treatments-xml/data/AD/04/98/AD0498777415FFA2FF1C2DE5FBA6FE90.xml

<document id="51662288DBF819D248C9FA1C56119193" ID="10.11646/zootaxa.4273.4.4SLASH11259" ID-CLB-Dataset="32890" ID-DOI="10.11646/zootaxa.4273.4.4" ID-GBIF-Dataset="61073b76-2847-4a9a-8b22-ae63c9ec8eba" ID-ISSN="1175-5326" ID-Zenodo-Dep="803754" ID-ZooBank="8CC36A32-5BCA-4BCE-AD6D-DB325C0F5D51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496822096739" checkinUser="plazi" docAuthor="Tutman, Pero, Freyhof, Jörg, Dulčić, Jakov, Glamuzina, Branko &amp; Geiger, Matthias" docDate="2017" docId="AD0498777415FFA2FF1C2DE5FBA6FE90" docLanguage="en" docName="zootaxa.4273.4.4SLASH11259.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lampetra soljani Tutman, Freyhof, Dulčić, Glamuzina &amp; Geiger, 2017, new species" docType="treatment" docVersion="7" lastPageNumber="540" masterDocId="513DE00F7410FFA8FF8B2D72FFFAFFD0" masterDocTitle="Lampetra soljani, a new brook lamprey from the southern Adriatic Sea basin (Petromyzontiformes: Petromyzontidae)" masterLastPageNumber="548" masterPageNumber="531" pageNumber="536" updateTime="1698440251818" updateUser="ExternalLinkService">
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<mods:title id="20A406951600783D41F9EB9683A2BDE0">Lampetra soljani, a new brook lamprey from the southern Adriatic Sea basin (Petromyzontiformes: Petromyzontidae)</mods:title>
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<mods:namePart id="08BED66C2947B545A64DF0EEDB1FE501">Tutman, Pero</mods:namePart>
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<mods:namePart id="DF7F20660C1B7E64DAA029EE47CAB538">Freyhof, Jörg</mods:namePart>
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<mods:namePart id="B40527440D6BFA6766B176B37C346A1F">Dulčić, Jakov</mods:namePart>
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<mods:namePart id="16B9F038CB09EF928A371C56CA98FA60">Glamuzina, Branko</mods:namePart>
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<mods:namePart id="423C1DEB70ECFEB5C2F10693D643AA6F">Geiger, Matthias</mods:namePart>
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<treatment id="AD0498777415FFA2FF1C2DE5FBA6FE90" ID-DOI="http://doi.org/10.5281/zenodo.6053348" ID-GBIF-Taxon="131812509" ID-Zenodo-Dep="6053348" LSID="urn:lsid:plazi:treatment:AD0498777415FFA2FF1C2DE5FBA6FE90" httpUri="http://treatment.plazi.org/id/AD0498777415FFA2FF1C2DE5FBA6FE90" lastPageId="10" lastPageNumber="540" pageId="5" pageNumber="536">
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<paragraph id="251229617415FFADFF1C2DE5FDF9FF62" blockId="5.[151,515,151,211]" box="[151,515,151,178]" pageId="5" pageNumber="536">
<heading id="7E5A9E0D7415FFADFF1C2DE5FDF9FF62" bold="true" box="[151,515,151,178]" fontSize="11" level="1" pageId="5" pageNumber="536" reason="1">
<emphasis id="17D9F5737415FFADFF1C2DE5FDF9FF62" bold="true" box="[151,515,151,178]" pageId="5" pageNumber="536">
<taxonomicName id="E2AD52E27415FFADFF1C2DE5FE9EFF61" box="[151,356,151,177]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="soljani" status="sp. nov.">
<emphasis id="17D9F5737415FFADFF1C2DE5FE9EFF61" bold="true" box="[151,356,151,177]" italics="true" pageId="5" pageNumber="536">Lampetra soljani</emphasis>
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, 
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<paragraph id="251229617415FFADFF1C2DC8FEEBFF03" blockId="5.[151,515,151,211]" box="[151,273,186,211]" pageId="5" pageNumber="536">
(
<figureCitation id="BD9635E47415FFADFF142DC8FEF3FF03" box="[159,265,186,211]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="5.[151,249,1228,1250]" captionStartId-1="5.[151,249,1675,1697]" captionStartId-2="5.[151,249,1977,1999]" captionStartId-3="6.[151,249,441,463]" captionStartId-4="6.[151,249,783,805]" captionTargetBox-0="[151,1436,464,1206]" captionTargetBox-1="[180,1392,1352,1637]" captionTargetBox-3="[151,1436,193,419]" captionTargetBox-4="[321,1266,573,761]" captionTargetId-0="figure@5.[151,1436,464,1206]" captionTargetId-1="figure@5.[151,1436,1330,1654]" captionTargetId-3="figure@6.[151,1436,193,419]" captionTargetId-4="figure@6.[321,1266,573,761]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="FIGURE 3. Lampetra soljani, from the top, ZFMK-ICH 103666, holotype, 109 mm TL; FSJF 3650, paratypes, 104 mm TL, 102 mm TL, 117 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-1="FIGURE 4. Lampetra soljani, FSJF 3650, paratypes, pre-spawning adults, from the top, 124 mm TL, 128 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-2="FIGURE 5. Lampetra soljani, FSJF 2184, paratype, freshly metamorphosed adult, 110 mm TL, Croatia: Norin at Konoba Vrilo; 21 Sept. 2006." captionText-3="FIGURE 6. Lampetra soljani, FSJF 1037, paratype, post-spawning adult, 111 mm TL, Bosnia-Herzegovina: Bregava north of Klepci; 12 May 2003." captionText-4="FIGURE 7. Lampetra soljani, not preserved, ammocoetes, about 130 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 17 Sep. 2014.." httpUri-0="https://zenodo.org/record/803764/files/figure.png" httpUri-1="https://zenodo.org/record/803766/files/figure.png" httpUri-3="https://zenodo.org/record/803768/files/figure.png" httpUri-4="https://zenodo.org/record/803770/files/figure.png" pageId="5" pageNumber="536">Figs. 3–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7415FFADFF1C2D8FFC90FE76" pageId="5" pageNumber="536" type="materials_examined">
<paragraph id="251229617415FFADFF1C2D8FFD23FEEA" blockId="5.[151,1436,253,422]" pageId="5" pageNumber="536">
<materialsCitation id="95C5233C7415FFADFF1C2D8FFD23FEEA" ID-GBIF-Occurrence="1563407827" collectingDate="2015-01-16" collectingDateMax="2015-01-17" collectingDateMin="2015-01-16" collectionCode="ZFMK-ICH" collectorName="P. Tutman" country="Bosnia and Herzegovina" location="Neretva River" municipality="Capljina" pageId="5" pageNumber="536" specimenCode="ZFMK-ICH 103666, 109" specimenCount="1" typeStatus="holotype">
<emphasis id="17D9F5737415FFADFF1C2D8FFEF1FEC6" bold="true" box="[151,267,253,278]" pageId="5" pageNumber="536">
<typeStatus id="FA1697C37415FFADFF1C2D8FFEFCFEC6" box="[151,262,253,278]" pageId="5" pageNumber="536" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="750B811A7415FFADFE992D8CFDD6FEC6" box="[274,556,253,278]" pageId="5" pageNumber="536">
<collectionCode id="43BCB1A47415FFADFE992D8CFE63FEC6" box="[274,409,254,278]" pageId="5" pageNumber="536">ZFMK-ICH</collectionCode>
<quantity id="E25584847415FFADFE2B2D8FFDD6FEC6" box="[416,556,253,278]" metricMagnitude="5" metricUnit="m" metricValue="1.03666109" pageId="5" pageNumber="536" unit="mm" value="1.03666109E8">103666, 109</quantity>
</specimenCode>
mm TL; 
<collectingCountry id="5DBA69F17415FFADFD1C2D8FFC55FEC6" box="[663,943,253,278]" name="Bosnia and Herzegovina" pageId="5" pageNumber="536">Bosnia and Herzegovina</collectingCountry>
: side-arm of 
<location id="20727FBA7415FFADFBCA2D8CFB24FEC6" LSID="urn:lsid:plazi:treatment:AD0498777415FFA2FF1C2DE5FBA6FE90:20727FBA7415FFADFBCA2D8CFB24FEC6" box="[1089,1246,253,278]" country="Bosnia and Herzegovina" municipality="Capljina" name="Neretva River" pageId="5" pageNumber="536">Neretva River</location>
east of 
<collectingMunicipality id="C576B31B7415FFADFABD2D8FFA62FEC6" box="[1334,1432,253,278]" pageId="5" pageNumber="536">Čapljina</collectingMunicipality>
, 43.106458 17.709536; 
<collectorName id="88584CB77415FFADFE102C50FDF4FEE9" box="[411,526,290,313]" pageId="5" pageNumber="536">P. Tutman</collectorName>
, 
<date id="51130FA17415FFADFD922C53FD2FFEEA" box="[537,725,289,314]" pageId="5" pageNumber="536" value="2015-01-16" valueMax="2015-01-17" valueMin="2015-01-16">
<collectingDate id="4157F6497415FFADFD922C53FD2FFEEA" box="[537,725,289,314]" pageId="5" pageNumber="536" value="2015-01-16" valueMax="2015-01-17" valueMin="2015-01-16">16–17 Jan. 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="251229617415FFADFF4C2C37FC90FE76" blockId="5.[151,1436,253,422]" pageId="5" pageNumber="536">
<materialsCitation id="95C5233C7415FFADFF4C2C37FC2CFE8E" ID-GBIF-Occurrence="1563407829" box="[199,982,325,350]" collectingDate="2015-01-16" collectingDateMax="2015-01-17" collectingDateMin="2015-01-16" collectionCode="FSJF" collectorName="P. Tutman" country="Bosnia and Herzegovina" location="Neretva River" municipality="Capljina" pageId="5" pageNumber="536" specimenCode="FSJF 3650, 12" specimenCount="1" typeStatus="paratype">
<emphasis id="17D9F5737415FFADFF4C2C37FEBCFE8E" bold="true" box="[199,326,325,350]" pageId="5" pageNumber="536">
<typeStatus id="FA1697C37415FFADFF4C2C37FEBBFE8E" box="[199,321,325,350]" pageId="5" pageNumber="536" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="750B811A7415FFADFEC42C34FE0DFE8E" box="[335,503,325,350]" pageId="5" pageNumber="536">
<collectionCode id="43BCB1A47415FFADFEC42C34FE73FE8E" box="[335,393,326,350]" pageId="5" pageNumber="536">FSJF</collectionCode>
3650, 12
</specimenCode>
, 105– 
<quantity id="E25584847415FFADFDB52C37FD66FE8D" box="[574,668,325,350]" metricMagnitude="-1" metricUnit="m" metricValue="1.26" pageId="5" pageNumber="536" unit="mm" value="126.0">126 mm</quantity>
TL; same data as holotype
</materialsCitation>
.—FSJF 1037, 
<quantity id="E25584847415FFADFB092C34FAE9FE8D" box="[1154,1299,326,350]" metricMagnitude="0" metricUnit="m" metricValue="1.1113" pageId="5" pageNumber="536" unit="mm" value="1111.3">1, 111.3 mm</quantity>
TL; 
<materialsCitation id="95C5233C7415FFADFAC52C37FA62FE51" ID-GBIF-Occurrence="1563407828" collectionCode="FSJF" country="Bosnia and Herzegovina" location="Bregava" municipality="Klepci" pageId="5" pageNumber="536" specimenCode="FSJF 2184, 1" specimenCount="1" typeStatus="paratype">
<collectingCountry id="5DBA69F17415FFADFAC52C37FEA0FE52" name="Bosnia and Herzegovina" pageId="5" pageNumber="536">Bosnia and Herzegovina</collectingCountry>
: lowermost stream 
<location id="20727FBA7415FFADFDBF2C18FD6BFE52" LSID="urn:lsid:plazi:treatment:AD0498777415FFA2FF1C2DE5FBA6FE90:20727FBA7415FFADFDBF2C18FD6BFE52" box="[564,657,362,386]" country="Bosnia and Herzegovina" municipality="Klepci" name="Bregava" pageId="5" pageNumber="536">Bregava</location>
north of 
<collectingMunicipality id="C576B31B7415FFADFD712C1BFCBDFE52" box="[762,839,361,386]" pageId="5" pageNumber="536">Klepci</collectingMunicipality>
, 43.052808 17.698089.— 
<specimenCode id="750B811A7415FFADFBE12C1BFAFBFE51" box="[1130,1281,361,386]" pageId="5" pageNumber="536">
<collectionCode id="43BCB1A47415FFADFBE12C1BFB5FFE52" box="[1130,1189,361,386]" pageId="5" pageNumber="536">FSJF</collectionCode>
2184, 
<quantity id="E25584847415FFADFB782C1BFAFBFE51" box="[1267,1281,361,385]" metricMagnitude="0" metricUnit="m" metricValue="1.1099999999999999" pageId="5" pageNumber="536" unit="mm" value="1110.0">1</quantity>
</specimenCode>
, 110 mm TL
</materialsCitation>
; Croatia: uppermost stream Norin at Prud, 43.095157 17,619551.
