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<document id="7AF873036926CEC6DFE09650FF52F1A2" ID-CLB-Dataset="31350" ID-DOI="10.11646/zootaxa.4353.2.12" ID-GBIF-Dataset="b621a553-5624-41a9-b439-0c542c3121e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1065156" ID-ZooBank="795A2001-4374-44F5-B731-28DDA7C5C062" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511423739668" checkinUser="plazi" docAuthor="Gyulai, Péter, Saldaitis, Aidas &amp; Vaitonis, Gintas" docDate="2017" docId="03AA8794FF96FF8B7DCEC56EFE7DFF7F" docLanguage="en" docName="zootaxa.4353.2.12.pdf" docOrigin="Zootaxa 4353 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gracilisinensis Gyulai &amp; Saldaitis &amp; Vaitonis 2017, gen. nov." docType="treatment" docUuid="8A0A09E2-5722-4F7D-A3AC-D6B2F324983B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="395" masterDocId="FF93FFECFF96FF897D59C313FFB3FF8B" masterDocTitle="A new genus of Noctuidae (Lepidoptera) from China with the description of two new species" masterLastPageNumber="398" masterPageNumber="393" pageNumber="393" updateTime="1698504174655" updateUser="ExternalLinkService">
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<mods:title id="89792DE27469CCF7EFF7645056D27F48">A new genus of Noctuidae (Lepidoptera) from China with the description of two new species</mods:title>
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<mods:namePart id="74A1112866427886037BBE5BB227BC35">Gyulai, Péter</mods:namePart>
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<mods:namePart id="9398AFB4CD75FBA2238610724247B6B9">Saldaitis, Aidas</mods:namePart>
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<mods:namePart id="B0B363074A9CD38F868D352D3156CFC9">Vaitonis, Gintas</mods:namePart>
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<mods:title id="2631CC721BC76EE83C47EC95C43E8161">Zootaxa</mods:title>
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<mods:part id="A4C145D5006183C957ABBA1D51B3EEAB">
<mods:date id="0BF69B728F1293895872EC85CE22B44C">2017</mods:date>
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<mods:number id="A4A28C35A1FD9E8FB0B2A36A7DC4AC97">2017-11-23</mods:number>
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<mods:number id="8EF4319D7676BA85BD345B23F47B2279">4353</mods:number>
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<mods:classification id="BFB9E42F843FB6720046CC670AAE631C">journal article</mods:classification>
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<mods:identifier id="AB17E9089B9CFC69F5282BDEBE2E8687" type="DOI">10.11646/zootaxa.4353.2.12</mods:identifier>
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<treatment id="03AA8794FF96FF8B7DCEC56EFE7DFF7F" ID-DOI="http://doi.org/10.5281/zenodo.6020939" ID-GBIF-Taxon="137541595" ID-Zenodo-Dep="6020939" LSID="urn:lsid:zoobank.org:act:8A0A09E2-5722-4F7D-A3AC-D6B2F324983B" httpUri="http://treatment.plazi.org/id/03AA8794FF96FF8B7DCEC56EFE7DFF7F" lastPageId="2" lastPageNumber="395" pageId="0" pageNumber="393">
<subSubSection id="C3196509FF96FF897DCEC56EFED1F93C" pageId="0" pageNumber="393" type="nomenclature">
<paragraph id="8BBC3682FF96FF897DCEC56EFD04F913" blockId="0.[151,1436,1627,2031]" box="[151,695,1661,1688]" pageId="0" pageNumber="393">
<heading id="D0F481EEFF96FF897DCEC56EFD04F913" bold="true" box="[151,695,1661,1688]" fontSize="11" level="1" pageId="0" pageNumber="393" reason="1">
<emphasis id="B977EA90FF96FF897DCEC56EFD04F913" bold="true" box="[151,695,1661,1688]" pageId="0" pageNumber="393">
<taxonomicName id="4C034D01FF96FF897DCEC56EFD88F913" authority="Gyulai &amp; Saldaitis" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[151,571,1661,1688]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF96FF897DCEC56EFEFFF91C" bold="true" box="[151,332,1661,1687]" italics="true" pageId="0" pageNumber="393">Gracilisinensis</emphasis>
Gyulai &amp; Saldaitis
</taxonomicName>
,
