</emphasis>
</taxonomicName>
(
<figureCitation id="DDDF2A4AFFA9652FFF3BBB3360F7FAAE" box="[159,231,1296,1321]" captionStart="FIGURE 2" captionStartId="3.[151,250,961,983]" captionTargetBox="[170,1410,193,946]" captionTargetId="figure@3.[170,1410,193,946]" captionTargetPageId="3" captionText="FIGURE 2. Shortest parsimony tree obtained with matrix on Appendix 1, using implied weights. Numbers above nodes indicate character numbers that have changed in this clade or terminal (i.e., apomorphies). Larger numbers below nodes indicate group support (GC differences from 300 replicates of jackknifing)." httpUri="https://zenodo.org/record/2617733/files/figure.png" pageId="3">Fig. 2</figureCitation>
), with good support. The three species share two synapomorphies: 1) a change in continuous character number 1 – Pedestal length/width ratio (from 1.12–1.40 to 1.05), thus the general shape of the pedestal change from subrectangular (plesiomorphy) to subquadrate; and 2) a change in color pattern of abdominal terga (character 7, discrete) from the state 0 (with][–shaped marks) to state 1 (marks closed, forming rectangle or circle).
</paragraph>
<paragraph id="455B36CFFFA9652FFF63BBBD611CFA7A" blockId="3.[151,1437,1083,1534]" pageId="3">
The key published in 
<bibRefCitation id="21754B3EFFA9652FFE18BBBD62C8FA30" author="Molineri, C. &amp; Salles, F. F." box="[444,728,1438,1463]" pageId="3" pagination="265 - 267" refId="ref4051" refString="Molineri, C. &amp; Salles, F. F. (2013) Phylogeny and biogeography of the ephemeral Campsurus (Ephemeroptera, Polymitarcyidae). Systematic Entomology, 38 (2), 265 - 267. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00656. x" type="journal article" year="2013">Molineri &amp; Salles (2013)</bibRefCitation>
should be modified as follows to include the new species here described. Please note that couplet 5 was also modified to include new data regarding 
<taxonomicName id="82E44D4CFFA9652FFB3CBBE06553FA5C" box="[1176,1347,1474,1499]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="gracilipenis">
<emphasis id="7790EADDFFA9652FFB3CBBE06553FA5C" box="[1176,1347,1474,1499]" italics="true" pageId="3">C. gracilipenis</emphasis>
</taxonomicName>
and 
<taxonomicName id="82E44D4CFFA9652FFA26BBE06114FA79" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="homaulus">
<emphasis id="7790EADDFFA9652FFA26BBE06114FA79" italics="true" pageId="3">C. homaulus</emphasis>
</taxonomicName>
:
</paragraph>
</subSubSection>
<subSubSection id="0DFE6544FFA9652FFF33B805658BF873" pageId="3" type="key">
<key id="317561FEFFA9652FFF33B805658BF873" pageId="3">
<keyStep id="FE102E6AFFA9652FFF33B805658BF913" pageId="3">
<paragraph id="455B36CFFFA9652FFF33B805658BF9DE" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B805658BF9DE" pageId="3">
1. Abdominal color pattern pale on terga I–III and very dark on terga IV–IX; secondary lobe of penes short, knoblike (see other characters in 
<bibRefCitation id="21754B3EFFA9652FFEC5B8676256F9DD" author="Dominguez, E. &amp; Molineri, C. &amp; Pescador, M. &amp; Hubbard, M. D. &amp; Nieto, C." box="[353,582,1604,1626]" pageId="3" refId="ref3900" refString="Dominguez, E., Molineri, C., Pescador, M., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity in Latin America. (ABLA Series. Uol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">Molineri &amp; Salles 2006</bibRefCitation>
)........................................................... 