</paragraph>
</subSubSection>
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<paragraph id="251229617415FFADFF1C29BEFAEBFAD0" blockId="5.[151,1436,1228,1280]" pageId="5" pageNumber="536">
<emphasis id="17D9F5737415FFADFF1C29BEFEE9FB31" bold="true" box="[151,275,1228,1250]" pageId="5" pageNumber="536">FIGURE 3.</emphasis>
<taxonomicName id="E2AD52E27415FFADFE9029BFFE33FB32" box="[283,457,1228,1250]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737415FFADFE9029BFFE33FB32" box="[283,457,1228,1250]" italics="true" pageId="5" pageNumber="536">Lampetra soljani</emphasis>
</taxonomicName>
, from the top, ZFMK-ICH 103666, holotype, 109 mm TL; FSJF 3650, paratypes, 104 mm TL, 102 mm TL, 117 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16–17 Jan. 2015.
</paragraph>
</caption>
<caption id="71D279E97415FFADFF1C2BF9FC75F910" httpUri="https://zenodo.org/record/803766/files/figure.png" pageId="5" pageNumber="536" targetBox="[180,1392,1352,1637]" targetPageId="5">
<paragraph id="251229617415FFADFF1C2BF9FC75F910" blockId="5.[151,1435,1675,1728]" pageId="5" pageNumber="536">
<emphasis id="17D9F5737415FFADFF1C2BF9FEE8F970" bold="true" box="[151,274,1675,1697]" pageId="5" pageNumber="536">FIGURE 4.</emphasis>
<taxonomicName id="E2AD52E27415FFADFE922BFEFE3CF971" box="[281,454,1675,1697]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737415FFADFE922BFEFE3CF971" box="[281,454,1675,1697]" italics="true" pageId="5" pageNumber="536">Lampetra soljani</emphasis>
</taxonomicName>
, FSJF 3650, paratypes, pre-spawning adults, from the top, 124 mm TL, 128 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16–17 Jan. 2015.
</paragraph>
</caption>
<caption id="71D279E97415FFADFF1C2ACBFE91F83E" pageId="5" pageNumber="536">
<paragraph id="251229617415FFADFF1C2ACBFE91F83E" blockId="5.[151,1436,1977,2030]" pageId="5" pageNumber="536">
<emphasis id="17D9F5737415FFADFF1C2ACBFEEFF81F" bold="true" box="[151,277,1977,1999]" pageId="5" pageNumber="536">FIGURE 5.</emphasis>
<taxonomicName id="E2AD52E27415FFADFE952AC8FE35F81F" box="[286,463,1977,1999]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737415FFADFE952AC8FE35F81F" box="[286,463,1977,1999]" italics="true" pageId="5" pageNumber="536">Lampetra soljani</emphasis>
</taxonomicName>
, FSJF 2184, paratype, freshly metamorphosed adult, 110 mm TL, Croatia: Norin at Konoba Vrilo; 21 Sept. 2006.
</paragraph>
</caption>
<caption id="71D279E97416FFAEFF1C2CCBFE8CFE3D" httpUri="https://zenodo.org/record/803768/files/figure.png" pageId="6" pageNumber="537" targetBox="[151,1436,193,419]" targetPageId="6">
<paragraph id="251229617416FFAEFF1C2CCBFE8CFE3D" blockId="6.[151,1435,441,493]" pageId="6" pageNumber="537">
<emphasis id="17D9F5737416FFAEFF1C2CCBFEE9FE1E" bold="true" box="[151,275,441,463]" pageId="6" pageNumber="537">FIGURE 6.</emphasis>
<taxonomicName id="E2AD52E27416FFAEFE922CC8FE3DFE1F" box="[281,455,441,463]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFE922CC8FE3DFE1F" box="[281,455,441,463]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
, FSJF 1037, paratype, post-spawning adult, 111 mm TL, Bosnia-Herzegovina: Bregava north of Klepci; 12 May 2003.
</paragraph>
</caption>
<caption id="71D279E97416FFAEFF1C2E7DFDEEFC93" httpUri="https://zenodo.org/record/803770/files/figure.png" pageId="6" pageNumber="537" targetBox="[321,1266,573,761]" targetPageId="6">
<paragraph id="251229617416FFAEFF1C2E7DFDEEFC93" blockId="6.[151,1436,783,835]" pageId="6" pageNumber="537">
<emphasis id="17D9F5737416FFAEFF1C2E7DFEE9FCF4" bold="true" box="[151,275,783,805]" pageId="6" pageNumber="537">FIGURE 7.</emphasis>
<taxonomicName id="E2AD52E27416FFAEFE902E62FE33FCF5" box="[283,457,783,805]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFE902E62FE33FCF5" box="[283,457,783,805]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
, not preserved, ammocoetes, about 130 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 17 Sep. 2014..
</paragraph>
</caption>
<subSubSection id="6DB77AEA7416FFAEFF4C2E19FCDFFADF" pageId="6" pageNumber="537" type="diagnosis">
<paragraph id="251229617416FFAEFF4C2E19FDF4FB73" blockId="6.[151,1437,875,2016]" pageId="6" pageNumber="537">
<emphasis id="17D9F5737416FFAEFF4C2E19FEBBFC54" bold="true" box="[199,321,875,900]" pageId="6" pageNumber="537">Diagnosis.</emphasis>
<taxonomicName id="E2AD52E27416FFAEFEDB2E1EFDE2FC53" box="[336,536,875,899]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFEDB2E1EFDE2FC53" box="[336,536,875,899]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
is distinguished from all other species of 
<taxonomicName id="E2AD52E27416FFAEFBA62E1EFB60FC53" box="[1069,1178,876,899]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="17D9F5737416FFAEFBA62E1EFB60FC53" box="[1069,1178,876,899]" italics="true" pageId="6" pageNumber="537">Lampetra</emphasis>
</taxonomicName>
species by having a marmorate flank pattern in live, fully grown ammocoetes, and in some adults (vs. plain colour pattern). It is further distinguished by a combination of characters. 
<taxonomicName id="E2AD52E27416FFAEFD102EC6FCA1FC1B" box="[667,859,947,971]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFD102EC6FCA1FC1B" box="[667,859,947,971]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="E2AD52E27416FFAEFBA72EC6FB3AFC1B" box="[1068,1216,947,971]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="17D9F5737416FFAEFBA72EC6FB3AFC1B" box="[1068,1216,947,971]" italics="true" pageId="6" pageNumber="537">L. lanceolata</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2AD52E27416FFAEFB7D2EC6FAA8FC1B" box="[1270,1362,948,971]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFB7D2EC6FAA8FC1B" box="[1270,1362,948,971]" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
and is distinguished from these two species by having three velar tentacles (vs. 
<quantity id="E25584847416FFAEFC542EA5FBF6FC20" box="[991,1036,983,1008]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="537" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFB9E2EAAFB50FC3F" box="[1045,1194,983,1007]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="17D9F5737416FFAEFB9E2EAAFB50FC3F" box="[1045,1194,983,1007]" italics="true" pageId="6" pageNumber="537">L. lanceolata</emphasis>
</taxonomicName>
, 
<quantity id="E25584847416FFAEFB302EA5FAFAFC20" box="[1211,1280,983,1008]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="6" pageNumber="537" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">5-7 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFA832EAAFA9CFC3F" box="[1288,1382,984,1007]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFA832EAAFA9CFC3F" box="[1288,1382,984,1007]" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
), no posterial teeth (vs. rarely absent, usually a single incomplete row with up to 23 teeth in 
<taxonomicName id="E2AD52E27416FFAEFB072E8EFB10FBC3" box="[1164,1258,1020,1043]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFB072E8EFB10FBC3" box="[1164,1258,1020,1043]" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
) and less trunk myomeres (trunk myomeres between last branchial opening and anus 54–57 vs. 
<quantity id="E25584847416FFAEFB9F296DFB8CFBE8" box="[1044,1142,1055,1080]" metricMagnitude="0" metricUnit="m" metricValue="1.5367" metricValueMax="1.6256" metricValueMin="1.4478" pageId="6" pageNumber="537" unit="in" value="60.5" valueMax="64.0" valueMin="57.0">57–64 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFBF72952FAF5FBE7" box="[1148,1295,1055,1079]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="17D9F5737416FFAEFBF72952FAF5FBE7" box="[1148,1295,1055,1079]" italics="true" pageId="6" pageNumber="537">L. lanceolata</emphasis>
</taxonomicName>
, 
<quantity id="E25584847416FFAEFA97296DFA85FBE8" box="[1308,1407,1055,1080]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" metricValueMax="1.5493999999999999" metricValueMin="1.4478" pageId="6" pageNumber="537" unit="in" value="59.0" valueMax="61.0" valueMin="57.0">57–61 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFA0E2952FF2FFB8B" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFA0E2952FF2FFB8B" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
; trunk myomeres between last branchial opening and first dorsal fin origin 28–32 vs. 
