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<document ID-DOI="http://doi.org/10.5281/zenodo.6079861" ID-GBIF-Dataset="54ea121d-32e1-4bc3-b503-6e54c101b3ed" ID-GBIF-Taxon="121268866" ID-Zenodo-Dep="6079861" checkinTime="1467565389598" checkinUser="plazi" docAuthor="Mark K. L. Wong &amp; Benoit Guénard" docDate="2016" docId="AF766F7D0F3CFF92FCC6FB35FC687824" docLanguage="en" docName="ANZ2016.66.1.002.pdf" docOrigin="Annales Zoologici 66 (1)" docStyle="DocumentStyle{}" docTitle="Aenictus seletarius Wong et Guenard, sp. nov." docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="41" masterDocId="534F17050F3FFF95FF9CFFDEFFBC7903" masterDocTitle="Aenictus seletarius, a New Species of Hypogaeic Army Ant from Singapore, with an Updated Key to the Aenictus minutulus Species Group (Hymenoptera: Formicidae: Dorylinae) from Southeast Asia" masterLastPageNumber="42" masterPageNumber="35" pageId="3" pageNumber="37" updateTime="1644861771794" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Aenictus seletarius, a New Species of Hypogaeic Army Ant from Singapore, with an Updated Key to the Aenictus minutulus Species Group (Hymenoptera: Formicidae: Dorylinae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mark K. L. Wong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benoit Guénard</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Annales Zoologici</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>35</mods:start>
<mods:end>42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3161/00034541Anz2016.66.1.002</mods:identifier>
<mods:identifier type="GBIF-Dataset">54ea121d-32e1-4bc3-b503-6e54c101b3ed</mods:identifier>
<mods:identifier type="Zenodo-Dep">269444</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6079861" ID-GBIF-Taxon="121268866" ID-Zenodo-Dep="6079861" LSID="urn:lsid:plazi:treatment:AF766F7D0F3CFF92FCC6FB35FC687824" httpUri="http://treatment.plazi.org/id/AF766F7D0F3CFF92FCC6FB35FC687824" lastPageId="7" lastPageNumber="41" pageId="3" pageNumber="37">
<subSubSection pageId="3" pageNumber="37" type="nomenclature">
<paragraph blockId="3.[858,1402,1259,1312]" box="[858,1402,1259,1281]" pageId="3" pageNumber="37">
<heading box="[858,1402,1259,1281]" centered="true" fontSize="9" level="6" pageId="3" pageNumber="37" reason="4">
<taxonomicName authority="Wong et Guenard" authorityName="Wong et Guenard" box="[858,1306,1259,1281]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="seletarius" status="sp. nov.">
<emphasis bold="true" box="[858,1099,1259,1281]" italics="true" pageId="3" pageNumber="37">Aenictus seletarius</emphasis>
Wong et Guénard
</taxonomicName>
<taxonomicNameLabel box="[1314,1402,1260,1281]" pageId="3" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="3.[858,1402,1259,1312]" box="[1082,1177,1290,1312]" pageId="3" pageNumber="37">
<figureCitation box="[1082,1177,1290,1312]" captionStart-0="Figures 2 – 4" captionStart-1="Figures 5 – 6" captionStartId-0="5.[143,211,1859,1876]" captionStartId-1="6.[570,639,1381,1398]" captionTargetBox-0="[347,1225,199,1814]" captionTargetBox-1="[570,1442,781,1363]" captionTargetId-0="figure@5.[346,1221,794,1300]" captionTargetId-1="figure@6.[569,1443,781,1364]" captionTargetPageId-0="5" captionText-0="Figures 2 – 4. Aenictus seletarius (holotype, ANTWEB 1009000, LKCNHM) from Singapore. (2) Profile view; (3) dorsal view; (4) profile view focusing" captionText-1="Figures 5 – 6. Aenictus seletarius (holotype, ANTWEB 1009000, LKCNHM) from Singapore. (5) Full-face view; (6) head view focusing on mandibles." httpUri-0="https://zenodo.org/record/269446/files/figure.png" httpUri-1="https://zenodo.org/record/269447/files/figure.png" pageId="3" pageNumber="37">
Figs
<number box="[1138,1177,1290,1312]" pageId="3" pageNumber="37" value="4.0" valueMax="6.0" valueMin="2.0">2–6</number>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="37" type="description">
<paragraph blockId="3.[816,1444,1352,1901]" pageId="3" pageNumber="37">
<emphasis bold="true" box="[854,936,1352,1374]" italics="true" pageId="3" pageNumber="37">
<typeStatus box="[854,928,1352,1373]" pageId="3" pageNumber="37">Types</typeStatus>
.
</emphasis>
<typeStatus box="[942,1044,1353,1375]" pageId="3" pageNumber="37">Holotype</typeStatus>
.Worker from
<collectionCode box="[1203,1351,1353,1374]" pageId="3" pageNumber="37">
<collectingCountry box="[1203,1351,1353,1374]" name="Singapore" pageId="3" pageNumber="37">SINGAPORE</collectingCountry>
</collectionCode>
, Seletar Trail,
<geoCoordinate box="[891,965,1384,1405]" direction="north" orientation="latitude" pageId="3" pageNumber="37" precision="925" value="1.3833333">
<number box="[891,900,1384,1405]" pageId="3" pageNumber="37" value="1.0">1</number>
°
<number box="[909,938,1384,1405]" pageId="3" pageNumber="37" value="23.0">23</number>
’N
</geoCoordinate>
;
<geoCoordinate box="[982,1082,1384,1405]" direction="east" orientation="longitude" pageId="3" pageNumber="37" precision="925" value="103.8">
<number box="[982,1023,1384,1405]" pageId="3" pageNumber="37" value="103.0">103</number>
°
<number box="[1031,1060,1384,1405]" pageId="3" pageNumber="37" value="48.0">48</number>
’E
</geoCoordinate>
, ca.