<taxonomicNameLabel id="A24457EBFF96FF897F10C56DFD04F913" box="[585,695,1662,1688]" pageId="0" pageNumber="393" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC3682FF96FF897DCEC5B3FED1F93C" blockId="0.[151,1436,1627,2031]" box="[151,354,1696,1719]" pageId="0" pageNumber="393">
(
<figureCitation id="13382A07FF96FF897DC7C5B3FF4CF93C" box="[158,255,1696,1719]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="0" pageNumber="393">Figs 1–4</figureCitation>
,
<figureCitation id="13382A07FF96FF897C53C5B3FE87F93C" box="[266,308,1696,1719]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="0" pageNumber="393">7–9</figureCitation>
,
<figureCitation id="13382A07FF96FF897C66C5B3FEE8F93C" box="[319,347,1696,1719]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="0" pageNumber="393">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3196509FF96FF897DCEC5D0FCCBF951" box="[151,888,1730,1754]" pageId="0" pageNumber="393" type="materials_examined">
<paragraph id="8BBC3682FF96FF897DCEC5D0FCCBF951" blockId="0.[151,1436,1627,2031]" box="[151,888,1730,1754]" pageId="0" pageNumber="393">
<emphasis id="B977EA90FF96FF897DCEC5D0FE9EF951" bold="true" box="[151,301,1731,1754]" pageId="0" pageNumber="393">Type species:</emphasis>
<taxonomicName id="4C034D01FF96FF897C6AC5D1FDA6F951" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[307,533,1730,1754]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="vaiva" status="sp. nov.">
<emphasis id="B977EA90FF96FF897C6AC5D1FDA6F951" box="[307,533,1730,1754]" italics="true" pageId="0" pageNumber="393">Gracilisinensis vaiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457EBFF96FF897F45C5D0FDC3F951" box="[540,624,1731,1754]" pageId="0" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C3196509FF96FF887D9EC5F6FA2FFA8F" lastPageId="1" lastPageNumber="394" pageId="0" pageNumber="393" type="diagnosis">
<paragraph id="8BBC3682FF96FF897D9EC5F6FD2FF864" blockId="0.[151,1436,1627,2031]" pageId="0" pageNumber="393">
<emphasis id="B977EA90FF96FF897D9EC5F6FE8FF977" bold="true" box="[199,316,1765,1788]" pageId="0" pageNumber="393">Diagnosis.</emphasis>
The new genus belongs to the subfamily
<taxonomicName id="4C034D01FF96FF897E5DC5F6FC36F977" authorityName="Grote" authorityYear="1896" box="[772,901,1765,1788]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="subFamily" subFamily="Psaphidinae">Psaphidinae</taxonomicName>
Grote and tribe
<taxonomicName id="4C034D01FF96FF897963C5F6FB5CF977" authority="Grote" authorityName="Grote" box="[1082,1263,1765,1788]" class="Insecta" family="Noctuidae" genus="Psaphidini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">Psaphidini Grote</taxonomicName>
based on a few shared characters, which are as follows: bipectinated antennae of males; in male genitalia the absence of the typical harpe, instead of it bearing a small ventrally positioned saccular appendage, which is originated from subventral part of the valva (shared feature with
<taxonomicName id="4C034D01FF96FF897C81C45EFD81F8EF" authorityName="Grote" authorityYear="1882" box="[472,562,1869,1892]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="subFamily" subFamily="Stiriinae">Stiriinae</taxonomicName>
and
<taxonomicName id="4C034D01FF96FF897F3DC45EFCAEF8EF" authorityName="Forbes &amp; Franclemont" authorityYear="1954" box="[612,797,1869,1892]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="subFamily" subFamily="Oncocnemidinae">Oncocnemidinae</taxonomicName>
), appearing independent from the distal part of the sacculus; a fine, rather anteriorly projecting costal extension in the distal part of the ventral edge of the valva and ample, membranous, somewhat dorsally everted vesica with a subbasal row of short but strong cornuti. In female genitalia the shared characters are the membranous but distally sclerotized ductus bursae, hardly prominent appendix bursae, and long, membranous, corpus bursae without signa.