<quantity id="E25584847416FFAEFB1C2931FB00FB8C" box="[1175,1274,1091,1116]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" metricValueMax="9.144" metricValueMin="8.128" pageId="6" pageNumber="537" unit="in" value="34.0" valueMax="36.0" valueMin="32.0">32–36 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFA8A2936FA6FFB8B" box="[1281,1429,1091,1115]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="17D9F5737416FFAEFA8A2936FA6FFB8B" box="[1281,1429,1091,1115]" italics="true" pageId="6" pageNumber="537">L. lanceolata</emphasis>
</taxonomicName>
, 
<quantity id="E25584847416FFAEFF1C2915FF03FB50" box="[151,249,1127,1152]" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="8.89" metricValueMin="8.128" pageId="6" pageNumber="537" unit="in" value="33.5" valueMax="35.0" valueMin="32.0">32–35 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFE8B291AFEA1FBAF" box="[256,347,1128,1151]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFE8B291AFEA1FBAF" box="[256,347,1128,1151]" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
; trunk myomeres between last branchial opening and second dorsal-fin origin 42–46 vs. 
<quantity id="E25584847416FFAEFAB12915FA66FB50" box="[1338,1436,1127,1152]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" metricValueMax="1.27" metricValueMin="1.1684" pageId="6" pageNumber="537" unit="in" value="48.0" valueMax="50.0" valueMin="46.0">46–50 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFF1C29FEFED0FB73" box="[151,298,1163,1187]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="17D9F5737416FFAEFF1C29FEFED0FB73" box="[151,298,1163,1187]" italics="true" pageId="6" pageNumber="537">L. lanceolata</emphasis>
</taxonomicName>
, 
<quantity id="E25584847416FFAEFEB229F9FE61FB74" box="[313,411,1163,1188]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" metricValueMax="1.2446" metricValueMin="1.143" pageId="6" pageNumber="537" unit="in" value="47.0" valueMax="49.0" valueMin="45.0">45–49 in</quantity>
<taxonomicName id="E2AD52E27416FFAEFE2929FEFE04FB73" box="[418,510,1164,1187]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="ninae">
<emphasis id="17D9F5737416FFAEFE2929FEFE04FB73" box="[418,510,1164,1187]" italics="true" pageId="6" pageNumber="537">L. ninae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="251229617416FFAEFF4C29C2FCDFFADF" blockId="6.[151,1437,875,2016]" pageId="6" pageNumber="537">
<taxonomicName id="E2AD52E27416FFAEFF4C29C2FE73FB17" box="[199,393,1199,1223]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFF4C29C2FE73FB17" box="[199,393,1199,1223]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="E2AD52E27416FFAEFD6F29C2FC7BFB17" box="[740,897,1199,1223]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="zanandreai">
<emphasis id="17D9F5737416FFAEFD6F29C2FC7BFB17" box="[740,897,1199,1223]" italics="true" pageId="6" pageNumber="537">L. zanandreai</emphasis>
</taxonomicName>
by lacking posterial teeth (vs. usually present, very rarely absent). 
<taxonomicName id="E2AD52E27416FFAEFE0929A6FD8DFB3B" box="[386,631,1235,1259]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="zanandreai">
<emphasis id="17D9F5737416FFAEFE0929A6FD8DFB3B" box="[386,631,1235,1259]" italics="true" pageId="6" pageNumber="537">Lampetra zanandreai</emphasis>
</taxonomicName>
has a single row of posterials with up to 20 teeth but there are also individuals without posterial teeth (
<bibRefCitation id="413C54907416FFAEFDA82985FCE2FADF" author="Hubbs" box="[547,792,1271,1296]" pageId="6" pageNumber="537" refString="Hubbs, C. L. &amp; Potter, I. C. (1971) Distribution, phylogeny and taxonomy. In: Hardisty, M. W. &amp; Potter, I. C. (Eds.), The biology of lampreys. Fol 1. Academic Press, London &amp; New York, pp. 1 - 65" type="journal article" year="1971">Hubbs &amp; Potter 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7416FFAEFF4C286EFB9EF9E0" pageId="6" pageNumber="537" type="reference_group">
<paragraph id="251229617416FFAEFF4C286EFB9EF9E0" blockId="6.[151,1437,875,2016]" pageId="6" pageNumber="537">
<taxonomicName id="E2AD52E27416FFAEFF4C286EFE73FAE3" box="[199,393,1307,1331]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFF4C286EFE73FAE3" box="[199,393,1307,1331]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="E2AD52E27416FFAEFD05286EFCE2FAE3" box="[654,792,1307,1331]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="fluviatilis">
<emphasis id="17D9F5737416FFAEFD05286EFCE2FAE3" box="[654,792,1307,1331]" italics="true" pageId="6" pageNumber="537">L. fluviatilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2AD52E27416FFAEFCDF286EFC3FFAE3" box="[852,965,1307,1331]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="planeri">
<emphasis id="17D9F5737416FFAEFCDF286EFC3FFAE3" box="[852,965,1307,1331]" italics="true" pageId="6" pageNumber="537">L. planeri</emphasis>
</taxonomicName>
by having a bicuspid middle endolateral tooth (vs. almost always tricuspid) and the infraoral lamina with 5 teeth (vs. usually 7–9, very rarely 5–6). The Portuguese species 
<taxonomicName id="E2AD52E27416FFAEFEFF2816FDE1FAAB" box="[372,539,1379,1403]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="alavariensis">
<emphasis id="17D9F5737416FFAEFEFF2816FDE1FAAB" box="[372,539,1379,1403]" italics="true" pageId="6" pageNumber="537">L. alavariensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E2AD52E27416FFAEFDA72816FD32FAAB" box="[556,712,1380,1403]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="auremensis">
<emphasis id="17D9F5737416FFAEFDA72816FD32FAAB" box="[556,712,1380,1403]" italics="true" pageId="6" pageNumber="537">L. auremensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2AD52E27416FFAEFC892816FC75FAAB" box="[770,911,1379,1403]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lusitanica">
<emphasis id="17D9F5737416FFAEFC892816FC75FAAB" box="[770,911,1379,1403]" italics="true" pageId="6" pageNumber="537">L. lusitanica</emphasis>
</taxonomicName>
often, but not always, have a bicuspid middle endolateral tooth. 
<taxonomicName id="E2AD52E27416FFAEFEEB28FAFDDAFA4F" box="[352,544,1415,1439]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737416FFAEFEEB28FAFDDAFA4F" box="[352,544,1415,1439]" italics="true" pageId="6" pageNumber="537">Lampetra soljani</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="E2AD52E27416FFAEFC9328FAFC44FA4F" box="[792,958,1415,1439]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="alavariensis">
<emphasis id="17D9F5737416FFAEFC9328FAFC44FA4F" box="[792,958,1415,1439]" italics="true" pageId="6" pageNumber="537">L. alavariensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E2AD52E27416FFAEFC4028FAFB9CFA4F" box="[971,1126,1416,1439]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="auremensis">
<emphasis id="17D9F5737416FFAEFC4028FAFB9CFA4F" box="[971,1126,1416,1439]" italics="true" pageId="6" pageNumber="537">L. auremensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2AD52E27416FFAEFB1728FAFADDFA4F" box="[1180,1319,1415,1439]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lusitanica">
<emphasis id="17D9F5737416FFAEFB1728FAFADDFA4F" box="[1180,1319,1415,1439]" italics="true" pageId="6" pageNumber="537">L. lusitanica</emphasis>
</taxonomicName>
by having 54–57 trunk myomeres between the last branchial opening and the anus (vs. 57–63). 
<taxonomicName id="E2AD52E27416FFAEFBCD28DEFADEFA13" box="[1094,1316,1451,1475]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="fluviatilis">
<emphasis id="17D9F5737416FFAEFBCD28DEFADEFA13" box="[1094,1316,1451,1475]" italics="true" pageId="6" pageNumber="537">Lampetra fluviatilis</emphasis>
</taxonomicName>
has 58–69 trunk myomeres (n= 883 from more than 14 sites; 
<bibRefCitation id="413C54907416FFAEFD6528BDFC64FA38" author="Hardisty" box="[750,926,1487,1512]" pageId="6" pageNumber="537" refString="Hardisty, M. W. (1986 a) Lampetra fluciatilis (Linnaeus, 1758). In: Holcik, J. (Ed.), The freshwater fishes of Europe. Fol. 1. Part I. Petromyzontiformes. Aula Verlag, Wiesbaden, pp. 249 - 278." type="journal article" year="1986" yearSuffix="a">Hardisty 1986a</bibRefCitation>
) and 
<taxonomicName id="E2AD52E27416FFAEFC6F28A2FBACFA37" box="[996,1110,1487,1511]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="planeri">
<emphasis id="17D9F5737416FFAEFC6F28A2FBACFA37" box="[996,1110,1487,1511]" italics="true" pageId="6" pageNumber="537">L. planeri</emphasis>
</taxonomicName>
has 54–69 trunk myomeres (n=2385 from 18 sites; 
<bibRefCitation id="413C54907416FFAEFE2E2881FDA3F9DC" author="Hardisty" box="[421,601,1523,1548]" pageId="6" pageNumber="537" refString="Hardisty, M. W. (1986 b) Lampetra planeri (Bloch, 1784). In: Holcik, J. (Ed.), The freshwater fishes of Europe. Fol. 1. Part I. Petromyzontiformes. Aula Verlag, Wiesbaden, pp. 279 - 304" type="journal article" year="1986" yearSuffix="b">Hardisty 1986b</bibRefCitation>
). However, 54 and 56 are minimum values of ammocoetes just from one site each and usually, the number of trunk myomeres ranges from 57–67 (mean 60–62)
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7416FFA0FF4C2B49FE83FF28" lastPageId="8" lastPageNumber="539" pageId="6" pageNumber="537" type="description">
<paragraph id="251229617416FFAEFF4C2B49FC97F830" blockId="6.[151,1437,875,2016]" pageId="6" pageNumber="537">
<emphasis id="17D9F5737416FFAEFF4C2B49FEA3F984" bold="true" box="[199,345,1595,1620]" pageId="6" pageNumber="537">Description.</emphasis>
For general appearance see 
<figureCitation id="BD9635E47416FFAEFD1D2B49FCF8F983" box="[662,770,1595,1620]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="5.[151,249,1228,1250]" captionStartId-1="5.[151,249,1675,1697]" captionStartId-2="5.[151,249,1977,1999]" captionStartId-3="6.[151,249,441,463]" captionStartId-4="6.[151,249,783,805]" captionTargetBox-0="[151,1436,464,1206]" captionTargetBox-1="[180,1392,1352,1637]" captionTargetBox-3="[151,1436,193,419]" captionTargetBox-4="[321,1266,573,761]" captionTargetId-0="figure@5.[151,1436,464,1206]" captionTargetId-1="figure@5.[151,1436,1330,1654]" captionTargetId-3="figure@6.[151,1436,193,419]" captionTargetId-4="figure@6.[321,1266,573,761]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="FIGURE 3. Lampetra soljani, from the top, ZFMK-ICH 103666, holotype, 109 mm TL; FSJF 3650, paratypes, 104 mm TL, 102 mm TL, 117 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-1="FIGURE 4. Lampetra soljani, FSJF 3650, paratypes, pre-spawning adults, from the top, 124 mm TL, 128 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-2="FIGURE 5. Lampetra soljani, FSJF 2184, paratype, freshly metamorphosed adult, 110 mm TL, Croatia: Norin at Konoba Vrilo; 21 Sept. 2006." captionText-3="FIGURE 6. Lampetra soljani, FSJF 1037, paratype, post-spawning adult, 111 mm TL, Bosnia-Herzegovina: Bregava north of Klepci; 12 May 2003." captionText-4="FIGURE 7. Lampetra soljani, not preserved, ammocoetes, about 130 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 17 Sep. 2014.." httpUri-0="https://zenodo.org/record/803764/files/figure.png" httpUri-1="https://zenodo.org/record/803766/files/figure.png" httpUri-3="https://zenodo.org/record/803768/files/figure.png" httpUri-4="https://zenodo.org/record/803770/files/figure.png" pageId="6" pageNumber="537">Figs. 3–7</figureCitation>
. Morphometric data of pre-spawning adults are provided in 
<tableCitation id="682F1CDA7416FFAEFF1C2B2DFF14F9A7" box="[151,238,1631,1656]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Morphometric data of L. soljani (holotype ZFMK-ICH 103666, paratypes FSJF 3650; n = 10). The calculations include the holotype." pageId="6" pageNumber="537">Table 2</tableCitation>
. Oral discs are shown in 
<figureCitation id="BD9635E47416FFAEFD992B2DFDADF9A7" box="[530,599,1631,1656]" captionStart="FIGURE 8" captionStartId="7.[151,249,1838,1860]" captionTargetBox="[151,1436,1282,1816]" captionTargetId="figure@7.[151,1436,1282,1816]" captionTargetPageId="7" captionText="FIGURE 8. Lampetra soljani, FSJF 3650, paratypes, about 120 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." httpUri="https://zenodo.org/record/803772/files/figure.png" pageId="6" pageNumber="537">Fig. 8</figureCitation>
. Trunk myomeres between last branchial opening and anus 54–57 (mode 55), between last branchial opening and origin of first dorsal-fin 28–32 (mode 28) and between last branchial opening and origin of second dorsal fin 42–46 (mode 43). Caudal fin spade like. Transverse lingual lamina not fully developed in individuals examined, with 0–4 (mode 2) unicuspid teeth on each side and one enlarged central tooth. In 14 individuals examined for this character, four without teeth on transverse lingual lamina and usually, there are 2–3 lateral teeth, 4 on one side in one individual. Longitudinal lingual laminae straight, each with 4–6 unicuspid teeth, no teeth seen in 12 out of 14 individuals examined. Supraoral lamina with two unicuspid teeth separated by a toothless bridge. Infraoral lamina with 5 teeth, outer two teeth bicuspid, inner three teeth unicuspid. Three bicuspid endolaterals on both sides of oral disc. Anterials in 1–2 rows, usually 1 row, inner row with 4–7 (mode 5 teeth) unicuspid teeth. Exolateral and posterial teeth absent. Three velar tentacles (n=6), middle one shorter than adjacent lateral tentacles, with tubercles on dorsal surface, without wings.