<quantity box="[1143,1193,1384,1405]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="37" unit="m" value="40.0">40m</quantity>
, subterranean pitfall trap,
<date box="[879,999,1414,1436]" pageId="3" pageNumber="37" value="2015-07-25">
<number box="[879,906,1414,1436]" pageId="3" pageNumber="37" value="25.0">25</number>
.vii.
<number box="[942,999,1414,1436]" pageId="3" pageNumber="37" value="2015.0">2015</number>
</date>
(Mark K. L. Wong), label “
<accessionNumber box="[1310,1436,1414,1436]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW250715" pageId="3" pageNumber="37">
MW
<number box="[1358,1436,1414,1436]" pageId="3" pageNumber="37" value="250715.0">
<date box="[1358,1436,1414,1436]" pageId="3" pageNumber="37" value="15-07-25">250715</date>
</number>
</accessionNumber>
-
<number box="[816,849,1446,1467]" pageId="3" pageNumber="37" value="1.1">1.1</number>
” (ANTWEB
<number box="[993,1086,1446,1468]" pageId="3" pageNumber="37" value="1009000.0">1009000</number>
); deposited in LKCNHM.
<typeStatus pageId="3" pageNumber="37">Paratypes</typeStatus>
. Three workers in total, all with the same collection data as
<typeStatus box="[960,1060,1508,1530]" pageId="3" pageNumber="37">holotype</typeStatus>
; deposited at SBSHKU.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="38" pageId="3" pageNumber="37" type="diagnosis">
<paragraph blockId="3.[816,1444,1352,1901]" pageId="3" pageNumber="37">
<emphasis bold="true" box="[854,987,1538,1560]" italics="true" pageId="3" pageNumber="37">Diagnosis.</emphasis>
Worker caste with important size variation. Head almost as wide as long, with side margins broadly convex. Masticatory margin of mandibles, medium-sized subapical tooth followed posteriorly by a distinct medium-sized denticle, and both the subapical tooth and the posterior denticle are of similar size. Posterodorsal corner of the propodeum strongly angular and followed by a concave propodeal declivity. Subpetiolar process well-developed and plough-shaped.
</paragraph>
<paragraph blockId="3.[816,1444,1352,1901]" lastBlockId="4.[143,772,180,1246]" lastPageId="4" lastPageNumber="38" pageId="3" pageNumber="37">
Measurements and indices.
<typeStatus box="[1179,1284,1818,1840]" pageId="3" pageNumber="37">Holotype</typeStatus>
: HL
<quantity box="[1340,1439,1817,1839]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="3" pageNumber="37" unit="mm" value="0.57">0.57 mm</quantity>
; HW
<quantity box="[866,965,1848,1870]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="37" unit="mm" value="0.52">0.52 mm</quantity>
; MaL
<quantity box="[1034,1133,1849,1870]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="37" unit="mm" value="0.33">0.33 mm</quantity>
; SL
<quantity box="[1183,1282,1849,1870]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="37" unit="mm" value="0.33">0.33 mm</quantity>
; WL
<quantity box="[1341,1439,1849,1870]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="37" unit="mm" value="0.82">0.82 mm</quantity>
; PNW
<quantity box="[882,983,1880,1901]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="37" unit="mm" value="0.32">0.32 mm</quantity>
;
<collectionCode box="[996,1048,1880,1901]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" name="National Museum" pageId="3" pageNumber="37">PNH</collectionCode>
<quantity box="[1058,1158,1880,1901]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="37" unit="mm" value="0.29">0.29 mm</quantity>
; MW
<quantity box="[1225,1326,1880,1901]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="37" unit="mm" value="0.17">0.17 mm</quantity>
; PTL
<quantity lastPageId="4" lastPageNumber="38" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="37" unit="mm" value="0.24">0.24 mm</quantity>
; PTW
<quantity box="[262,360,180,202]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="38" unit="mm" value="0.17">0.17 mm</quantity>
;
<collectionCode box="[371,422,181,202]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13854" name="Perth Museum and Art Gallery" pageId="4" pageNumber="38">PTH</collectionCode>
<quantity box="[429,528,180,202]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="38" unit="mm" value="0.27">0.27 mm</quantity>
; TL
<quantity box="[578,673,180,202]" metricMagnitude="-3" metricUnit="m" metricValue="2.83" pageId="4" pageNumber="38" unit="mm" value="2.83">2.83 mm</quantity>
(stinger not included);
<collectionCode box="[314,361,211,232]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12552" name="Agricultural Development and Advisory Service" pageId="4" pageNumber="38">PPL</collectionCode>
<quantity box="[371,472,211,232]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="38" unit="mm" value="0.19">0.19 mm</quantity>
; PPW
<quantity box="[550,651,211,233]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="38" unit="mm" value="0.16">0.16 mm</quantity>
; PPH
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="38" unit="mm" value="0.23">0.23 mm</quantity>
; CI
<number box="[231,260,242,263]" pageId="4" pageNumber="38" value="92.0">92</number>
, SI
<number box="[301,330,241,263]" pageId="4" pageNumber="38" value="64.0">64</number>
,
<date box="[341,420,241,263]" pageId="4" pageNumber="38" value="1965-05">
MaI
<number box="[391,420,241,263]" pageId="4" pageNumber="38" value="65.0">65</number>
</date>
, PI
<number box="[462,491,242,263]" pageId="4" pageNumber="38" value="70.0">70</number>
,
<collectionCode box="[502,542,242,263]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14887" name="National Pingtung University of Science and Technology" pageId="4" pageNumber="38">PPI</collectionCode>
<number box="[548,577,242,263]" pageId="4" pageNumber="38" value="81.0">81</number>
.