<taxonomicName id="82E44D4CFFA9652FFABDB867658BF9DE" box="[1305,1435,1604,1625]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="fuliginatus">
<emphasis id="7790EADDFFA9652FFABDB867658BF9DE" box="[1305,1435,1604,1625]" italics="true" pageId="3">C. fuliginatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="455B36CFFFA9652FFF33B841658BF913" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B841658BF913" pageId="3">
- Abdominal color pattern uniform on terga except darker on posterior terga IX or VIII–IX; secondary lobe of penes distinct (
<figureCitation id="DDDF2A4AFFA9652FFF4CB85D612DF913" box="[232,317,1662,1684]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="3">Figs. 1A</figureCitation>
, 
<figureCitation id="DDDF2A4AFFA9652FFEE3B85D6142F913" box="[327,338,1662,1684]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="3">3</figureCitation>
E–D); FW length variable; pedestals variable..................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE102E6AFFA9652FFF33B8BF658BF96B" pageId="3">
<paragraph id="455B36CFFFA9652FFF33B8BF658CF935" blockId="3.[151,1436,1574,2036]" box="[151,1436,1692,1714]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B8BF658CF935" box="[151,1436,1692,1714]" pageId="3">2. Pedestal subrectangular, length/width ratio = 1.12–1.40; abdominal terga with submedian][–shaped gray mark............ 3</keyLead>
</paragraph>
<paragraph id="455B36CFFFA9652FFF33B899658BF96B" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B899658BF96B" pageId="3">
- Pedestal subquadrate (
<figureCitation id="DDDF2A4AFFA9652FFE18B89A6204F948" box="[444,532,1721,1743]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="3">Figs. 1A</figureCitation>
, 
<figureCitation id="DDDF2A4AFFA9652FFD85B89A6250F948" box="[545,576,1721,1743]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="3">3D</figureCitation>
), length/width ratio = 1.05 or less; abdominal terga more widely pigmented with gray, except small rectangular to ovate pale mark at each side (
<figureCitation id="DDDF2A4AFFA9652FFCA2B8F56342F96B" box="[774,850,1750,1772]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="3">Fig. 3A</figureCitation>
)................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE102E6AFFA9652FFF33B8D7658BF8A0" pageId="3">
<paragraph id="455B36CFFFA9652FFF33B8D7658BF88E" blockId="3.[151,1436,1574,2036]" box="[151,1435,1780,1802]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B8D7658BF88E" box="[151,1435,1780,1802]" pageId="3">
3. Pedestals very long (length/width ratio&gt; 2.1) and with the apical inner margin strongly projected posteriorly...... 
<taxonomicName id="82E44D4CFFA9652FFAE6B8D7658BF88E" box="[1346,1435,1780,1801]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="yavari">
<emphasis id="7790EADDFFA9652FFAE6B8D7658BF88E" box="[1346,1435,1780,1801]" italics="true" pageId="3">C. yavari</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="455B36CFFFA9652FFF33B931658BF8A0" blockId="3.[151,1436,1574,2036]" box="[151,1435,1809,1831]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B931658BF8A0" box="[151,1435,1809,1831]" pageId="3">
- Pedestals shorter (length/width ratio&gt; 1.7), inner apical margin broad, not so strongly projected.............. 