</paragraph>
<caption id="71D279E97417FFAFFF1C2DEAFE04FF1F" pageId="7" pageNumber="538">
<paragraph id="251229617417FFAFFF1C2DEAFE04FF1F" blockId="7.[151,1435,151,207]" pageId="7" pageNumber="538">
<emphasis id="17D9F5737417FFAFFF1C2DEAFEEAFF7F" bold="true" box="[151,272,152,175]" pageId="7" pageNumber="538">TABLE 2.</emphasis>
Morphometric data of 
<taxonomicName id="E2AD52E27417FFAFFDA72DEAFD60FF7F" box="[556,666,151,175]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737417FFAFFDA72DEAFD60FF7F" box="[556,666,151,175]" italics="true" pageId="7" pageNumber="538">L. soljani</emphasis>
</taxonomicName>
(holotype ZFMK-ICH 103666, paratypes FSJF 3650; n=10). The calculations include the holotype.
</paragraph>
</caption>
<paragraph id="251229617417FFAFFD5D2D98FAC2FB67" pageId="7" pageNumber="538">
<table id="57ADDBC174170057FF142D98FAC2FB67" box="[159,1336,234,1207]" gridcols="6" gridrows="24" pageId="7" pageNumber="538">
<tr id="9B9D2B2374170057FF142D98FAC2FED0" box="[159,1336,234,256]" gridrow="0" pageId="7" pageNumber="538" rowspan-0="1" rowspan-4="1" rowspan-5="1">
<th id="D84C425F74170057FD5D2D98FCD5FED0" box="[726,815,234,256]" gridcol="1" gridrow="0" pageId="7" pageNumber="538">holotype</th>
<th id="D84C425F74170057FC1F2D98FBFDFED0" box="[916,1031,234,256]" gridcol="2" gridrow="0" pageId="7" pageNumber="538">holotype &amp;</th>
<th id="D84C425F74170057FB862D98FB94FED0" box="[1037,1134,234,256]" gridcol="3" gridrow="0" pageId="7" pageNumber="538">paratypes</th>
</tr>
<tr id="9B9D2B2374170057FF142C66FAC2FEFA" box="[159,1336,276,298]" gridrow="1" pageId="7" pageNumber="538" rowspan-1="1" rowspan-4="1">
<th id="D84C425F74170057FF142C66FD38FEFA" box="[159,706,276,298]" gridcol="0" gridrow="1" pageId="7" pageNumber="538">Total length</th>
<td id="D84C425F74170057FC1F2C66FBFDFEFA" box="[916,1031,276,298]" gridcol="2" gridrow="1" pageId="7" pageNumber="538">mean</td>
<td id="D84C425F74170057FB862C66FB94FEFA" box="[1037,1134,276,298]" gridcol="3" gridrow="1" pageId="7" pageNumber="538">range</td>
<td id="D84C425F74170057FA932C66FAC2FEFA" box="[1304,1336,276,298]" gridcol="5" gridrow="1" pageId="7" pageNumber="538">SD</td>
</tr>
<tr id="9B9D2B2374170057FF142C4FFAC2FE83" box="[159,1336,317,339]" gridrow="2" pageId="7" pageNumber="538" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="D84C425F74170057FF142C4FFD38FE83" box="[159,706,317,339]" gridcol="0" gridrow="2" pageId="7" pageNumber="538">In percent of total length</th>
</tr>
<tr id="9B9D2B2374170057FF142C14FAC2FEAC" box="[159,1336,358,380]" gridrow="3" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142C14FD38FEAC" box="[159,706,358,380]" gridcol="0" gridrow="3" pageId="7" pageNumber="538">Prebranchial length</th>
<td id="D84C425F74170057FD5D2C14FCD5FEAC" box="[726,815,358,380]" gridcol="1" gridrow="3" pageId="7" pageNumber="538">11.0</td>
<td id="D84C425F74170057FC1F2C14FBFDFEAC" box="[916,1031,358,380]" gridcol="2" gridrow="3" pageId="7" pageNumber="538">10.6</td>
<td id="D84C425F74170057FB862C14FB94FEAC" box="[1037,1134,358,380]" gridcol="3" gridrow="3" pageId="7" pageNumber="538">10.1</td>
<td id="D84C425F74170057FB2E2C14FB2BFEAC" box="[1189,1233,358,380]" gridcol="4" gridrow="3" pageId="7" pageNumber="538">11.4</td>
<td id="D84C425F74170057FA932C14FAC2FEAC" box="[1304,1336,358,380]" gridcol="5" gridrow="3" pageId="7" pageNumber="538">0.5</td>
</tr>
<tr id="9B9D2B2374170057FF142CE2FAC2FE76" box="[159,1336,400,422]" gridrow="4" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142CE2FD38FE76" box="[159,706,400,422]" gridcol="0" gridrow="4" pageId="7" pageNumber="538">Branchial length</th>
<td id="D84C425F74170057FD5D2CE2FCD5FE76" box="[726,815,400,422]" gridcol="1" gridrow="4" pageId="7" pageNumber="538">10.5</td>
<td id="D84C425F74170057FC1F2CE2FBFDFE76" box="[916,1031,400,422]" gridcol="2" gridrow="4" pageId="7" pageNumber="538">10.5</td>
<td id="D84C425F74170057FB862CE2FB94FE76" box="[1037,1134,400,422]" gridcol="3" gridrow="4" pageId="7" pageNumber="538">9.9</td>
<td id="D84C425F74170057FB2E2CE2FB2BFE76" box="[1189,1233,400,422]" gridcol="4" gridrow="4" pageId="7" pageNumber="538">11.4</td>
<td id="D84C425F74170057FA932CE2FAC2FE76" box="[1304,1336,400,422]" gridcol="5" gridrow="4" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF142CCBFAC2FE1F" box="[159,1336,441,463]" gridrow="5" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142CCBFD38FE1F" box="[159,706,441,463]" gridcol="0" gridrow="5" pageId="7" pageNumber="538">Interbranchial opening length</th>
<td id="D84C425F74170057FD5D2CCBFCD5FE1F" box="[726,815,441,463]" gridcol="1" gridrow="5" pageId="7" pageNumber="538">1.2</td>
<td id="D84C425F74170057FC1F2CCBFBFDFE1F" box="[916,1031,441,463]" gridcol="2" gridrow="5" pageId="7" pageNumber="538">1.7</td>
<td id="D84C425F74170057FB862CCBFB94FE1F" box="[1037,1134,441,463]" gridcol="3" gridrow="5" pageId="7" pageNumber="538">1.4</td>
<td id="D84C425F74170057FB2E2CCBFB2BFE1F" box="[1189,1233,441,463]" gridcol="4" gridrow="5" pageId="7" pageNumber="538">2.0</td>
<td id="D84C425F74170057FA932CCBFAC2FE1F" box="[1304,1336,441,463]" gridcol="5" gridrow="5" pageId="7" pageNumber="538">0.2</td>
</tr>
<tr id="9B9D2B2374170057FF142C90FAC2FE28" box="[159,1336,482,504]" gridrow="6" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142C90FD38FE28" box="[159,706,482,504]" gridcol="0" gridrow="6" pageId="7" pageNumber="538">Branchial depth</th>
<td id="D84C425F74170057FD5D2C90FCD5FE28" box="[726,815,482,504]" gridcol="1" gridrow="6" pageId="7" pageNumber="538">6.3</td>
<td id="D84C425F74170057FC1F2C90FBFDFE28" box="[916,1031,482,504]" gridcol="2" gridrow="6" pageId="7" pageNumber="538">6.1</td>
<td id="D84C425F74170057FB862C90FB94FE28" box="[1037,1134,482,504]" gridcol="3" gridrow="6" pageId="7" pageNumber="538">5.6</td>
<td id="D84C425F74170057FB2E2C90FB2BFE28" box="[1189,1233,482,504]" gridcol="4" gridrow="6" pageId="7" pageNumber="538">6.6</td>
<td id="D84C425F74170057FA932C90FAC2FE28" box="[1304,1336,482,504]" gridcol="5" gridrow="6" pageId="7" pageNumber="538">0.3</td>
</tr>
<tr id="9B9D2B2374170057FF142F7EFAC2FDF2" box="[159,1336,524,546]" gridrow="7" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142F7EFD38FDF2" box="[159,706,524,546]" gridcol="0" gridrow="7" pageId="7" pageNumber="538">Disc length</th>
<td id="D84C425F74170057FD5D2F7EFCD5FDF2" box="[726,815,524,546]" gridcol="1" gridrow="7" pageId="7" pageNumber="538">5.0</td>
<td id="D84C425F74170057FC1F2F7EFBFDFDF2" box="[916,1031,524,546]" gridcol="2" gridrow="7" pageId="7" pageNumber="538">4.7</td>
<td id="D84C425F74170057FB862F7EFB94FDF2" box="[1037,1134,524,546]" gridcol="3" gridrow="7" pageId="7" pageNumber="538">4.3</td>
<td id="D84C425F74170057FB2E2F7EFB2BFDF2" box="[1189,1233,524,546]" gridcol="4" gridrow="7" pageId="7" pageNumber="538">5.2</td>
<td id="D84C425F74170057FA932F7EFAC2FDF2" box="[1304,1336,524,546]" gridcol="5" gridrow="7" pageId="7" pageNumber="538">0.3</td>
</tr>
<tr id="9B9D2B2374170057FF142F47FAC2FD9B" box="[159,1336,565,587]" gridrow="8" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142F47FD38FD9B" box="[159,706,565,587]" gridcol="0" gridrow="8" pageId="7" pageNumber="538">Eye length</th>
<td id="D84C425F74170057FD5D2F47FCD5FD9B" box="[726,815,565,587]" gridcol="1" gridrow="8" pageId="7" pageNumber="538">1.8</td>
<td id="D84C425F74170057FC1F2F47FBFDFD9B" box="[916,1031,565,587]" gridcol="2" gridrow="8" pageId="7" pageNumber="538">1.9</td>