<typeStatus box="[588,704,242,263]" pageId="4" pageNumber="38">Paratypes</typeStatus>
(n=
<number box="[756,770,242,263]" pageId="4" pageNumber="38" value="3.0">3</number>
measured): HL
<quantity box="[325,484,272,294]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="6.3" metricValueMin="4.6" pageId="4" pageNumber="38" unit="mm" value="0.545" valueMax="0.63" valueMin="0.46">0.46–0.63 mm</quantity>
; HW
<quantity box="[548,707,272,294]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="6.0" metricValueMin="4.2" pageId="4" pageNumber="38" unit="mm" value="0.51" valueMax="0.6" valueMin="0.42">0.42–0.60 mm</quantity>
; MaL
<quantity box="[143,307,303,325]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.5999999999999996" metricValueMin="2.7" pageId="4" pageNumber="38" unit="mm" value="0.315" valueMax="0.36" valueMin="0.27">0.27–0.36 mm</quantity>
; SL
<quantity box="[369,533,303,325]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.7" metricValueMin="2.6" pageId="4" pageNumber="38" unit="mm" value="0.315" valueMax="0.37" valueMin="0.26">0.26–0.37 mm</quantity>
; WL
<quantity box="[603,767,303,325]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="9.1" metricValueMin="6.7" pageId="4" pageNumber="38" unit="mm" value="0.79" valueMax="0.91" valueMin="0.67">0.67–0.91 mm</quantity>
; PNW
<quantity box="[208,367,333,355]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.5999999999999996" metricValueMin="2.6" pageId="4" pageNumber="38" unit="mm" value="0.31" valueMax="0.36" valueMin="0.26">0.26–0.36 mm</quantity>
;
<collectionCode box="[378,430,334,355]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" name="National Museum" pageId="4" pageNumber="38">PNH</collectionCode>
<quantity box="[438,597,334,355]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="4" pageNumber="38" unit="mm" value="0.28" valueMax="0.34" valueMin="0.22">0.22–0.34 mm</quantity>
; PTL
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.3" metricValueMax="2.7" metricValueMin="1.9" pageId="4" pageNumber="38" unit="mm" value="0.23" valueMax="0.27" valueMin="0.19">0.19–0.27 mm</quantity>
; PTW
<quantity box="[284,456,364,386]" metricMagnitude="-4" metricUnit="m" metricValue="1.65" metricValueMax="1.9" metricValueMin="1.4" pageId="4" pageNumber="38" unit="mm" value="0.165" valueMax="0.19" valueMin="0.14">0.14–0.19 mm</quantity>
;
<collectionCode box="[476,529,365,386]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13854" name="Perth Museum and Art Gallery" pageId="4" pageNumber="38">PTH</collectionCode>
<quantity box="[545,717,364,386]" metricMagnitude="-4" metricUnit="m" metricValue="2.8000000000000003" metricValueMax="3.3000000000000003" metricValueMin="2.3000000000000003" pageId="4" pageNumber="38" unit="mm" value="0.28" valueMax="0.33" valueMin="0.23">0.23–0.33 mm</quantity>
; TL
<quantity box="[143,300,395,416]" metricMagnitude="-3" metricUnit="m" metricValue="2.745" metricValueMax="3.18" metricValueMin="2.31" pageId="4" pageNumber="38" unit="mm" value="2.745" valueMax="3.18" valueMin="2.31">2.31–3.18 mm</quantity>
(sting not included);
<collectionCode box="[551,598,395,416]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12552" name="Agricultural Development and Advisory Service" pageId="4" pageNumber="38">PPL</collectionCode>
<quantity box="[607,767,395,416]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="4" pageNumber="38" unit="mm" value="0.2" valueMax="0.23" valueMin="0.17">0.17–0.23 mm</quantity>
; PPW
<quantity box="[210,372,426,447]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7999999999999998" metricValueMin="1.4" pageId="4" pageNumber="38" unit="mm" value="0.16" valueMax="0.18" valueMin="0.14">0.14–0.18 mm</quantity>
; PPH
<quantity box="[449,611,426,447]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.8" metricValueMin="2.0" pageId="4" pageNumber="38" unit="mm" value="0.24" valueMax="0.28" valueMin="0.2">0.20–0.28 mm</quantity>
; CI
<number box="[663,731,426,448]" pageId="4" pageNumber="38" value="93.5" valueMax="96.0" valueMin="91.0">91–96</number>
, SI
<number box="[143,212,456,478]" pageId="4" pageNumber="38" value="62.5" valueMax="64.0" valueMin="61.0">61–64</number>
,
<date box="[223,301,456,478]" pageId="4" pageNumber="38" value="1959-05">
MaI
<number box="[274,301,456,478]" pageId="4" pageNumber="38" value="61.5" valueMax="64.0" valueMin="59.0">59</number>
</date>
–64, PI
<number box="[385,453,456,478]" pageId="4" pageNumber="38" value="71.5" valueMax="74.0" valueMin="69.0">69–74</number>
,
<collectionCode box="[465,505,457,478]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14887" name="National Pingtung University of Science and Technology" pageId="4" pageNumber="38">PPI</collectionCode>
<number box="[513,581,456,477]" pageId="4" pageNumber="38" value="80.5" valueMax="83.0" valueMin="78.0">78–83</number>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="40" pageId="4" pageNumber="38" type="description">
<paragraph blockId="4.[143,772,180,1246]" pageId="4" pageNumber="38">
<emphasis bold="true" box="[181,423,487,509]" pageId="4" pageNumber="38">Worker description</emphasis>
.