</paragraph>
<paragraph id="8BBC3682FF97FF887D9EC38BFE21FBF2" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
The most closely related genus is
<taxonomicName id="4C034D01FF97FF887F6DC384FC9DFF24" authority="Warren" authorityName="Warren" box="[564,814,151,175]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887F6DC384FD6AFF24" box="[564,729,151,175]" italics="true" pageId="1" pageNumber="394">Diphtherocome</emphasis>
Warren
</taxonomicName>
(
<figureCitation id="13382A07FF97FF887E64C38BFC2EFF24" box="[829,925,152,175]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="1" pageNumber="394">Figs 5, 6</figureCitation>
), by some external characters (colour and wing pattern), however mostly by the similar genitalia morphological structure: in males the base configuration of the saccular and ventral extensions and the subbasal row of short but strong cornuti; in females the configuration of antrum – ductus bursae complex, the shallow appendix bursae, the long, saccular corpus bursae and the absence of the signae. The two species of
<taxonomicName id="4C034D01FF97FF887C57C232FE1CFEB2" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[270,431,289,313]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF97FF887C57C232FE1CFEB2" box="[270,431,289,313]" italics="true" pageId="1" pageNumber="394">Gracilisinensis</emphasis>
</taxonomicName>
(
<figureCitation id="13382A07FF97FF887C9BC231FD97FEB2" box="[450,548,290,313]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="1" pageNumber="394">Figs 1–4</figureCitation>
) differ from the species of
<taxonomicName id="4C034D01FF97FF887E05C232FBB2FEB2" box="[860,1025,289,313]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887E05C232FBB2FEB2" box="[860,1025,289,313]" italics="true" pageId="1" pageNumber="394">Diphtherocome</emphasis>
</taxonomicName>
by smaller size, much more broadly bipectinated antennae with longer rami in males and the more prominent orbicular and reniform stigmata of forewing. In the male genitalia, the shape of the valvae of the
<typeStatus id="54B88820FF97FF887F90C27BFD44FEF4" box="[713,759,360,383]" pageId="1" pageNumber="394">type</typeStatus>
species of the new genus (
<figureCitation id="13382A07FF97FF887975C27BFB3CFEF4" box="[1068,1167,360,383]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">Figs 7–9</figureCitation>
) resembles those of the
<taxonomicName id="4C034D01FF97FF887DCEC29AFE91FE2A" box="[151,290,393,417]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="discibrunnea">
<emphasis id="B977EA90FF97FF887DCEC29AFE91FE2A" box="[151,290,393,417]" italics="true" pageId="1" pageNumber="394">discibrunnea</emphasis>
</taxonomicName>
species group of
<taxonomicName id="4C034D01FF97FF887C85C29AFD32FE2A" box="[476,641,393,417]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887C85C29AFD32FE2A" box="[476,641,393,417]" italics="true" pageId="1" pageNumber="394">Diphtherocome</emphasis>
</taxonomicName>
(
<taxonomicName id="4C034D01FF97FF887FD6C29AFB97FE2A" authority="(Moore)" authorityName="Moore" box="[655,1060,393,417]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="discibrunnea">
<emphasis id="B977EA90FF97FF887FD6C29AFC76FE2A" box="[655,965,393,417]" italics="true" pageId="1" pageNumber="394">Diphtherocome discibrunnea</emphasis>
(Moore)
</taxonomicName>
,
<taxonomicName id="4C034D01FF97FF887968C29AFA24FE2A" authority="Wileman" authorityName="Wileman" box="[1073,1431,393,417]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B977EA90FF97FF887968C29AFA9DFE2A" box="[1073,1326,393,417]" italics="true" pageId="1" pageNumber="394">Diphtherocome pulchra</emphasis>
Wileman
</taxonomicName>
,
<taxonomicName id="4C034D01FF97FF887DCEC2BFFDA1FE4F" authority="(Leech)" authorityName="Leech" box="[151,530,428,452]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="marmorea">
<emphasis id="B977EA90FF97FF887DCEC2BFFE06FE4F" box="[151,437,428,452]" italics="true" pageId="1" pageNumber="394">Diphtherocome marmorea</emphasis>
(Leech)
</taxonomicName>
(
<figureCitation id="13382A07FF97FF887F7FC2BEFDC4FE4F" box="[550,631,429,452]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="1" pageNumber="394">Fig. 14</figureCitation>
) and
<taxonomicName id="4C034D01FF97FF887FE5C2BFFB53FE4F" authority="Gyulai, Ronkay &amp; Ronkay" authorityName="Gyulai, Ronkay &amp; Ronkay" box="[700,1248,428,452]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="ligula">
<emphasis id="B977EA90FF97FF887FE5C2BFFC19FE4F" box="[700,938,428,452]" italics="true" pageId="1" pageNumber="394">Diphtherocome ligula</emphasis>
Gyulai, Ronkay &amp; Ronkay
</taxonomicName>
(
<figureCitation id="13382A07FF97FF8879AAC2BEFA8FFE4F" box="[1267,1340,429,452]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="1" pageNumber="394">Figs 5</figureCitation>
,
<figureCitation id="13382A07FF97FF887815C2BEFADAFE4F" box="[1356,1385,429,452]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">10</figureCitation>
,
<figureCitation id="13382A07FF97FF887821C2BEFA20FE4F" box="[1400,1427,429,452]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="1" pageNumber="394">13</figureCitation>
) (