<td id="D84C425F74170057FB862F47FB94FD9B" box="[1037,1134,565,587]" gridcol="3" gridrow="8" pageId="7" pageNumber="538">1.6</td>
<td id="D84C425F74170057FB2E2F47FB2BFD9B" box="[1189,1233,565,587]" gridcol="4" gridrow="8" pageId="7" pageNumber="538">2.0</td>
<td id="D84C425F74170057FA932F47FAC2FD9B" box="[1304,1336,565,587]" gridcol="5" gridrow="8" pageId="7" pageNumber="538">0.1</td>
</tr>
<tr id="9B9D2B2374170057FF142F2CFAC2FDA4" box="[159,1336,606,628]" gridrow="9" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142F2CFD38FDA4" box="[159,706,606,628]" gridcol="0" gridrow="9" pageId="7" pageNumber="538">Interocular width</th>
<td id="D84C425F74170057FD5D2F2CFCD5FDA4" box="[726,815,606,628]" gridcol="1" gridrow="9" pageId="7" pageNumber="538">3.5</td>
<td id="D84C425F74170057FC1F2F2CFBFDFDA4" box="[916,1031,606,628]" gridcol="2" gridrow="9" pageId="7" pageNumber="538">3.4</td>
<td id="D84C425F74170057FB862F2CFB94FDA4" box="[1037,1134,606,628]" gridcol="3" gridrow="9" pageId="7" pageNumber="538">2.7</td>
<td id="D84C425F74170057FB2E2F2CFB2BFDA4" box="[1189,1233,606,628]" gridcol="4" gridrow="9" pageId="7" pageNumber="538">4.1</td>
<td id="D84C425F74170057FA932F2CFAC2FDA4" box="[1304,1336,606,628]" gridcol="5" gridrow="9" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF142FFAFAC2FD4E" box="[159,1336,648,670]" gridrow="10" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142FFAFD38FD4E" box="[159,706,648,670]" gridcol="0" gridrow="10" pageId="7" pageNumber="538">Snout length</th>
<td id="D84C425F74170057FD5D2FFAFCD5FD4E" box="[726,815,648,670]" gridcol="1" gridrow="10" pageId="7" pageNumber="538">6.3</td>
<td id="D84C425F74170057FC1F2FFAFBFDFD4E" box="[916,1031,648,670]" gridcol="2" gridrow="10" pageId="7" pageNumber="538">6.2</td>
<td id="D84C425F74170057FB862FFAFB94FD4E" box="[1037,1134,648,670]" gridcol="3" gridrow="10" pageId="7" pageNumber="538">5.5</td>
<td id="D84C425F74170057FB2E2FFAFB2BFD4E" box="[1189,1233,648,670]" gridcol="4" gridrow="10" pageId="7" pageNumber="538">6.7</td>
<td id="D84C425F74170057FA932FFAFAC2FD4E" box="[1304,1336,648,670]" gridcol="5" gridrow="10" pageId="7" pageNumber="538">0.3</td>
</tr>
<tr id="9B9D2B2374170057FF142FC3FAC2FD17" box="[159,1336,689,711]" gridrow="11" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142FC3FD38FD17" box="[159,706,689,711]" gridcol="0" gridrow="11" pageId="7" pageNumber="538">Postocular length</th>
<td id="D84C425F74170057FD5D2FC3FCD5FD17" box="[726,815,689,711]" gridcol="1" gridrow="11" pageId="7" pageNumber="538">3.1</td>
<td id="D84C425F74170057FC1F2FC3FBFDFD17" box="[916,1031,689,711]" gridcol="2" gridrow="11" pageId="7" pageNumber="538">3.0</td>
<td id="D84C425F74170057FB862FC3FB94FD17" box="[1037,1134,689,711]" gridcol="3" gridrow="11" pageId="7" pageNumber="538">2.7</td>
<td id="D84C425F74170057FB2E2FC3FB2BFD17" box="[1189,1233,689,711]" gridcol="4" gridrow="11" pageId="7" pageNumber="538">3.3</td>
<td id="D84C425F74170057FA932FC3FAC2FD17" box="[1304,1336,689,711]" gridcol="5" gridrow="11" pageId="7" pageNumber="538">0.2</td>
</tr>
<tr id="9B9D2B2374170057FF142FA8FAC2FD20" box="[159,1336,730,752]" gridrow="12" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142FA8FD38FD20" box="[159,706,730,752]" gridcol="0" gridrow="12" pageId="7" pageNumber="538">Prenostril length</th>
<td id="D84C425F74170057FD5D2FA8FCD5FD20" box="[726,815,730,752]" gridcol="1" gridrow="12" pageId="7" pageNumber="538">4.7</td>
<td id="D84C425F74170057FC1F2FA8FBFDFD20" box="[916,1031,730,752]" gridcol="2" gridrow="12" pageId="7" pageNumber="538">4.8</td>
<td id="D84C425F74170057FB862FA8FB94FD20" box="[1037,1134,730,752]" gridcol="3" gridrow="12" pageId="7" pageNumber="538">4.4</td>
<td id="D84C425F74170057FB2E2FA8FB2BFD20" box="[1189,1233,730,752]" gridcol="4" gridrow="12" pageId="7" pageNumber="538">5.1</td>
<td id="D84C425F74170057FA932FA8FAC2FD20" box="[1304,1336,730,752]" gridcol="5" gridrow="12" pageId="7" pageNumber="538">0.3</td>
</tr>
<tr id="9B9D2B2374170057FF142E76FAC2FCCA" box="[159,1336,772,794]" gridrow="13" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142E76FD38FCCA" box="[159,706,772,794]" gridcol="0" gridrow="13" pageId="7" pageNumber="538">Trunk length</th>
<td id="D84C425F74170057FD5D2E76FCD5FCCA" box="[726,815,772,794]" gridcol="1" gridrow="13" pageId="7" pageNumber="538">50.6</td>
<td id="D84C425F74170057FC1F2E76FBFDFCCA" box="[916,1031,772,794]" gridcol="2" gridrow="13" pageId="7" pageNumber="538">51.2</td>
<td id="D84C425F74170057FB862E76FB94FCCA" box="[1037,1134,772,794]" gridcol="3" gridrow="13" pageId="7" pageNumber="538">50.3</td>
<td id="D84C425F74170057FB2E2E76FB2BFCCA" box="[1189,1233,772,794]" gridcol="4" gridrow="13" pageId="7" pageNumber="538">52.2</td>
<td id="D84C425F74170057FA932E76FAC2FCCA" box="[1304,1336,772,794]" gridcol="5" gridrow="13" pageId="7" pageNumber="538">0.6</td>
</tr>
<tr id="9B9D2B2374170057FF142E5FFAC2FC93" box="[159,1336,813,835]" gridrow="14" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142E5FFD38FC93" box="[159,706,813,835]" gridcol="0" gridrow="14" pageId="7" pageNumber="538">Trunk depth</th>
<td id="D84C425F74170057FD5D2E5FFCD5FC93" box="[726,815,813,835]" gridcol="1" gridrow="14" pageId="7" pageNumber="538">6.5</td>
<td id="D84C425F74170057FC1F2E5FFBFDFC93" box="[916,1031,813,835]" gridcol="2" gridrow="14" pageId="7" pageNumber="538">6.0</td>
<td id="D84C425F74170057FB862E5FFB94FC93" box="[1037,1134,813,835]" gridcol="3" gridrow="14" pageId="7" pageNumber="538">5.5</td>
<td id="D84C425F74170057FB2E2E5FFB2BFC93" box="[1189,1233,813,835]" gridcol="4" gridrow="14" pageId="7" pageNumber="538">6.7</td>
<td id="D84C425F74170057FA932E5FFAC2FC93" box="[1304,1336,813,835]" gridcol="5" gridrow="14" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF142E24FAC2FCBC" box="[159,1336,854,876]" gridrow="15" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142E24FD38FCBC" box="[159,706,854,876]" gridcol="0" gridrow="15" pageId="7" pageNumber="538">Tail length</th>
<td id="D84C425F74170057FD5D2E24FCD5FCBC" box="[726,815,854,876]" gridcol="1" gridrow="15" pageId="7" pageNumber="538">28.5</td>
<td id="D84C425F74170057FC1F2E24FBFDFCBC" box="[916,1031,854,876]" gridcol="2" gridrow="15" pageId="7" pageNumber="538">28.2</td>
<td id="D84C425F74170057FB862E24FB94FCBC" box="[1037,1134,854,876]" gridcol="3" gridrow="15" pageId="7" pageNumber="538">25.9</td>
<td id="D84C425F74170057FB2E2E24FB2BFCBC" box="[1189,1233,854,876]" gridcol="4" gridrow="15" pageId="7" pageNumber="538">30.1</td>
<td id="D84C425F74170057FA932E24FAC2FCBC" box="[1304,1336,854,876]" gridcol="5" gridrow="15" pageId="7" pageNumber="538">1.3</td>
</tr>
<tr id="9B9D2B2374170057FF142EF2FAC2FC46" box="[159,1336,896,918]" gridrow="16" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142EF2FD38FC46" box="[159,706,896,918]" gridcol="0" gridrow="16" pageId="7" pageNumber="538">First dorsal-fin length</th>
<td id="D84C425F74170057FD5D2EF2FCD5FC46" box="[726,815,896,918]" gridcol="1" gridrow="16" pageId="7" pageNumber="538">13.5</td>
<td id="D84C425F74170057FC1F2EF2FBFDFC46" box="[916,1031,896,918]" gridcol="2" gridrow="16" pageId="7" pageNumber="538">13.6</td>
<td id="D84C425F74170057FB862EF2FB94FC46" box="[1037,1134,896,918]" gridcol="3" gridrow="16" pageId="7" pageNumber="538">12.0</td>
<td id="D84C425F74170057FB2E2EF2FB2BFC46" box="[1189,1233,896,918]" gridcol="4" gridrow="16" pageId="7" pageNumber="538">18.3</td>