<emphasis bold="true" box="[439,509,487,508]" pageId="4" pageNumber="38">Head.</emphasis>
Head in full-face view almost as wide as long (CI
<number box="[452,523,518,540]" pageId="4" pageNumber="38" value="93.5" valueMax="96.0" valueMin="91.0">91–96</number>
), side margins broadly convex, posterior margin slightly convex to almost straight and approximately
<number box="[478,487,576,590]" pageId="4" pageNumber="38" value="3.0">3</number>
/
<number box="[495,504,592,606]" pageId="4" pageNumber="38" value="4.0">4</number>
of HW, posterior corners broadly rounded. Antennal scape curved and enlarged in their posterior half, relative size to head moderate (SI
<number box="[297,368,671,693]" pageId="4" pageNumber="38" value="62.5" valueMax="64.0" valueMin="61.0">61–64</number>
), slightly extending to over the midpoint of head length; antennal segments longer than broad; length of segments II–IX continuously increasing; apical segment X longer than VIII and IX combined; last two segments forming indistinct club. Frontal carina distinct, surpassing posterior margin of antennal torulus. Clypeus short, its anterior margin convex and without denticles. Basal margin of mandible with denticles that gradually reduce in size toward base of mandible. Masticatory margin of mandible with large acute apical tooth, followed posteriorly by a medium-sized subapical tooth, one mediumsized denticle and one small denticle, a medium-sized basal tooth; basal margin with
<number box="[499,538,1070,1091]" pageId="4" pageNumber="38" value="3.5" valueMax="4.0" valueMin="3.0">3–4</number>
small denticles.
</paragraph>
<paragraph blockId="4.[143,772,180,1246]" lastBlockId="4.[816,1444,180,1246]" pageId="4" pageNumber="38">
<emphasis bold="true" box="[181,318,1101,1122]" pageId="4" pageNumber="38">Mesosoma.</emphasis>
In profile, promesonotum convex, sloping gradually to the metanotal groove; mesopleuron relatively short, demarcated from metapleuron by distinct groove. In profile, dorsal outline of propodeum flat to weakly convex nearing the posterior corner. Posterior part of propodeum forming the propodeal declivity nearly at right angle with propodeal dorsum, and separated from the latter by an angular edge; overhanging declivity of propodeum is strongly concave and encircled with thin but distinct rim. Metapleural gland bulla well-developed, its maximum diameter about
<number box="[1082,1115,364,385]" pageId="4" pageNumber="38" value="1.3">1.3</number>
times as long as distance between propodeal spiracle and most proximate part of metapleural gland bulla.
</paragraph>
<paragraph blockId="4.[816,1444,180,1246]" pageId="4" pageNumber="38">
<emphasis bold="true" box="[854,988,456,477]" pageId="4" pageNumber="38">Metasoma.</emphasis>
In profile, petiole excluding subpetiolar process slightly higher than long and with triangular shape; petiole node with steep anterior face and broadly convex dorsal outline; subpetiolar process welldeveloped and of an irregular quadrilateral shape (plough-shaped) with roughly angular apex posteriorly oriented and a slightly concave posterior lateral margin. Size of subpetiolar process approximately
<number box="[1384,1393,668,682]" pageId="4" pageNumber="38" value="1.0">1</number>
/
<number box="[1401,1410,684,698]" pageId="4" pageNumber="38" value="5.0">5</number>
of petiole height and
<number box="[1034,1043,699,713]" pageId="4" pageNumber="38" value="2.0">2</number>
/
<number box="[1050,1059,714,728]" pageId="4" pageNumber="38" value="5.0">5</number>
of petiole length, its ventral outline broadly convex and its ventralmost part with thin almost transparent lamella. In profile postpetiole has a square shape with rounded corners; dorsal outline of postpetiole node flat to weakly convex; postpetiolar process developed and pointing anteriorly with round- ed to weakly angular apex. First gastral tergite and sternite long, extending over half the total length of the gaster.
</paragraph>
<paragraph blockId="4.[816,1444,180,1246]" lastBlockId="5.[143,1444,1858,1902]" lastPageId="5" lastPageNumber="39" pageId="4" pageNumber="38">
<emphasis bold="true" box="[854,985,978,1000]" pageId="4" pageNumber="38">Sculpture.</emphasis>
Head entirely smooth and shiny. Mandibles superficially striate at the base. Basal portion of antennal scape (approximately
<number box="[1260,1269,1037,1051]" pageId="4" pageNumber="38" value="1.0">1</number>
/
<number box="[1276,1285,1052,1066]" pageId="4" pageNumber="38" value="3.0">3</number>
of SL) reticulate transitioning to smooth and shiny on its last
<number box="[1418,1427,1067,1081]" pageId="4" pageNumber="38" value="2.0">2</number>
/
<number box="[1434,1443,1083,1097]" pageId="4" pageNumber="38" value="3.0">3</number>
portion. Mesosoma finely reticulate with exception of pronotum and parts of metapleuron; pronotum smooth and shiny on dorsum and sides but finely reticulate towards the posterior edge; metapleuron smooth and shiny on anterior median portion but otherwise finely on the propodeum, petiole and subpetiole.
</paragraph>
<caption httpUri="https://zenodo.org/record/269445/files/figure.png" pageId="4" pageNumber="38" targetBox="[153,1443,1318,1841]" targetPageId="4">
<paragraph blockId="4.[143,1443,1857,1902]" pageId="4" pageNumber="38">
Figure 1. Left side map: Known global distribution of
<taxonomicName box="[617,701,1859,1876]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[617,701,1859,1876]" italics="true" pageId="4" pageNumber="38">Aenictus</emphasis>
</taxonomicName>
species (modified from antmaps.org). Right side box: Distribution of
<taxonomicName box="[1316,1443,1859,1876]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[1316,1443,1859,1876]" italics="true" pageId="4" pageNumber="38">A. minutulus</emphasis>
</taxonomicName>
group species in Southeast Asia with the inclusion of
<taxonomicName box="[624,746,1884,1901]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[624,746,1884,1901]" italics="true" pageId="4" pageNumber="38">A. seletarius</emphasis>
</taxonomicName>
from Singapore (modified after Jaitrong and Hashimoto 2012, Jaitrong 2015).
