treatments-xml/data/84/23/1A/84231AB179B69C69D9FD0A15E0CB37F7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.505.9862" ID-GBIF-Dataset="2e13b190-dff9-41bd-b2db-d6f2b4c82731" ID-PMC="PMC4453233" ID-Pensoft-Pub="1313-2970-505-1" ID-PubMed="26052236" ID-ZBK="7F1C641D389940BD8E9B1F812D4509D1" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-505-1" ModsDocOrigin="ZooKeys 505" ModsDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)" checkinTime="1451244309637" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauries, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2015" docId="84231AB179B69C69D9FD0A15E0CB37F7" docLanguage="en" docName="ZooKeys 505: 1-34" docOrigin="ZooKeys 505" docSource="http://dx.doi.org/10.3897/zookeys.505.9862" docTitle="Eutrichodesmus tenuis Golovatch, Geoffroy, Mauries &amp; VandenSpiegel, sp. n." docType="treatment" docUuid="B7247170-99F8-4A4F-AED5-440344A4E881" docUuidSource="ZooBank" docVersion="4" lastPageNumber="6" masterDocId="FFCBFF9AFFE6FFEDFFA5644EFF808470" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="6" updateTime="1668160367798" updateUser="ExternalLinkService">
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<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mauries, Jean-Paul</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>VandenSpiegel, Didier</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
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<mods:number>505</mods:number>
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<mods:start>1</mods:start>
<mods:end>34</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.505.9862</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.505.9862</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-505-1</mods:identifier>
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<treatment ID-GBIF-Taxon="152060115" LSID="urn:lsid:zoobank.org:act:B7247170-99F8-4A4F-AED5-440344A4E881" httpUri="http://treatment.plazi.org/id/84231AB179B69C69D9FD0A15E0CB37F7" lastPageNumber="6" pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">Taxon classification Animalia Polydesmida Haplodesmidae</paragraph>
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<subSubSection pageId="5" pageNumber="6" type="nomenclature">
<paragraph pageId="5" pageNumber="6">
<taxonomicName LSID="http://zoobank.org/B7247170-99F8-4A4F-AED5-440344A4E881" authority="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus tenuis" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tenuis">
Eutrichodesmus tenuis Golovatch, Geoffroy, 
<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
&amp; VandenSpiegel
</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
Figs 5, 6
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="type material">
<paragraph pageId="5" pageNumber="6">Type material.</paragraph>
<paragraph pageId="5" pageNumber="6">Holotype ♂ (MNHN JC 369), China, Guizhou Prov., Guanling County, Yong Ning Town, Cave Yun Dong (Cloud Cave), 01.VIII.2005, leg. L. Latella &amp; D. Avesani.</paragraph>
<paragraph pageId="5" pageNumber="6">Paratypes: 1 ♀, 2 subadult ♀ (MNHN JC 369), 1 subadult ♀ (SEM), same data, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="name">
<paragraph pageId="5" pageNumber="6">Name.</paragraph>
<paragraph pageId="5" pageNumber="6">To emphasize the relatively slender body due to subvertical paraterga; adjective.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="6">Differs from congeners by the large body size, clearly elevated mid-dorsal regions of most metaterga, coupled with narrow, strongly declivous, subvertical paraterga and a simple, falcate gonopod telopodite carrying a long, spiniform, distofemoral process (see also Key below).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="description">
<paragraph pageId="5" pageNumber="6">Description.</paragraph>
<paragraph pageId="5" pageNumber="6">Length of adults ca 14-15 mm, width 1.8 and 2.5 mm (♂ holotype) or 1.6 and 2.1 mm (♀ paratype) on midbody pro- and metazonae, respectively. Coloration entirely pallid, sometimes (♀ paratype) with traces of reddish earth material on metaterga.</paragraph>
<paragraph pageId="5" pageNumber="6">
All characters as in 
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus triangularis" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="triangularis">Eutrichodesmus triangularis</taxonomicName>
sp. n., except as follows.