<td id="D84C425F74170057FA932EF2FAC2FC46" box="[1304,1336,896,918]" gridcol="5" gridrow="16" pageId="7" pageNumber="538">1.8</td>
</tr>
<tr id="9B9D2B2374170057FF142EDBFAC2FC6F" box="[159,1336,937,959]" gridrow="17" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142EDBFD38FC6F" box="[159,706,937,959]" gridcol="0" gridrow="17" pageId="7" pageNumber="538">First dorsal-fin depth</th>
<td id="D84C425F74170057FD5D2EDBFCD5FC6F" box="[726,815,937,959]" gridcol="1" gridrow="17" pageId="7" pageNumber="538">2.5</td>
<td id="D84C425F74170057FC1F2EDBFBFDFC6F" box="[916,1031,937,959]" gridcol="2" gridrow="17" pageId="7" pageNumber="538">1.8</td>
<td id="D84C425F74170057FB862EDBFB94FC6F" box="[1037,1134,937,959]" gridcol="3" gridrow="17" pageId="7" pageNumber="538">1.4</td>
<td id="D84C425F74170057FB2E2EDBFB2BFC6F" box="[1189,1233,937,959]" gridcol="4" gridrow="17" pageId="7" pageNumber="538">2.7</td>
<td id="D84C425F74170057FA932EDBFAC2FC6F" box="[1304,1336,937,959]" gridcol="5" gridrow="17" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF142EA0FAC2FC38" box="[159,1336,978,1000]" gridrow="18" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142EA0FD38FC38" box="[159,706,978,1000]" gridcol="0" gridrow="18" pageId="7" pageNumber="538">Second dorsal-fin length</th>
<td id="D84C425F74170057FD5D2EA0FCD5FC38" box="[726,815,978,1000]" gridcol="1" gridrow="18" pageId="7" pageNumber="538">25.0</td>
<td id="D84C425F74170057FC1F2EA0FBFDFC38" box="[916,1031,978,1000]" gridcol="2" gridrow="18" pageId="7" pageNumber="538">26.1</td>
<td id="D84C425F74170057FB862EA0FB94FC38" box="[1037,1134,978,1000]" gridcol="3" gridrow="18" pageId="7" pageNumber="538">23.5</td>
<td id="D84C425F74170057FB2E2EA0FB2BFC38" box="[1189,1233,978,1000]" gridcol="4" gridrow="18" pageId="7" pageNumber="538">27.8</td>
<td id="D84C425F74170057FA932EA0FAC2FC38" box="[1304,1336,978,1000]" gridcol="5" gridrow="18" pageId="7" pageNumber="538">1.5</td>
</tr>
<tr id="9B9D2B2374170057FF142E8EFAC2FBC2" box="[159,1336,1020,1042]" gridrow="19" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142E8EFD38FBC2" box="[159,706,1020,1042]" gridcol="0" gridrow="19" pageId="7" pageNumber="538">Second dorsal-fin depth</th>
<td id="D84C425F74170057FD5D2E8EFCD5FBC2" box="[726,815,1020,1042]" gridcol="1" gridrow="19" pageId="7" pageNumber="538">4.0</td>
<td id="D84C425F74170057FC1F2E8EFBFDFBC2" box="[916,1031,1020,1042]" gridcol="2" gridrow="19" pageId="7" pageNumber="538">3.2</td>
<td id="D84C425F74170057FB862E8EFB94FBC2" box="[1037,1134,1020,1042]" gridcol="3" gridrow="19" pageId="7" pageNumber="538">2.5</td>
<td id="D84C425F74170057FB2E2E8EFB2BFBC2" box="[1189,1233,1020,1042]" gridcol="4" gridrow="19" pageId="7" pageNumber="538">3.8</td>
<td id="D84C425F74170057FA932E8EFAC2FBC2" box="[1304,1336,1020,1042]" gridcol="5" gridrow="19" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF142957FAC2FBEB" box="[159,1336,1061,1083]" gridrow="20" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF142957FD38FBEB" box="[159,706,1061,1083]" gridcol="0" gridrow="20" pageId="7" pageNumber="538">Caudal-fin length</th>
<td id="D84C425F74170057FD5D2957FCD5FBEB" box="[726,815,1061,1083]" gridcol="1" gridrow="20" pageId="7" pageNumber="538">11.0</td>
<td id="D84C425F74170057FC1F2957FBFDFBEB" box="[916,1031,1061,1083]" gridcol="2" gridrow="20" pageId="7" pageNumber="538">9.7</td>
<td id="D84C425F74170057FB862957FB94FBEB" box="[1037,1134,1061,1083]" gridcol="3" gridrow="20" pageId="7" pageNumber="538">8.2</td>
<td id="D84C425F74170057FB2E2957FB2BFBEB" box="[1189,1233,1061,1083]" gridcol="4" gridrow="20" pageId="7" pageNumber="538">10.4</td>
<td id="D84C425F74170057FA932957FAC2FBEB" box="[1304,1336,1061,1083]" gridcol="5" gridrow="20" pageId="7" pageNumber="538">0.7</td>
</tr>
<tr id="9B9D2B2374170057FF14293CFAC2FBB4" box="[159,1336,1102,1124]" gridrow="21" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF14293CFD38FBB4" box="[159,706,1102,1124]" gridcol="0" gridrow="21" pageId="7" pageNumber="538">Maximum caudal-fin depth</th>
<td id="D84C425F74170057FD5D293CFCD5FBB4" box="[726,815,1102,1124]" gridcol="1" gridrow="21" pageId="7" pageNumber="538">5.6</td>
<td id="D84C425F74170057FC1F293CFBFDFBB4" box="[916,1031,1102,1124]" gridcol="2" gridrow="21" pageId="7" pageNumber="538">5.0</td>
<td id="D84C425F74170057FB86293CFB94FBB4" box="[1037,1134,1102,1124]" gridcol="3" gridrow="21" pageId="7" pageNumber="538">4.5</td>
<td id="D84C425F74170057FB2E293CFB2BFBB4" box="[1189,1233,1102,1124]" gridcol="4" gridrow="21" pageId="7" pageNumber="538">5.8</td>
<td id="D84C425F74170057FA93293CFAC2FBB4" box="[1304,1336,1102,1124]" gridcol="5" gridrow="21" pageId="7" pageNumber="538">0.4</td>
</tr>
<tr id="9B9D2B2374170057FF14290AFAC2FB5E" box="[159,1336,1144,1166]" gridrow="22" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF14290AFD38FB5E" box="[159,706,1144,1166]" gridcol="0" gridrow="22" pageId="7" pageNumber="538">Tail depth at junction of second dorsal and caudal fins</th>
<td id="D84C425F74170057FD5D290AFCD5FB5E" box="[726,815,1144,1166]" gridcol="1" gridrow="22" pageId="7" pageNumber="538">3.1</td>
<td id="D84C425F74170057FC1F290AFBFDFB5E" box="[916,1031,1144,1166]" gridcol="2" gridrow="22" pageId="7" pageNumber="538">2.9</td>
<td id="D84C425F74170057FB86290AFB94FB5E" box="[1037,1134,1144,1166]" gridcol="3" gridrow="22" pageId="7" pageNumber="538">2.7</td>
<td id="D84C425F74170057FB2E290AFB2BFB5E" box="[1189,1233,1144,1166]" gridcol="4" gridrow="22" pageId="7" pageNumber="538">3.3</td>
<td id="D84C425F74170057FA93290AFAC2FB5E" box="[1304,1336,1144,1166]" gridcol="5" gridrow="22" pageId="7" pageNumber="538">0.2</td>
</tr>
<tr id="9B9D2B2374170057FF1429D3FAC2FB67" box="[159,1336,1185,1207]" gridrow="23" pageId="7" pageNumber="538">
<th id="D84C425F74170057FF1429D3FD38FB67" box="[159,706,1185,1207]" gridcol="0" gridrow="23" pageId="7" pageNumber="538">Tail depth at maximum caudal-fin depth</th>
<td id="D84C425F74170057FD5D29D3FCD5FB67" box="[726,815,1185,1207]" gridcol="1" gridrow="23" pageId="7" pageNumber="538">1.5</td>
<td id="D84C425F74170057FC1F29D3FBFDFB67" box="[916,1031,1185,1207]" gridcol="2" gridrow="23" pageId="7" pageNumber="538">1.3</td>
<td id="D84C425F74170057FB8629D3FB94FB67" box="[1037,1134,1185,1207]" gridcol="3" gridrow="23" pageId="7" pageNumber="538">1.0</td>
<td id="D84C425F74170057FB2E29D3FB2BFB67" box="[1189,1233,1185,1207]" gridcol="4" gridrow="23" pageId="7" pageNumber="538">1.8</td>
<td id="D84C425F74170057FA9329D3FAC2FB67" box="[1304,1336,1185,1207]" gridcol="5" gridrow="23" pageId="7" pageNumber="538">0.2</td>
</tr>
</table>
</paragraph>
<caption id="71D279E97417FFAFFF1C2A5CFE0BF8B3" httpUri="https://zenodo.org/record/803772/files/figure.png" pageId="7" pageNumber="538" targetBox="[151,1436,1282,1816]" targetPageId="7">
<paragraph id="251229617417FFAFFF1C2A5CFE0BF8B3" blockId="7.[151,1436,1838,1891]" pageId="7" pageNumber="538">
<emphasis id="17D9F5737417FFAFFF1C2A5CFEE8F893" bold="true" box="[151,274,1838,1860]" pageId="7" pageNumber="538">FIGURE 8.</emphasis>
<taxonomicName id="E2AD52E27417FFAFFE912A5DFE33F894" box="[282,457,1838,1860]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737417FFAFFE912A5DFE33F894" box="[282,457,1838,1860]" italics="true" pageId="7" pageNumber="538">Lampetra soljani</emphasis>
</taxonomicName>
, FSJF 3650, paratypes, about 120 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16–17 Jan. 2015.