<bibRefCitation id="EF924B73FF97FF887DC7C2C3FE6DFE6C" author="Hreblay" box="[158,478,464,487]" pageId="1" pageNumber="394" refString="Hreblay, M. &amp; Kononenko, V. S. (1997) The Chinese species of the Diphterocome pallida and D. discibrunnea species-groups (Lepidoptera, Noctuidae, Acronictinae) in the collection of the Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Esperiana, 7, 669 - 686." type="journal article" year="1997">Hreblay &amp; Kononenko 1997</bibRefCitation>
;
<bibRefCitation id="EF924B73FF97FF887CB4C2C3FCE8FE6C" author="Gyulai" box="[493,859,464,487]" pageId="1" pageNumber="394" refString="Gyulai, P., Ronkay, L. &amp; Ronkay, G. (2014) New species of Noctuidae (Lepidoptera) from Asia. Esperiana, 19, 187 - 208." type="journal article" year="2014">Gyulai, Ronkay &amp; Ronkay 2014</bibRefCitation>
). However, the members of the new genus lack the typical harpe but bear very long and fine hairs in high density particularly in the dorsal, setose side; a slight saccular appendage and a small digitate process (which sometimes leaning back inward) in the distal part of the ventral edge. The “saccular appendage” is a small, ventrally positioned harpe/clasper, which appears as mostly independent from the distal part of the sacculus; and the digitate process is a fine, rather anteriorly projecting costal extension, which may be curved backwards to the ventral surface of the valva. The vesica with the strong cornuti field is a shared feature within some other species groups of
<taxonomicName id="4C034D01FF97FF887CCDC18CFD8AFD3C" box="[404,569,671,695]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887CCDC18CFD8AFD3C" box="[404,569,671,695]" italics="true" pageId="1" pageNumber="394">Diphtherocome</emphasis>
</taxonomicName>
(e.g. the
<taxonomicName id="4C034D01FF97FF887FC7C18CFCFAFD3C" authority="(Moore)" authorityName="Moore" box="[670,841,671,695]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B977EA90FF97FF887FC7C18CFD59FD3C" box="[670,746,671,695]" italics="true" pageId="1" pageNumber="394">pallida</emphasis>
(Moore)
</taxonomicName>
species group (
<figureCitation id="13382A07FF97FF887EA9C1B3FB8FFD3C" box="[1008,1084,672,695]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">Fig. 11</figureCitation>
) (
<bibRefCitation id="EF924B73FF97FF88790AC1B3FA23FD3C" author="Hreblay" box="[1107,1424,672,695]" pageId="1" pageNumber="394" refString="Hreblay, M. &amp; Kononenko, V. S. (1997) The Chinese species of the Diphterocome pallida and D. discibrunnea species-groups (Lepidoptera, Noctuidae, Acronictinae) in the collection of the Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Esperiana, 7, 669 - 686." type="journal article" year="1997">Hreblay &amp; Kononenko, 1997</bibRefCitation>
), while the structure of the female genitalia (
<figureCitation id="13382A07FF97FF887F35C1D1FD09FD52" box="[620,698,706,729]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="1" pageNumber="394">Fig. 12</figureCitation>
) appears closer to that of the
<taxonomicName id="4C034D01FF97FF887958C1D2FB3FFD52" box="[1025,1164,705,729]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="discibrunnea">
<emphasis id="B977EA90FF97FF887958C1D2FB3FFD52" box="[1025,1164,705,729]" italics="true" pageId="1" pageNumber="394">discibrunnea</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="4C034D01FF97FF887995C1D1FA98FD52" box="[1228,1323,705,729]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="ligula">
<emphasis id="B977EA90FF97FF887995C1D1FA98FD52" box="[1228,1323,705,729]" italics="true" pageId="1" pageNumber="394">D. ligula</emphasis>
</taxonomicName>
(
<figureCitation id="13382A07FF97FF887865C1D1FA3AFD52" box="[1340,1417,706,729]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="1" pageNumber="394">Fig. 13</figureCitation>
)) species group. The
<typeStatus id="54B88820FF97FF887C32C1F6FE2AFD77" box="[363,409,741,764]" pageId="1" pageNumber="394">type</typeStatus>
species of
<taxonomicName id="4C034D01FF97FF887F4CC1F7FD09FD77" box="[533,698,740,764]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887F4CC1F7FD09FD77" box="[533,698,740,764]" italics="true" pageId="1" pageNumber="394">Diphtherocome</emphasis>
</taxonomicName>
is
<taxonomicName id="4C034D01FF97FF887F85C1F6FCFAFD77" box="[732,841,740,764]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B977EA90FF97FF887F85C1F6FCFAFD77" box="[732,841,740,764]" italics="true" pageId="1" pageNumber="394">D. pallida</emphasis>
</taxonomicName>
(
<figureCitation id="13382A07FF97FF887E03C1F6FC13FD77" box="[858,928,741,764]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="1" pageNumber="394">Figs 6</figureCitation>
,
<figureCitation id="13382A07FF97FF887EF4C1F6FC7BFD77" box="[941,968,741,764]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">11</figureCitation>
), in which the vesica with the large, partly interrupted, comb- or brush-like surface structure of the carina penis seems to be the most resembling those of the two new species of the
<emphasis id="B977EA90FF97FF887C20C03AFD91FCCA" box="[377,546,809,833]" italics="true" pageId="1" pageNumber="394">
<taxonomicName id="4C034D01FF97FF887C20C03AFDADFCCA" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[377,542,809,833]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">Gracilisinensis</taxonomicName>
.