</paragraph>
</caption>
<caption box="[143,1444,1858,1877]" httpUri="https://zenodo.org/record/269446/files/figure.png" pageId="5" pageNumber="39" targetBox="[347,1225,199,1814]" targetPageId="5">
<paragraph blockId="5.[143,1444,1858,1902]" box="[143,1444,1858,1877]" pageId="5" pageNumber="39">
Figures 2–4.
<taxonomicName box="[257,442,1859,1876]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[257,442,1859,1876]" italics="true" pageId="5" pageNumber="39">
<taxonomicName box="[257,341,1859,1876]" class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="39" phylum="Arthropoda" rank="genus">Aenictus</taxonomicName>
seletarius
</emphasis>
</taxonomicName>
(holotype, ANTWEB1009000, LKCNHM) from Singapore. (2) Profile view; (3) dorsal view; (4) profile view focusing
</paragraph>
</caption>
<paragraph blockId="6.[143,539,180,1373]" pageId="6" pageNumber="40">reticulate. Petiole including sub- petiolar process finely reticulate with the exception of a smooth and shiny spot anterodorsally. Postpetiole finely reticulate, with flat surface on dorsum smooth and shiny. Gaster entirely smooth and shiny. Legs entirely smooth and shiny.</paragraph>
<paragraph blockId="6.[143,539,180,1373]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,334,456,477]" pageId="6" pageNumber="40">Pubescence.</emphasis>
Head and body, except sides of mesosoma, with abundant suberect standing hairs with lengths of
<quantity box="[314,440,548,570]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="6" pageNumber="40" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
on head dorsum and
<quantity box="[294,438,579,600]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="6" pageNumber="40" unit="mm" value="0.115" valueMax="0.13" valueMin="0.1">0.1–0.13 mm</quantity>
on dor- sum of meso- and metasoma. Shorter decumbent pubescence also present in between longer hairs. Antennal scapes and legs with abundant, decumbent pilosity.
</paragraph>
<paragraph blockId="6.[143,539,180,1373]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,330,763,785]" pageId="6" pageNumber="40">Colouration.</emphasis>
Dark amber colo- uration on head, most of antennae, mesosoma, petiole and most of postpetiole, with darkest brown colouration on the reticulated propodeum. Tip of antennal seg- ment X, entire legs, entire gaster and dorsum of postpetiole node with lighter yellow colouration.
</paragraph>
<paragraph blockId="6.[143,539,180,1373]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,271,1041,1063]" pageId="6" pageNumber="40">Castes.</emphasis>
Worker caste displays variation in body size. Apart from size variation, values of the differ- ent measurement indices are gen- erally consistent among the work- ers measured, thus indicating an absence of allometric growth. Oth- er morphological features such as sculpture, pubescence and colour- ation remain constant among the specimens examined. Male and
</paragraph>
<paragraph blockId="6.[143,385,1382,1434]" box="[143,385,1382,1404]" pageId="6" pageNumber="40">female are unknown.</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/269447/files/figure.png" pageId="6" pageNumber="40" targetBox="[570,1442,781,1363]" targetPageId="6">
<paragraph blockId="6.[570,1445,1379,1424]" box="[570,1445,1379,1398]" pageId="6" pageNumber="40">
Figures
<number box="[651,685,1381,1398]" pageId="6" pageNumber="40" value="5.5" valueMax="6.0" valueMin="5.0">5–6</number>
.
<taxonomicName box="[700,896,1381,1398]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[700,896,1381,1398]" italics="true" pageId="6" pageNumber="40">Aenictus seletarius</emphasis>
</taxonomicName>
(
<typeStatus box="[915,995,1381,1398]" pageId="6" pageNumber="40">holotype</typeStatus>
, ANTWEB
<number box="[1100,1171,1381,1398]" pageId="6" pageNumber="40" value="1009000.0">1009000</number>
, LKCNHM) from
<collectingCountry box="[1347,1441,1381,1398]" name="Singapore" pageId="6" pageNumber="40">Singapore</collectingCountry>
.
</paragraph>
<subSubSection pageId="6" pageNumber="40" type="etymology">
<paragraph blockId="6.[143,1256,1405,1466]" lastBlockId="6.[143,771,1474,1899]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,325,1412,1434]" italics="true" pageId="6" pageNumber="40">Etymology.</emphasis>
The species epi- thet is derived from the collection locality, Seletar forest, which contains some of Singa- pore’s last primary and old secondary habitats. The species epithet is a noun, and thus invariant.
</paragraph>
</subSubSection>
<paragraph blockId="6.[143,1256,1405,1466]" box="[759,1256,1405,1424]" pageId="6" pageNumber="40">
(
<number box="[766,777,1405,1424]" pageId="6" pageNumber="40" value="5.0">5</number>
) Full-face view; (
<number box="[931,943,1405,1424]" pageId="6" pageNumber="40" value="6.0">6</number>
) head view focusing on mandibles.
</paragraph>
</caption>
<subSubSection pageId="6" pageNumber="40" type="distribution">
<paragraph blockId="6.[143,771,1474,1899]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,345,1567,1589]" italics="true" pageId="6" pageNumber="40">Distribution.</emphasis>
Southeast Asia. Only known from
<collectingCountry box="[143,263,1598,1620]" name="Singapore" pageId="6" pageNumber="40">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="40" type="biology_ecology">
<paragraph blockId="6.[143,771,1474,1899]" lastBlockId="6.[816,1444,1474,1902]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[181,541,1628,1651]" italics="true" pageId="6" pageNumber="40">
Ecology.