</paragraph>
</caption>
<paragraph id="251229617417FFA0FF4C2AF8FE83FF28" blockId="7.[151,1436,1930,2027]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538">
<emphasis id="17D9F5737417FFAFFF4C2AF8FEB5F873" bold="true" box="[199,335,1930,1955]" pageId="7" pageNumber="538">Coloration.</emphasis>
Ammocoetes (
<figureCitation id="BD9635E47417FFAFFE732AF8FDC1F873" box="[504,571,1930,1955]" captionStart="FIGURE 7" captionStartId="6.[151,249,783,805]" captionTargetBox="[321,1266,573,761]" captionTargetId="figure@6.[321,1266,573,761]" captionTargetPageId="6" captionText="FIGURE 7. Lampetra soljani, not preserved, ammocoetes, about 130 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 17 Sep. 2014.." httpUri="https://zenodo.org/record/803770/files/figure.png" pageId="7" pageNumber="538">Fig. 7</figureCitation>
) are brownish along back with lighter undersides and a marmorate pattern on flank. Tail dark brown. Fins hyaline or pale yellowish-brown. Area around gill openings as well as in front of them reddish from blood vessels. Adults with brown back and upper flank, marmorate in some individuals, belly and lower flank silvery, yellowish in preserved individuals. All fins yellowish. A dark grey blotch near apex of second dorsal fin in some individuals (
<figureCitation id="BD9635E47418FFA0FD922DCEFD77FF05" box="[537,653,188,213]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="5.[151,249,1228,1250]" captionStartId-1="5.[151,249,1675,1697]" captionStartId-2="5.[151,249,1977,1999]" captionTargetBox-0="[151,1436,464,1206]" captionTargetBox-1="[180,1392,1352,1637]" captionTargetId-0="figure@5.[151,1436,464,1206]" captionTargetId-1="figure@5.[151,1436,1330,1654]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 3. Lampetra soljani, from the top, ZFMK-ICH 103666, holotype, 109 mm TL; FSJF 3650, paratypes, 104 mm TL, 102 mm TL, 117 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-1="FIGURE 4. Lampetra soljani, FSJF 3650, paratypes, pre-spawning adults, from the top, 124 mm TL, 128 mm TL; Bosnia and Herzegovina: side-arm of Neretva River east of Čapljina; 16 – 17 Jan. 2015." captionText-2="FIGURE 5. Lampetra soljani, FSJF 2184, paratype, freshly metamorphosed adult, 110 mm TL, Croatia: Norin at Konoba Vrilo; 21 Sept. 2006." httpUri-0="https://zenodo.org/record/803764/files/figure.png" httpUri-1="https://zenodo.org/record/803766/files/figure.png" pageId="8" pageNumber="539">Figs. 3–5</figureCitation>
). Lateral-line neuromasts unpigmented on ventral surface and in prebranchial region.
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7418FFA0FF4C2C76FF1AFACE" pageId="8" pageNumber="539" type="distribution">
<paragraph id="251229617418FFA0FF4C2C76FC5BFE58" blockId="8.[151,1436,151,392]" pageId="8" pageNumber="539">
<emphasis id="17D9F5737418FFA0FF4C2C76FE9AFECD" bold="true" box="[199,352,260,285]" pageId="8" pageNumber="539">Distribution.</emphasis>
<taxonomicName id="E2AD52E27418FFA0FEE42C77FDC2FECC" box="[367,568,260,284]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737418FFA0FEE42C77FDC2FECC" box="[367,568,260,284]" italics="true" pageId="8" pageNumber="539">Lampetra soljani</emphasis>
</taxonomicName>
is only known from the Neretva drainage in 
<collectingCountry id="5DBA69F17418FFA0FBF72C76FB35FECD" box="[1148,1231,260,285]" name="Croatia" pageId="8" pageNumber="539">Croatia</collectingCountry>
and 
<collectingCountry id="5DBA69F17418FFA0FA9D2C76FED0FE90" name="Bosnia and Herzegovina" pageId="8" pageNumber="539">Bosnia and Herzegovina</collectingCountry>
. Lamprey from the Morača River drainage including Zeta River are preliminary identified as 
<taxonomicName id="E2AD52E27418FFA0FA0E2C5BFF19FEB4" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737418FFA0FA0E2C5BFF19FEB4" italics="true" pageId="8" pageNumber="539">L. soljani</emphasis>
</taxonomicName>
based on molecular characters. The Morača River flows to Lake Skadar in 
<collectingCountry id="5DBA69F17418FFA0FBA22C3FFB4FFEB5" box="[1065,1205,333,357]" name="Serbia" pageId="8" pageNumber="539">Montenegro</collectingCountry>
. The actually known distribution area is shown in 
<figureCitation id="BD9635E47418FFA0FE502C1DFDE6FE58" box="[475,540,367,392]" captionStart="FIGURE 9" captionStartId="8.[151,250,1111,1133]" captionTargetBox="[151,1436,445,1089]" captionTargetId="figure@8.[151,1436,445,1089]" captionTargetPageId="8" captionText="FIGURE 9. Records of L. soljani. Numbers correspond to locality data given in Table 3." httpUri="https://zenodo.org/record/803774/files/figure.png" pageId="8" pageNumber="539">Fig. 9</figureCitation>
based on records listed in 
<tableCitation id="682F1CDA7418FFA0FCCC2C1DFC66FE58" box="[839,924,367,392]" captionStart="TABLE 3" captionStartId="9.[151,239,292,315]" captionText="TABLE 3. Findings of L. soljani. The locality numbers correspond to numbers in Fig. 9." pageId="8" pageNumber="539">Table 3</tableCitation>
.
</paragraph>
<caption id="71D279E97418FFA0FF1C2925FBE0FBBD" box="[151,1050,1111,1133]" httpUri="https://zenodo.org/record/803774/files/figure.png" pageId="8" pageNumber="539" targetBox="[151,1436,445,1089]" targetPageId="8">
<paragraph id="251229617418FFA0FF1C2925FBE0FBBD" blockId="8.[151,1050,1111,1133]" box="[151,1050,1111,1133]" pageId="8" pageNumber="539">
<emphasis id="17D9F5737418FFA0FF1C2925FEE8FBBC" bold="true" box="[151,274,1111,1133]" pageId="8" pageNumber="539">FIGURE 9.</emphasis>
Records of 
<taxonomicName id="E2AD52E27418FFA0FE06292AFE17FBBD" box="[397,493,1111,1133]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737418FFA0FE06292AFE17FBBD" box="[397,493,1111,1133]" italics="true" pageId="8" pageNumber="539">L. soljani</emphasis>
</taxonomicName>
. Numbers correspond to locality data given in Table 3.
</paragraph>
</caption>
<paragraph id="251229617418FFA0FF4C29EBFF1AFACE" blockId="8.[151,1437,1177,2031]" pageId="8" pageNumber="539">
<emphasis id="17D9F5737418FFA0FF4C29EBFE44FB62" bold="true" box="[199,446,1177,1202]" pageId="8" pageNumber="539">Habitat and biology.</emphasis>
Ammocoetes are found in silt, muddy or fine sandy-muddy sediment in river sections with slow or no current, in shallows at banks or backwaters at water depth of about 
<quantity id="E25584847418FFA0FB1E29CFFAF7FB06" box="[1173,1293,1213,1238]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="8" pageNumber="539" unit="cm" value="30.0" valueMax="50.0" valueMin="10.0">10–50 cm</quantity>
. Adults and ammocoetes are commonly caught together. First post-metamorphosis individual was found in September (FSJF 2184).
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7418FFA0FF4C285BFBB6FAB6" pageId="8" pageNumber="539" type="etymology">
<paragraph id="251229617418FFA0FF4C285BFBB6FAB6" blockId="8.[151,1437,1177,2031]" pageId="8" pageNumber="539">
<emphasis id="17D9F5737418FFA0FF4C285BFEB4FA92" bold="true" box="[199,334,1321,1346]" pageId="8" pageNumber="539">Etymology.</emphasis>
Named in honour of Tonko Šoljan (
<date id="51130FA17418FFA0FC9E2858FC6CFA92" box="[789,918,1322,1346]" pageId="8" pageNumber="539" value="1907" valueMax="1980">1907–1980</date>
) in appreciation of his contribution to the knowledge and development of ichthyology in 
<collectingCountry id="5DBA69F17418FFA0FD23283FFD00FAB6" box="[680,762,1357,1382]" name="Croatia" pageId="8" pageNumber="539">Croatia</collectingCountry>
and 
<collectingCountry id="5DBA69F17418FFA0FCBB283FFBB2FAB6" box="[816,1096,1357,1382]" name="Bosnia and Herzegovina" pageId="8" pageNumber="539">Bosnia and Herzegovina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6DB77AEA7418FFA2FF4C2803FBA6FE90" lastPageId="10" lastPageNumber="541" pageId="8" pageNumber="539" type="discussion">
<paragraph id="251229617418FFA0FF4C2803FE56FA26" blockId="8.[151,1437,1177,2031]" pageId="8" pageNumber="539">
<emphasis id="17D9F5737418FFA0FF4C2803FEC1FA5A" bold="true" box="[199,315,1393,1418]" pageId="8" pageNumber="539">Remarks.</emphasis>
Lampreys from the Morača River drainage are identified as 
<taxonomicName id="E2AD52E27418FFA0FB9F2800FB78FA5A" box="[1044,1154,1394,1418]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737418FFA0FB9F2800FB78FA5A" box="[1044,1154,1394,1418]" italics="true" pageId="8" pageNumber="539">L. soljani</emphasis>
</taxonomicName>
based on the presented molecular characters. No adult individuals were available to us to test, if these agree with the diagnostic characters given here. Therefore, the identification is preliminary and should be tested in the future, when well-preserved adults become available.
</paragraph>
<paragraph id="251229617418FFA0FF4C2B73FAABF9B1" blockId="8.[151,1437,1177,2031]" pageId="8" pageNumber="539">
Posterial teeth are often deeply embedded in brook lampreys and we were unable to x-ray the mental disc in 
<emphasis id="17D9F5737418FFA0FA0E2B70FF11F9EE" italics="true" pageId="8" pageNumber="539">
<taxonomicName id="E2AD52E27418FFA0FA0E2B70FF1DF9EE" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="soljani">L. soljani</taxonomicName>
.
</emphasis>
Due to the availability of so few adult individuals, we also did not apply any clearing and staining methods. Therefore, we cannot full exclude, that deeply embedded posterials, not visible externally, might be present.
</paragraph>
<paragraph id="251229617418FFA1FF4C2B1DFB96FF28" blockId="8.[151,1437,1177,2031]" lastBlockId="9.[151,1436,151,248]" lastPageId="9" lastPageNumber="540" pageId="8" pageNumber="539">
Lampreys are still a challenge for taxonomists and a good example for cryptic species exhibiting only few characters that can be reliably measured or counted (no fin rays, no scales, no ossified structures). Furthermore, lampreys show strong allometric growth after metamorphosis (
<bibRefCitation id="413C54907418FFA0FCD32BC4FE34F922" author="Krappe" pageId="8" pageNumber="539" refString="Krappe, M. (2004) Quantitative Analysen populationsbiologischer Phanomene im Lebenszyklus des Bachneunauges Lampetra planeri (Bloch 1784). Inaugural-Dissertation, Universitat Rostock, 241 + XXXIII pp." type="book" year="2004">
Krappe 2004; see also differences in size and shape of body and fins in 
<figureCitation id="BD9635E47418FFA0FEE42BABFE55F922" box="[367,431,1753,1778]" captionStart="FIGURE 5" captionStartId="5.[151,249,1977,1999]" captionText="FIGURE 5. Lampetra soljani, FSJF 2184, paratype, freshly metamorphosed adult, 110 mm TL, Croatia: Norin at Konoba Vrilo; 21 Sept. 2006." pageId="8" pageNumber="539">Fig. 5</figureCitation>
+6
</bibRefCitation>
) limiting morphometric studies to individuals in exactly the same phase of life. 