</emphasis>
The female genitalia of
<taxonomicName id="4C034D01FF97FF887E1AC039FC06FCCA" box="[835,949,809,833]" class="Insecta" family="Noctuidae" genus="Diphtherocome" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B977EA90FF97FF887E1AC039FC06FCCA" box="[835,949,809,833]" italics="true" pageId="1" pageNumber="394">D. pallida</emphasis>
</taxonomicName>
is considerably distinctive by the strongly sclerotized antrum, ductus bursae and folded-ribbed, broadly rounded appendix bursae and the large pouch-like corpus bursae. On the other hand, it is worth to note that no such genitalia character complex set is known to occur in the Eurasiatic fauna. At first sight, the two species of
<taxonomicName id="4C034D01FF97FF887FE2C082FCEEFC22" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[699,861,913,937]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF97FF887FE2C082FCEEFC22" box="[699,861,913,937]" italics="true" pageId="1" pageNumber="394">Gracilisinensis</emphasis>
</taxonomicName>
are also similar to some species of the genus
<taxonomicName id="4C034D01FF97FF88780CC081FEB9FC47" authority="Staudinger" authorityName="Staudinger" class="Insecta" family="Noctuidae" genus="Victrix" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF88780CC081FA28FC22" box="[1365,1435,914,937]" italics="true" pageId="1" pageNumber="394">Victrix</emphasis>
Staudinger
</taxonomicName>
of the subfamily
<taxonomicName id="4C034D01FF97FF887C8DC0A6FDECFC47" authorityName="Guenée" authorityYear="1852" box="[468,607,949,972]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="subFamily" subFamily="Bryophilinae">Bryophilinae</taxonomicName>
however, these species are generally larger. The largest species of
<taxonomicName id="4C034D01FF97FF887816C0A6FA26FC47" box="[1359,1429,949,972]" class="Insecta" family="Noctuidae" genus="Victrix" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887816C0A6FA26FC47" box="[1359,1429,949,972]" italics="true" pageId="1" pageNumber="394">Victrix</emphasis>
</taxonomicName>
, including the type-species, which is the largest taxon of the genus, may exceed the wingspan
<quantity id="4CFB9B67FF97FF8879DAC0CBFB7CFC64" box="[1155,1231,984,1007]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="394" unit="mm" value="37.0">37 mm</quantity>
which is the upper limit of the size of
<taxonomicName id="4C034D01FF97FF887C31C0EAFDBAFB9A" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[360,521,1017,1041]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF97FF887C31C0EAFDBAFB9A" box="[360,521,1017,1041]" italics="true" pageId="1" pageNumber="394">Gracilisinensis</emphasis>
</taxonomicName>
. The size of specimens together with broader wings and conspicuously bipectinated antennae of the males are characteristic and diagnose the genus
<emphasis id="B977EA90FF97FF887E02C70FFBB7FBBF" box="[859,1028,1052,1076]" italics="true" pageId="1" pageNumber="394">
<taxonomicName id="4C034D01FF97FF887E02C70FFBB3FBBF" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[859,1024,1052,1076]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">Gracilisinensis</taxonomicName>
.