<taxonomicName id="82E44D4CFFA9652FFA8DB931658BF8A0" ID-CoL="QF3J" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[1321,1435,1810,1831]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="albifilum">
<emphasis id="7790EADDFFA9652FFA8DB931658BF8A0" box="[1321,1435,1810,1831]" italics="true" pageId="3">C. albifilum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE102E6AFFA9652FFF33B90D658CF8F8" pageId="3">
<paragraph id="455B36CFFFA9652FFF33B90D658CF8E5" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B90D658CF8E5" pageId="3">
4. Body and wings large to medium–sized (FW length&gt; 
<quantity id="821C9B2AFFA9652FFD40B90D6321F8C3" box="[740,817,1838,1860]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" unit="mm" value="9.5">9.5 mm</quantity>
); secondary lobe of penes reduced, short (
<figureCitation id="DDDF2A4AFFA9652FFB67B90D6508F8C3" box="[1219,1304,1838,1860]" captionStart="FIGURES 1" captionStartId="2.[151,264,1924,1947]" captionTargetBox="[163,1436,455,1898]" captionTargetId="figure@2.[151,1436,447,1901]" captionTargetPageId="2" captionText="FIGURES 1A–G. Campsurus vichada sp. nov.: A, male genitalia, v.v.; B, female sternum VIII, C, same, detail of sockets. Campsurus cristales sp. nov.: D, male genitalia, v.v.; E, same, l.v.; F, female sternum VIII, G, same, detail of sockets." httpUri="https://zenodo.org/record/2617731/files/figure.png" pageId="3">Figs. 1A</figureCitation>
, 
<figureCitation id="DDDF2A4AFFA9652FFA87B90D653DF8C3" box="[1315,1325,1838,1860]" captionStart="FIGURES 3" captionStartId="4.[151,264,1948,1971]" captionTargetBox="[201,1383,195,1924]" captionTargetId="figure@4.[201,1383,195,1924]" captionTargetPageId="4" captionText="FIGURES 3A–D. Campsurus vichada sp. nov.: A, male imago, dorsal view (PSP= Postero Scutal Protuberances); B, female imago, d.v.; C, female mesosternum; D, male mesosternum (mfp= mesofurcasternal plates); E, male genitalia, v.v. (ml= main lobe; sl= secondary lobe); D, detail of pedestal." httpUri="https://zenodo.org/record/2617735/files/figure.png" pageId="3">3</figureCitation>
D–E).................................................................................. 
<taxonomicName id="82E44D4CFFA9652FFB2EB96F6559F8E6" authority="Molineri &amp; Granados-Martinez, 2019" authorityName="Molineri &amp; Granados-Martinez" authorityYear="2019" box="[1162,1353,1868,1889]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="vichada" status="sp. nov.">
<emphasis id="7790EADDFFA9652FFB2EB96F6559F8E6" box="[1162,1353,1868,1889]" italics="true" pageId="3">Campsurus vichada</emphasis>
</taxonomicName>
<emphasis id="7790EADDFFA9652FFAEBB96F658CF8E5" bold="true" box="[1359,1436,1868,1890]" pageId="3">
<taxonomicNameLabel id="6CA357A6FFA9652FFAEBB96F658CF8E5" box="[1359,1436,1868,1890]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="455B36CFFFA9652FFF33B949658CF8F8" blockId="3.[151,1436,1574,2036]" box="[151,1436,1897,1919]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B949658CF8F8" box="[151,1436,1897,1919]" pageId="3">- Body and wings medium to small in size (FW length &lt;9.0 mm); secondary lobe of penes relatively long................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE102E6AFFA9652FFF33B9A5658BF873" pageId="3">
<paragraph id="455B36CFFFA9652FFF33B9A5658BF83E" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B9A5658BF83E" pageId="3">
5. Penes long and slender (length/width mostly&gt; 0.85); basal inner membrane of penes very short in relation to total length of penean lobe (ratio &lt;0.28).................................................................... 
<taxonomicName id="82E44D4CFFA9652FFAA9B987658BF83E" box="[1293,1435,1956,1977]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="gracilipenis">
<emphasis id="7790EADDFFA9652FFAA9B987658BF83E" box="[1293,1435,1956,1977]" italics="true" pageId="3">C. gracilipenis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="455B36CFFFA9652FFF33B9E1658BF873" blockId="3.[151,1436,1574,2036]" pageId="3">
<keyLead id="FE1595FAFFA9652FFF33B9E1658BF873" pageId="3">
- Penes shorter (length/width mostly &lt;0.80); basal inner membrane of penes not so short in relation to total length of penean lobe (ratio&gt;0.32)............................................................................. 
<taxonomicName id="82E44D4CFFA9652FFA86B9FC658BF873" box="[1314,1435,2015,2036]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" phylum="Arthropoda" rank="species" species="homaulos">
<emphasis id="7790EADDFFA9652FFA86B9FC658BF873" box="[1314,1435,2015,2036]" italics="true" pageId="3">C. homaulos</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>