<taxonomicName box="[298,541,1629,1651]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">Aenictus seletarius</taxonomicName>
</emphasis>
was collected from a tropical lowland primary and old growth secondary rainforest in
<collectingCountry box="[292,409,1691,1713]" name="Singapore" pageId="6" pageNumber="40">Singapore</collectingCountry>
proximally located (&lt;
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) to a freshwater catchment. As individuals were collected with subterranean pitfall traps
<date box="[512,581,1753,1775]" pageId="6" pageNumber="40" value="2015-09">
set
<quantity box="[555,581,1753,1775]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="40" unit="cm" value="15.0">15</quantity>
</date>
cm beneath the soil surface,
<taxonomicName box="[293,451,1783,1805]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[293,451,1783,1805]" italics="true" pageId="6" pageNumber="40">A. seletarius</emphasis>
</taxonomicName>
likely exhibits a hypogaeic lifestyle similar to many other
<taxonomicName box="[511,619,1815,1836]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[511,619,1815,1836]" italics="true" pageId="6" pageNumber="40">Aenictus</emphasis>
</taxonomicName>
species. Ad- ditionally, we found over thirty specimens of a small (TL ca.
<quantity box="[250,326,1877,1898]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="40" unit="mm" value="4.0">4 mm</quantity>
), eyeless unidentified
<taxonomicName box="[605,771,1876,1898]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[605,771,1876,1898]" italics="true" pageId="6" pageNumber="40">Pseudolasius</emphasis>
</taxonomicName>
species in the same traps in which the
<taxonomicName box="[1284,1443,1474,1496]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[1284,1443,1474,1496]" italics="true" pageId="6" pageNumber="40">A. seletarius</emphasis>
</taxonomicName>
individuals were collected.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="41" pageId="6" pageNumber="40" type="discussion">
<paragraph blockId="6.[816,1444,1474,1902]" lastBlockId="7.[143,771,180,1902]" lastPageId="7" lastPageNumber="41" pageId="6" pageNumber="40">
<emphasis bold="true" box="[854,973,1537,1559]" pageId="6" pageNumber="40">
<emphasis bold="true" box="[854,965,1537,1559]" italics="true" pageId="6" pageNumber="40">Remarks</emphasis>
.
</emphasis>
The new species
<taxonomicName box="[1182,1340,1537,1559]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[1182,1340,1537,1559]" italics="true" pageId="6" pageNumber="40">A. seletarius</emphasis>
</taxonomicName>
displays substantial variation in body size among workers (TL
<quantity box="[816,975,1600,1621]" metricMagnitude="-3" metricUnit="m" metricValue="2.745" metricValueMax="3.18" metricValueMin="2.31" pageId="6" pageNumber="40" unit="mm" value="2.745" valueMax="3.18" valueMin="2.31">2.31–3.18 mm</quantity>
). This was also observed by Jaitrong and
<bibRefCitation author="Jaitrong, W. &amp; Y. Hashimoto" box="[816,1023,1631,1653]" journalOrPublisher="Zootaxa" pageId="6" pageNumber="40" pagination="29 - 44" part="3426" refString="Jaitrong, W. and Y. Hashimoto. 2012. Revision of the Aenictus minutulus species group (Hymenoptera: Formicidae: Aenictinae) from Southeast Asia. Zootaxa, 3426: 29 - 44." title="Revision of the Aenictus minutulus species group (Hymenoptera: Formicidae: Aenictinae) from Southeast Asia" type="journal article" year="2012">
Hashimoto (
<number box="[958,1015,1631,1653]" pageId="6" pageNumber="40" value="2012.0">2012</number>
)
</bibRefCitation>
in
<taxonomicName box="[1068,1237,1630,1652]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[1068,1237,1630,1652]" italics="true" pageId="6" pageNumber="40">A. minutulus</emphasis>
</taxonomicName>
(TL
<quantity box="[1301,1436,1631,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="6" pageNumber="40" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.7–2.4 mm</quantity>
) and
<taxonomicName box="[868,1089,1661,1683]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis bold="true" box="[868,1089,1661,1683]" italics="true" pageId="6" pageNumber="40">A. changmaianus</emphasis>
</taxonomicName>
(TL
<quantity box="[1149,1297,1662,1684]" metricMagnitude="-3" metricUnit="m" metricValue="2.275" metricValueMax="2.6" metricValueMin="1.95" pageId="6" pageNumber="40" unit="mm" value="2.275" valueMax="2.6" valueMin="1.95">1.95–2.6 mm</quantity>
). Among the
<taxonomicName box="[816,980,1692,1714]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[816,980,1692,1714]" italics="true" pageId="6" pageNumber="40">A. minutulus</emphasis>
</taxonomicName>
species group,
<taxonomicName box="[1158,1314,1692,1714]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[1158,1314,1692,1714]" italics="true" pageId="6" pageNumber="40">A. seletarius</emphasis>
</taxonomicName>
is morpho- logically most similar to
<taxonomicName box="[1103,1269,1724,1746]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[1103,1269,1724,1746]" italics="true" pageId="6" pageNumber="40">A. minutulus</emphasis>
</taxonomicName>
. Excluding the latter, individual workers of
<taxonomicName box="[1150,1308,1755,1777]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[1150,1308,1755,1777]" italics="true" pageId="6" pageNumber="40">A. seletarius</emphasis>
</taxonomicName>
can be dis- tinguished from other species by the dentition on the masticatory margin of their mandibles, where the medium-sized subapical tooth is followed posteriorly by a distinct medium-sized denticle, and both the subapical tooth and the posterior denticle are of similar size (
<figureCitation box="[256,330,211,233]" captionStart="Figures 5 – 6" captionStartId="6.[570,639,1381,1398]" captionTargetBox="[570,1442,781,1363]" captionTargetId="figure@6.[569,1443,781,1364]" captionText="Figures 5 – 6. Aenictus seletarius (holotype, ANTWEB 1009000, LKCNHM) from Singapore. (5) Full-face view; (6) head view focusing on mandibles." httpUri="https://zenodo.org/record/269447/files/figure.png" pageId="7" pageNumber="41">
Fig.