</emphasis>
The features of the male and female genitalia support the assignment of the latter two genera to two different subfamilies:
<taxonomicName id="4C034D01FF97FF887973C72CFB78FBDC" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[1066,1227,1087,1111]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF97FF887973C72CFB78FBDC" box="[1066,1227,1087,1111]" italics="true" pageId="1" pageNumber="394">Gracilisinensis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C034D01FF97FF8879B7C753FADCFBDC" authorityName="Grote" authorityYear="1896" box="[1262,1391,1088,1111]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="subFamily" subFamily="Psaphidinae">Psaphidinae</taxonomicName>
and
<taxonomicName id="4C034D01FF97FF887DCEC771FF6FFBF2" box="[151,220,1122,1145]" class="Insecta" family="Noctuidae" genus="Victrix" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B977EA90FF97FF887DCEC771FF6FFBF2" box="[151,220,1122,1145]" italics="true" pageId="1" pageNumber="394">Victrix</emphasis>
</taxonomicName>
to
<taxonomicName id="4C034D01FF97FF887DA6C771FE3DFBF2" authorityName="Guenée" authorityYear="1852" box="[255,398,1122,1145]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="subFamily" subFamily="Bryophilinae">Bryophilinae</taxonomicName>
.
</paragraph>
<paragraph id="8BBC3682FF97FF887D9EC796FA2FFA8F" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
Finally, it is worth to mention, that although the external features of
<taxonomicName id="4C034D01FF97FF887EE4C797FBEDFB17" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[957,1118,1156,1180]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B977EA90FF97FF887EE4C797FBEDFB17" box="[957,1118,1156,1180]" italics="true" pageId="1" pageNumber="394">Gracilisinensis</emphasis>
</taxonomicName>
can differ significantly from some colourful genera of the subfamily
<taxonomicName id="4C034D01FF97FF887F66C7BBFD28FB34" authorityName="Grote" authorityYear="1882" box="[575,667,1192,1215]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="subFamily" subFamily="Stiriinae">Stiriinae</taxonomicName>
, confined only to North and Central America (
<bibRefCitation id="EF924B73FF97FF8879D0C7BBFAB4FB34" author="Poole" box="[1161,1287,1192,1215]" pageId="1" pageNumber="394" refString="Poole, R. W. (1995) Noctuoidea, Noctuidae (part). In: Dominick, R. B. (Ed.), The Moths of America North of Mexico. 26.1. The Wedge Entomological Research Foundation, Washington, D. C., 249 pp." type="book" year="1995">
<collectingRegion id="49C7F860FF97FF8879D0C7BBFB74FB34" box="[1161,1223,1192,1215]" country="United Kingdom" name="Poole" pageId="1" pageNumber="394">Poole</collectingRegion>
1995
</bibRefCitation>
), the genitalia structure and particularly those of the males display remarkable resemblance. For instance, the shape of the valvae of
<taxonomicName id="4C034D01FF97FF887DCEC7FFFE11FA8F" baseAuthorityName="H. Edwards" baseAuthorityYear="1882" box="[151,418,1260,1284]" class="Insecta" family="Noctuidae" genus="Plagiomimicus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="aureolum">
<emphasis id="B977EA90FF97FF887DCEC7FFFE11FA8F" box="[151,418,1260,1284]" italics="true" pageId="1" pageNumber="394">Plagiomimicus aureolum</emphasis>
</taxonomicName>
(in:
<bibRefCitation id="EF924B73FF97FF887C8DC7FEFDE3FA8F" author="Poole" box="[468,592,1261,1284]" pageId="1" pageNumber="394" refString="Poole, R. W. (1995) Noctuoidea, Noctuidae (part). In: Dominick, R. B. (Ed.), The Moths of America North of Mexico. 26.1. The Wedge Entomological Research Foundation, Washington, D. C., 249 pp." type="book" year="1995">
<collectingRegion id="49C7F860FF97FF887C8DC7FEFDA3FA8F" box="[468,528,1261,1284]" country="United Kingdom" name="Poole" pageId="1" pageNumber="394">Poole</collectingRegion>
1995
</bibRefCitation>
: plate J,
<figureCitation id="13382A07FF97FF887FF2C7FEFC92FA8F" box="[683,801,1261,1284]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">Figs 10, 11</figureCitation>
) almost overlaps with that of
<taxonomicName id="4C034D01FF97FF887907C7FFFAF2FA8F" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[1118,1345,1260,1284]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="moha" status="sp. nov.">
<emphasis id="B977EA90FF97FF887907C7FFFAF2FA8F" box="[1118,1345,1260,1284]" italics="true" pageId="1" pageNumber="394">Gracilisinensis moha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457EBFF97FF88781EC7FEFA2FFA8F" box="[1351,1436,1261,1284]" pageId="1" pageNumber="394" rank="species">