<bibRefCitation id="413C54907418FFA0FAC12BA8FF1BF8C6" author="Krappe" pageId="8" pageNumber="539" refString="Krappe, M. (2004) Quantitative Analysen populationsbiologischer Phanomene im Lebenszyklus des Bachneunauges Lampetra planeri (Bloch 1784). Inaugural-Dissertation, Universitat Rostock, 241 + XXXIII pp." type="book" year="2004">Krappe (2004)</bibRefCitation>
report growth of the disk and shrinkage of the body and tail between 5-15% after metamorphosis for different body proportions in 
<taxonomicName id="E2AD52E27418FFA0FE6D2A50FDAFF8EA" box="[486,597,1826,1850]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="planeri">
<emphasis id="17D9F5737418FFA0FE6D2A50FDAFF8EA" box="[486,597,1826,1850]" italics="true" pageId="8" pageNumber="539">L. planeri</emphasis>
</taxonomicName>
. Therefore, the traditional taxonomy in lampreys is mainly based on their dentition and the number of myomeres. Given that brook lampreys do not feed as adults, and hypothesized that they would use their teeth similarly to their predatory congeners, it can be speculated that the variation in tooth formulae and dentition might be a result of relaxed selection pressure. However, this hypothesis deserves a thorough testing and is far beyond the aims of our study. All brook lampreys have less and smaller teeth compared to their predatory congeners (
<bibRefCitation id="413C54907418FFA0FE7F2AA7FD6BF83E" author="Renaud" box="[500,657,2005,2030]" pageId="8" pageNumber="539" refString="Renaud, C. B. (2011) Lampreys of the World. An annotated and illustrated catalogue of lamprey species known to date. FAO Species Catalogue for Fishery Purposes, No. 5, i - vi + 1 - 109." type="book chapter" year="2011">Renaud 2011</bibRefCitation>
; 
<bibRefCitation id="413C54907418FFA0FD292AA7FC31F83E" author="Kottelat" box="[674,971,2005,2030]" pageId="8" pageNumber="539" refString="Kottelat, M. &amp; Freyhof, J. (2007) Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin, 646 pp." type="book" year="2007">Kottelat &amp; Freyhof 2007</bibRefCitation>
). Potentially, all non-feeding species of 
<taxonomicName id="E2AD52E27419FFA1FF1C2DEBFEFEFF60" box="[151,260,153,176]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="9" pageNumber="540" phylum="Chordata" rank="genus">
<emphasis id="17D9F5737419FFA1FF1C2DEBFEFEFF60" box="[151,260,153,176]" italics="true" pageId="9" pageNumber="540">Lampetra</emphasis>
</taxonomicName>
are satellite species of 
<taxonomicName id="E2AD52E27419FFA1FD852DEBFD6CFF60" box="[526,662,152,176]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="fluviatilis">
<emphasis id="17D9F5737419FFA1FD852DEBFD6CFF60" box="[526,662,152,176]" italics="true" pageId="9" pageNumber="540">L. fluviatilis</emphasis>
</taxonomicName>
and have been isolated at different time periods from this predatory lamprey. It is likely, that all brook lampreys reduce the teeth and the degree of reduction might be related to the time, brook lamprey species or populations are isolated from their predatory congeners.
</paragraph>
<caption id="71D279E97419FFA1FF1C2C56FBBDFEEB" box="[151,1095,291,315]" pageId="9" pageNumber="540">
<paragraph id="251229617419FFA1FF1C2C56FBBDFEEB" blockId="9.[151,1095,291,315]" box="[151,1095,291,315]" pageId="9" pageNumber="540">
<emphasis id="17D9F5737419FFA1FF1C2C56FEF9FEEB" bold="true" box="[151,259,292,315]" pageId="9" pageNumber="540">TABLE 3</emphasis>
. Findings of 
<taxonomicName id="E2AD52E27419FFA1FE1A2C56FE0CFEEB" box="[401,502,291,315]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F5737419FFA1FE1A2C56FE0CFEEB" box="[401,502,291,315]" italics="true" pageId="9" pageNumber="540">L. soljani</emphasis>
</taxonomicName>
. The locality numbers correspond to numbers in Fig. 9.
</paragraph>
</caption>
<paragraph id="251229617419FFA2FF142C25FBA6FE90" blockId="9.[159,1088,342,364]" lastBlockId="10.[151,1436,151,320]" lastPageId="10" lastPageNumber="541" pageId="9" pageNumber="540">
Nr. Country Drainage Locality Coordinates Source With regard to the paucity of morphological characters compared to bony fishes, the inclusion of molecular genetic characters into standard taxonomic works in lampreys is of great value. A comprehensive, genome wide comparison of a predatory/non-feeding species pair from the same location (
<taxonomicName id="E2AD52E27419FFA1FC7F2BB0FB87F909" box="[1012,1149,1729,1753]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="fluviatilis">
<emphasis id="17D9F5737419FFA1FC7F2BB0FB87F909" box="[1012,1149,1729,1753]" italics="true" pageId="9" pageNumber="540">L. fluviatilis</emphasis>
</taxonomicName>
with 
<taxonomicName id="E2AD52E27419FFA1FB362BB0FAD6F909" box="[1213,1324,1729,1753]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="planeri">
<emphasis id="17D9F5737419FFA1FB362BB0FAD6F909" box="[1213,1324,1729,1753]" italics="true" pageId="9" pageNumber="540">L. planeri</emphasis>
</taxonomicName>
) revealed strong genetic differentiation, despite the existence of shared mitochondrial haplotypes in these two particular populations (
<bibRefCitation id="413C54907419FFA1FEA52A78FDD9F8F2" author="Mateus" box="[302,547,1801,1826]" pageId="9" pageNumber="540" refString="Mateus, C. S., Stange, M., Berner, D., Roesti, M., Quintella, B. R., Alves, M. J., Almeida, P. J. &amp; Salzburger, W. (2013 b) Strong genome-wide divergence between sympatric European river and brook lampreys. Current biology, 23 (15), R 649 - R 650. https: // doi. org / 10.1016 / j. cub. 2013.06.026" type="journal article" year="2013" yearSuffix="b">
Mateus 
<emphasis id="17D9F5737419FFA1FE052A78FE3FF8F1" box="[398,453,1801,1825]" italics="true" pageId="9" pageNumber="540">et al</emphasis>
. 2013b
</bibRefCitation>
). In line with these findings, Rougemont 
<emphasis id="17D9F5737419FFA1FB922A78FBB4F8F1" box="[1049,1102,1801,1825]" italics="true" pageId="9" pageNumber="540">et al</emphasis>
. recently (2017) used RAD sequencing for a genome wide screening of SNP in nine different population species pairs in France and demonstrated again a strong genomic differentiation between the two species, but also showed levels of ongoing gene flow to correlate with geographic connectivity. All known brook lampreys included herein show differences in their mitochondrial DNA sequences between them (COI - this study; control region – 
<bibRefCitation id="413C54907419FFA1FBFD2AEBFABDF862" author="Pereira" box="[1142,1351,1945,1970]" pageId="9" pageNumber="540" refString="Pereira, A. M., Robalo, J. I., Freyhof, J., Maia, C., Fonseca, J. P., Valente, A. &amp; Almada, V. C. (2010) Phylogeographical analysis reveals multiple conservation units in brook lampreys of Portuguese streams. Journal of Fish Biology, 77, 361 - 371. https: // doi. org / 10.1111 / j. 1095 - 8649.2010.02675. x" type="journal article" year="2010">
Pereira 
<emphasis id="17D9F5737419FFA1FB472AE8FAFFF861" box="[1228,1285,1945,1969]" italics="true" pageId="9" pageNumber="540">et al.</emphasis>
2010
</bibRefCitation>
; cytB – 
<bibRefCitation id="413C54907419FFA1FF1C2ACCFE83F806" author="Mateus" box="[151,377,1981,2006]" pageId="9" pageNumber="540" refString="Mateus, C. S., Alves, M. J., Quintella, B. R. &amp; Almeida, P. R. (2013 a) Three new cryptic species of the lamprey genus Lampetra Bonnaterre, 1788 (Petromyzontiformes: Petromyzontidae) from the Iberian Peninsula. Contributions to Zoology, 82, 37 - 53." type="journal article" year="2013" yearSuffix="a">
Mateus 
<emphasis id="17D9F5737419FFA1FF7A2ACCFED3F805" box="[241,297,1981,2005]" italics="true" pageId="9" pageNumber="540">et al.</emphasis>
2013a
</bibRefCitation>
), but no study exists that included the potential predatory sister taxon of the Eastern European species, so we do not know if the genome wide strong differentiation documented in 
<bibRefCitation id="413C5490741AFFA2FBF92DEAFA90FF60" box="[1138,1386,151,176]" pageId="10" pageNumber="541" refString="Mateus, C. S., Stange, M., Berner, D., Roesti, M., Quintella, B. R., Alves, M. J., Almeida, P. J. &amp; Salzburger, W. (2013 b) Strong genome-wide divergence between sympatric European river and brook lampreys. Current biology, 23 (15), R 649 - R 650. https: // doi. org / 10.1016 / j. cub. 2013.06.026" type="journal article">
Mateus 
<emphasis id="17D9F573741AFFA2FB462DEBFAFAFF60" box="[1229,1280,152,176]" italics="true" pageId="10" pageNumber="541">et al</emphasis>
. (2013b)
</bibRefCitation>
and Rougemont et al. (2017) is the rule or an exception. Interestingly, while our samples of 
<taxonomicName id="E2AD52E2741AFFA2FB132DCFFAFEFF04" box="[1176,1284,188,212]" class="Cephalaspidomorphi" family="Petromyzontidae" genus="Lampetra" kingdom="Animalia" order="Petromyzontiformes" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="soljani">
<emphasis id="17D9F573741AFFA2FB132DCFFAFEFF04" box="[1176,1284,188,212]" italics="true" pageId="10" pageNumber="541">L. soljani</emphasis>
</taxonomicName>
show only a single, potentially diagnostic nucleotide substitution (position 460 C vs T) in their DNA barcode region, the four individuals analysed from the Morača River drainage exhibit two additional diagnostic positions, which are unique among all included specimens and can be perceived in the haplotype network (
<figureCitation id="BD9635E4741AFFA2FB822C55FBB5FE90" box="[1033,1103,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1569,1591]" captionTargetBox="[176,1410,194,1526]" captionTargetId="figure@4.[175,1411,193,1527]" captionTargetPageId="4" captionText="FIGURE 2. Median-joining network of the COI haplotypes. Circle size corresponds to sample size; one bar indicates an additional mutational step. Small black circles represent median vectors, i. e., hypothetical, reconstructed haplotypes." httpUri="https://zenodo.org/record/803762/files/figure.png" pageId="10" pageNumber="541">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>