<number box="[314,330,211,233]" pageId="7" pageNumber="41" value="6.0">6</number>
</figureCitation>
). This is contrary to the pattern of mandibular dentition of
<taxonomicName box="[433,657,241,263]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis bold="true" box="[433,657,241,263]" italics="true" pageId="7" pageNumber="41">A. changmaianus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minimus">
<emphasis bold="true" italics="true" pageId="7" pageNumber="41">A. minimus</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[221,245,273,294]" italics="true" pageId="7" pageNumber="41">A.</emphasis>
sp.
<number box="[294,320,273,295]" pageId="7" pageNumber="41" value="56.0">56</number>
of WJT and
<taxonomicName box="[468,664,272,294]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis bold="true" box="[468,664,272,294]" italics="true" pageId="7" pageNumber="41">A. subterraneus</emphasis>
</taxonomicName>
, where a medium-sized subapical tooth is followed posteriorly by a distinctly smaller denticle, as well as
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis bold="true" italics="true" pageId="7" pageNumber="41">A. peguensis</emphasis>
</taxonomicName>
, where both the subapical tooth and subsequent denticles are small in size. However, important variation in mandibular patterns can be observed in ants as blunting of the denticles with usage what may result in slight variation in dentition patterns (as observed in some
<typeStatus box="[214,329,520,541]" pageId="7" pageNumber="41">paratypes</typeStatus>
and non-type specimens). Therefore, relying on mandibular dentition alone for species determination is not ideal. In consideration of the above,
<taxonomicName box="[222,377,612,634]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[222,377,612,634]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
may be further distinguished from the
<taxonomicName box="[186,350,643,665]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[186,350,643,665]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
group species including the morphologically similar
<taxonomicName box="[361,542,674,696]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[361,542,674,696]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
by several other notable characters outlined below.
</paragraph>
<paragraph blockId="7.[143,771,180,1902]" pageId="7" pageNumber="41">
In full face view (
<figureCitation box="[379,446,736,758]" captionStart="Figures 5 – 6" captionStartId="6.[570,639,1381,1398]" captionTargetBox="[570,1442,781,1363]" captionTargetId="figure@6.[569,1443,781,1364]" captionText="Figures 5 – 6. Aenictus seletarius (holotype, ANTWEB 1009000, LKCNHM) from Singapore. (5) Full-face view; (6) head view focusing on mandibles." httpUri="https://zenodo.org/record/269447/files/figure.png" pageId="7" pageNumber="41">
Fig.
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</figureCitation>
),
<taxonomicName box="[468,625,736,758]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[468,625,736,758]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
displays the most square-shaped head among all
<taxonomicName box="[599,771,767,789]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[599,771,767,789]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
group species, as its head is almost as wide as it is long, CI
<number box="[237,306,829,851]" pageId="7" pageNumber="41" value="93.5" valueMax="96.0" valueMin="91.0">91–96</number>
; the side margins of its head are broadly convex and its posterior occipital margin is approximately
<number box="[228,237,888,902]" pageId="7" pageNumber="41" value="3.0">3</number>
/
<number box="[244,253,903,917]" pageId="7" pageNumber="41" value="4.0">4</number>
the length of its HW. Although
<taxonomicName box="[614,771,891,912]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis bold="true" box="[614,771,891,912]" italics="true" pageId="7" pageNumber="41">A. peguensis</emphasis>
</taxonomicName>
also possesses a head that is almost as wide as it is long, CI
<number box="[236,304,953,975]" pageId="7" pageNumber="41" value="89.0" valueMax="96.0" valueMin="82.0">82–96</number>
, the head shape of this species as well as that of
<taxonomicName box="[238,392,983,1004]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minimus">
<emphasis bold="true" box="[238,392,983,1004]" italics="true" pageId="7" pageNumber="41">A. minimus</emphasis>
</taxonomicName>
and
<emphasis bold="true" box="[463,486,983,1004]" italics="true" pageId="7" pageNumber="41">A.</emphasis>
sp.
<number box="[548,574,983,1005]" pageId="7" pageNumber="41" value="56.0">56</number>
of WJT are all markedly rounded; the side margins of their heads are strongly convex and the respective lengths of their posterior occipital margins are no more than
<number box="[692,701,1073,1087]" pageId="7" pageNumber="41" value="2.0">2</number>
/
<number box="[709,718,1089,1103]" pageId="7" pageNumber="41" value="3.0">3</number>
the lengths of their HW. In relation to
<taxonomicName box="[558,718,1107,1129]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[558,718,1107,1129]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
, the heads of the remaining
<taxonomicName box="[429,598,1138,1160]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[429,598,1138,1160]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
group species are comparatively longer than wide (
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis bold="true" italics="true" pageId="7" pageNumber="41">A. changmaianus</emphasis>
</taxonomicName>
, CI
<number box="[261,330,1200,1222]" pageId="7" pageNumber="41" value="82.0" valueMax="89.0" valueMin="75.0">75–89</number>
;
<taxonomicName box="[341,505,1199,1221]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[341,505,1199,1221]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
, CI
<number box="[550,616,1200,1222]" pageId="7" pageNumber="41" value="83.0" valueMax="90.0" valueMin="76.0">76–90</number>
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis bold="true" italics="true" pageId="7" pageNumber="41">A. subterraneus</emphasis>
</taxonomicName>
, CI
<number box="[252,320,1231,1253]" pageId="7" pageNumber="41" value="86.5" valueMax="87.0" valueMin="86.0">86–87</number>
) and their heads appear slightly more elongate than that of
<emphasis bold="true" box="[422,594,1261,1283]" italics="true" pageId="7" pageNumber="41">
<taxonomicName box="[422,589,1261,1283]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">A. seletarius</taxonomicName>
.