<emphasis id="B977EA90FF97FF88781EC7FEFA26FA8F" bold="true" box="[1351,1429,1261,1284]" pageId="1" pageNumber="394">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3196509FF97FF887D9EC603FAFFF857" pageId="1" pageNumber="394" type="description">
<paragraph id="8BBC3682FF97FF887D9EC603FA25F992" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
<emphasis id="B977EA90FF97FF887D9EC603FEE1FAAC" bold="true" box="[199,338,1296,1319]" pageId="1" pageNumber="394">Description.</emphasis>
Medium-sized species, wingspan
<quantity id="4CFB9B67FF97FF887F99C603FC86FAAC" box="[704,821,1296,1319]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.7" metricValueMin="2.9" pageId="1" pageNumber="394" unit="mm" value="33.0" valueMax="37.0" valueMin="29.0">29–37 mm</quantity>
, with thin, gracile body (
<figureCitation id="13382A07FF97FF887919C603FB2DFAAC" box="[1088,1182,1296,1319]" captionStart="FIGURES 1–6" captionStartId="2.[151,258,1644,1666]" captionTargetBox="[152,1435,302,1608]" captionTargetId="figure@2.[151,1436,298,1620]" captionTargetPageId="2" captionText="FIGURES 1–6. Gracilisinensis &amp; Diphtherocome spp. adults. 1. G. vaiva gen. nov. &amp; sp. nov., male, HT, China, W. Sichuan, PGY4048m(coll.PGM);2.G.vaiva gen. nov. &amp; sp.n., female, PT, China,W.Sichuan,PGY4049f (coll.PGM); 3. G.vaiva gen. nov. &amp; sp. nov., male, PT,China, W.Sichuan (coll. AFM); 4. G.moha gen. nov. &amp; sp. nov., male, HT, China, Shaanxi, PGY4588m (coll. PGM); 5. D. ligula, male, HT, China, Sichuan, PGY3841m (coll. PGM); 6. D. pallida, male, China, W. Sichuan, PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065158/files/figure.png" pageId="1" pageNumber="394">Figs 1–4</figureCitation>
<emphasis id="B977EA90FF97FF8879C7C603FB14FAAC" bold="true" box="[1182,1191,1296,1319]" pageId="1" pageNumber="394">)</emphasis>
. Last palpomere white tipped, frons and collar green or greenish, but paler on the older ones. Male antennae bipectinated, pale ochre, those of the females filiform, brown and white segmented. Forewings with pointed apex, costa somewhat arched, ground colour dark green in the fresh specimens, paler or ochre in the older ones, variegated with pale violet and brown. Transversal lines double, antemedial line somewhat zigzag, postmedial line arched, subterminal line the most conspicuous, being white in the basal segment, bearing a more or less large blackish patch in the inner side, which is more conspicuous in the males. Orbicular and reniform stigmata green or greenish, prominent, while the claviform stigmata a slight irregular blackish patch. Hindwings whitish brown, with slight brown cellular spot, sinuous medial line and darker marginal area.
</paragraph>
<paragraph id="8BBC3682FF97FF887D9EC537FAA3F887" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
<emphasis id="B977EA90FF97FF887D9EC537FED4F9B7" box="[199,359,1572,1596]" italics="true" pageId="1" pageNumber="394">Male genitalia</emphasis>
(
<figureCitation id="13382A07FF97FF887C20C536FE6CF9B7" box="[377,479,1573,1596]" captionStart="FIGURES7–11" captionStartId="3.[151,258,1655,1677]" captionTargetBox="[154,1434,194,1629]" captionTargetId="figure@3.[151,1436,194,1634]" captionTargetPageId="3" captionText="FIGURES7–11. Gracilisinensis &amp; Diphtherocome spp.male genitalia.7. G.vaiva gen. nov. &amp;sp. nov.,HT,W.Sichuan,PGY4048m (coll.PGM);8. G.vaiva gen.nov. &amp;sp.nov., PT,W.Sichuan,PGY3033m(coll.ASV);9.G.moha gen.nov. &amp;sp. nov.,HT,Shaanxi, PGY4588m(coll. PGM);10.D.ligula, HT, China, Sichuan,PGY3841m (coll.PGM); 11. D. pallida, China,W.Sichuan,PGY4187m (coll. PGM)." httpUri="https://zenodo.org/record/1065160/files/figure.png" pageId="1" pageNumber="394">Figs 7–9</figureCitation>
). They can be characterized by the medium long, straight, slightly lancet uncus, with pointed thin tip apically; tegumen developed, elongated; broadly heart shaped juxta with slight medial depression, laterally the broadest, dorsally with two slight appendages; V-shaped vinculum; absence of the typical harpe and corona; distally somewhat slenderer, apically pointed valvae, bearing very long and fine hairs in high density particularly in the dorsal, setosed side, a small saccular appendage and a slight pollex-like process in the distal part of the ventral edge (which sometimes leaning back inwards); medially curved, sclerotized aedeagus with a sclerotized ventral carinal bar and ample, membranous, somewhat dorsally everted vesica with a subbasal row of short but strong cornuti.
</paragraph>
<paragraph id="8BBC3682FF97FF887D9EC404FAB6F81C" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
<emphasis id="B977EA90FF97FF887D9EC404FECEF8A4" box="[199,381,1815,1839]" italics="true" pageId="1" pageNumber="394">Female genitalia</emphasis>
(
<figureCitation id="13382A07FF97FF887CC8C40BFE50F8A4" box="[401,483,1816,1839]" captionStart="FIGURES 12–14" captionStartId="4.[151,258,1900,1922]" captionTargetBox="[163,1432,893,1868]" captionTargetId="figure@4.[151,1436,885,1880]" captionTargetPageId="4" captionText="FIGURES 12–14. Gracilisinensis &amp; Diphtherocome spp. female genitalia. 12. G. vaiva gen. nov. &amp; sp. nov., PT, W. Sichuan, PGY4049f (coll.PGM); 13. D.ligula, PT,China, Sichuan, PGY3857f (coll.PGM); 14. D.marmorea, China, W.Sichuan, PGY3071f (coll. PGM)." httpUri="https://zenodo.org/record/1065162/files/figure.png" pageId="1" pageNumber="394">Fig. 12</figureCitation>
). Ovipositor long, anterior and posterior apophyses fine, the last ones shorter; antrum somewhat flower shaped, sclerotized, broad, with deep medial incision distally; ductus bursae almost straight, membranous anteriorly and sclerotized distally, evenly broad, with longitudinal sclerotized folds in the wall; appendix bursae hardly prominent, flat on the posterior part of the very long, membranous, evenly slim corpus bursae, without signa.
</paragraph>
<paragraph id="8BBC3682FF97FF887D9EC4B1FAFFF857" blockId="1.[151,1436,151,2012]" pageId="1" pageNumber="394">
The two new species of the new genus are somewhat similar to each other, however they can be easily separated by remarkable external differences and in male genitalia characters. The female of the
<taxonomicName id="4C034D01FF97FF887948C4D6FBDAF857" authority="Gyulai &amp; Saldaitis &amp; Vaitonis, 2017" authorityName="Gyulai &amp; Saldaitis &amp; Vaitonis" authorityYear="2017" box="[1041,1129,1988,2012]" class="Insecta" family="Noctuidae" genus="Gracilisinensis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="moha" status="sp. nov.">
<emphasis id="B977EA90FF97FF887948C4D6FBDAF857" box="[1041,1129,1988,2012]" italics="true" pageId="1" pageNumber="394">G. moha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457EBFF97FF887929C4D6FB71F857" box="[1136,1218,1989,2012]" pageId="1" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3196509FF94FF8B7D9EC38BFD94FF24" box="[199,551,152,175]" pageId="2" pageNumber="395" type="distribution">
<paragraph id="8BBC3682FF94FF8B7D9EC38BFD94FF24" blockId="2.[151,1436,152,244]" box="[199,551,152,175]" pageId="2" pageNumber="395">
<emphasis id="B977EA90FF94FF8B7D9EC38BFEE1FF24" bold="true" box="[199,338,152,175]" pageId="2" pageNumber="395">Distribution</emphasis>
. Endemic to
<collectingCountry id="F3147612FF94FF8B7CB9C38BFD90FF24" box="[480,547,152,175]" name="China" pageId="2" pageNumber="395">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196509FF94FF8B7D9EC3A9FE7DFF7F" pageId="2" pageNumber="395" type="etymology">
<paragraph id="8BBC3682FF94FF8B7D9EC3A9FE7DFF7F" blockId="2.[151,1436,152,244]" pageId="2" pageNumber="395">
<emphasis id="B977EA90FF94FF8B7D9EC3A9FEF3FF5A" bold="true" box="[199,320,186,209]" pageId="2" pageNumber="395">Etymology</emphasis>
. “
<emphasis id="B977EA90FF94FF8B7C04C3AAFE07FF5A" box="[349,436,185,209]" italics="true" pageId="2" pageNumber="395">Gracilis</emphasis>
” in the Latin means thin, slender, graceful, elegant, referring to the thin body; “
<emphasis id="B977EA90FF94FF8B7865C3A9FA23FF5A" box="[1340,1424,186,209]" italics="true" pageId="2" pageNumber="395">sinensis</emphasis>
” means: Chinese, from
<collectingCountry id="F3147612FF94FF8B7CDEC3CEFE79FF7F" box="[391,458,221,244]" name="China" pageId="2" pageNumber="395">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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