</emphasis>
The antennal scape of
<taxonomicName box="[248,406,1292,1314]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[248,406,1292,1314]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
is also relatively short in comparison to its head width, SI
<number box="[474,543,1323,1345]" pageId="7" pageNumber="41" value="62.5" valueMax="64.0" valueMin="61.0">61–64</number>
, in contrast to most other species in the
<taxonomicName box="[412,586,1354,1376]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[412,586,1354,1376]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
species group (
<taxonomicName box="[153,371,1385,1407]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis bold="true" box="[153,371,1385,1407]" italics="true" pageId="7" pageNumber="41">A. changmaianus</emphasis>
</taxonomicName>
, SI
<number box="[414,482,1385,1407]" pageId="7" pageNumber="41" value="70.0" valueMax="71.0" valueMin="69.0">69–71</number>
;
<taxonomicName box="[492,656,1385,1407]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis bold="true" box="[492,656,1385,1407]" italics="true" pageId="7" pageNumber="41">A. minutulus</emphasis>
</taxonomicName>
, SI
<number box="[698,767,1385,1407]" pageId="7" pageNumber="41" value="67.5" valueMax="68.0" valueMin="67.0">67–68</number>
;
<taxonomicName box="[143,304,1416,1437]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis bold="true" box="[143,304,1416,1437]" italics="true" pageId="7" pageNumber="41">A. peguensis</emphasis>
</taxonomicName>
, SI
<number box="[358,427,1416,1437]" pageId="7" pageNumber="41" value="76.5" valueMax="79.0" valueMin="74.0">74–79</number>
;
<taxonomicName box="[442,643,1416,1438]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis bold="true" box="[442,643,1416,1438]" italics="true" pageId="7" pageNumber="41">A. subterraneus</emphasis>
</taxonomicName>
, SI
<number box="[698,766,1416,1438]" pageId="7" pageNumber="41" value="77.0" valueMax="79.0" valueMin="75.0">75–79</number>
;
<emphasis bold="true" box="[143,167,1447,1468]" italics="true" pageId="7" pageNumber="41">A.</emphasis>
sp.
<number box="[228,254,1447,1469]" pageId="7" pageNumber="41" value="56.0">56</number>
of WJT, SI
<number box="[413,483,1447,1469]" pageId="7" pageNumber="41" value="70.5" valueMax="74.0" valueMin="67.0">67–74</number>
), but similar to that of
<taxonomicName box="[143,290,1478,1499]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minimus">
<emphasis bold="true" box="[143,290,1478,1499]" italics="true" pageId="7" pageNumber="41">A. minimus</emphasis>
</taxonomicName>
(SI
<number box="[338,408,1478,1500]" pageId="7" pageNumber="41" value="63.5" valueMax="64.0" valueMin="63.0">63–64</number>
).
</paragraph>
<paragraph blockId="7.[143,771,180,1902]" lastBlockId="7.[816,1444,180,295]" pageId="7" pageNumber="41">
In profile view (
<figureCitation box="[371,527,1509,1531]" captionStart="Figures 2 – 4" captionStartId="5.[143,211,1859,1876]" captionTargetBox="[347,1225,199,1814]" captionTargetId="figure@5.[346,1221,794,1300]" captionTargetPageId="5" captionText="Figures 2 – 4. Aenictus seletarius (holotype, ANTWEB 1009000, LKCNHM) from Singapore. (2) Profile view; (3) dorsal view; (4) profile view focusing" httpUri="https://zenodo.org/record/269446/files/figure.png" pageId="7" pageNumber="41">
Figs
<number box="[433,446,1509,1530]" pageId="7" pageNumber="41" value="2.0">2</number>
and
<number box="[511,527,1509,1531]" pageId="7" pageNumber="41" value="4.0">4</number>
</figureCitation>
), a strongly angular posterodorsal corner of the propodeum, a concave propodeal declivity, and a flat anterior face of the petiole distinguish
<taxonomicName box="[321,479,1601,1623]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[321,479,1601,1623]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
from
<taxonomicName box="[550,771,1601,1623]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis bold="true" box="[550,771,1601,1623]" italics="true" pageId="7" pageNumber="41">A. changmaianus</emphasis>
</taxonomicName>
and
<emphasis bold="true" box="[200,378,1632,1654]" italics="true" pageId="7" pageNumber="41">
<taxonomicName box="[200,371,1632,1654]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="minutulus">A. minutulus</taxonomicName>
,
</emphasis>
whose individuals have a more rounded posterodorsal corner of the propodeum, a weakly concave propodeal declivity, and a more rounded or broadly convex anterodorsal face of the petiole. In addition to the flat anterior face of the petiole in
<taxonomicName box="[254,409,1786,1808]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">
<emphasis bold="true" box="[254,409,1786,1808]" italics="true" pageId="7" pageNumber="41">A. seletarius</emphasis>
</taxonomicName>
, a less pronounced postpetiolar process also distinguishes this new species from
<taxonomicName box="[143,360,1848,1870]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis bold="true" box="[143,360,1848,1870]" italics="true" pageId="7" pageNumber="41">A. subterraneus</emphasis>
</taxonomicName>
, which has a rounded anterodorsal face and a longer and slightly more acute postpetiolar process. Another subtle difference between the two species is the helcium, which appears to be more elongate in
<taxonomicName box="[1127,1335,242,264]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis bold="true" box="[1127,1335,242,264]" italics="true" pageId="7" pageNumber="41">A. subterraneus</emphasis>
</taxonomicName>
than in
<emphasis bold="true" box="[816,980,273,295]" italics="true" pageId="7" pageNumber="41">
<taxonomicName box="[816,975,273,295]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="seletarius">A. seletarius</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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