</paragraph>
<paragraph pageId="5" pageNumber="6">
Body with 20 segments (♂, ♀), conglobation pattern typical of 
<normalizedToken originalValue="“doratodesmoids”">&quot;doratodesmoids&quot;</normalizedToken>
, volvation apparently being complete because of particularly strongly declivous and short paraterga. Antennae short and clavate (Fig. 5H, K). Collum not covering the head from above, fore margin clearly lobulate and slightly elevated, with 4-5 transverse rows of small, but evident tuberculations, only frontal- and caudalmost rows being regular. Metaterga behind collum with three transverse, rather irregular, mixostictic rows of similarly evident, rounded, setigerous tuberculations extending onto paraterga, usually about 10-11+10-11 per row (Fig. 5 
<normalizedToken originalValue="A–G">A-G</normalizedToken>
); starting with midbody segments, middle rows clearly enlarged and elevated mid-dorsad, increasingly clearly so towards segment 18 as well (Fig. 5 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
); a few caudomarginal lobulations evident only on paraterga (Fig. 5 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
); limbus microcrenulate (Fig. 5N). Paraterga with evident shoulders anteriorly, very strongly declivous, subvertical, directed ventrolaterad at about 75-80° to even more strongly declined sides above paraterga (Fig. 5L), broad, tips lying clearly below level of venter, usually rather vaguely tri- or quadrilobate laterally, without anterolaterals (Fig. 5 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
, M). Paraterga 2 strongly enlarged, directed ventrad (Fig. 5A, B, E, I), lateral margin broadly rounded, with numerous, very small lobulations, caudal margin with a few lobes located near schism, both schism and hyposchism being small (Fig. 5B). Tergal setae very short, 2-segmented, apical part usually phylloid (Fig. 5P, Q). Pore formula normal, ozopores distinct, located dorsally on small porosteles in posterior 1/3 of paratergite and well removed from lateral margin (Fig. 5 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
, M). Epiproct finger-shaped, densely tuberculate (Fig. 5D, G, J). Hypoproct subtrapeziform (Fig. 5J).
</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figure 5. 
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus tenuis" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tenuis">Eutrichodesmus tenuis</taxonomicName>
sp. n., subadult ♀ paratype; A habitus, lateral view B, E, H anterior part of body, lateral, dorsal and ventral views, respectively C, F, I midbody segments, lateral, dorsal and ventral views, respectively D, G, J posterior part of body, lateral, dorsal and ventral views, respectively K head, ventral view L cross-section of a midbody segment, caudal view M poriferous midbody paratergite, lateral view N limbus, lateral view O, P, Q tergal setae, various views R midbody leg. Scale bars: 0.5 mm (
<normalizedToken originalValue="A–I">A-I</normalizedToken>
, L), 0.2 mm (J, K), 0.1 mm (M, O, R), 0.02 mm (N, P). 0.01 mm (Q).
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="6">Sterna usually with a rather deep, narrow depression between coxae (Fig. 5I). Legs short and crassate, about half as long as body height (Fig. 5L), all podomeres except tarsi finely micropapillate (Figs 5R, 6A).</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figure 6. 
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus tenuis" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tenuis">Eutrichodesmus tenuis</taxonomicName>
sp. n., ♂ holotype; A leg 9, lateral view B left gonopod, mesal view. Scale bar: 0.2 mm. Designation in text.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="6">Gonopods (Fig. 6B) very simple. Coxae subquadrate, large, micropapillate and rather densely setose on lateral face, with only a small round lobe caudolaterally. Telopodite considerably longer than coxite, suberect, setose over its basal half until base of a prominent, spiniform, simple, distofemoral process (dp), the latter situated at about halfway along telopodite, acropodite strongly falcate, twisted, subacuminate, simple, devoid of outgrowths; seminal groove terminating subapically at base of a hairpad.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="remark">
<paragraph pageId="5" pageNumber="6">Remark.</paragraph>
<paragraph pageId="5" pageNumber="6">
More information on this cave and its fauna can be found in 
<bibRefCitation author="Latella, L" journalOrPublisher="Memorie del Museo Civico di Storia Naturale di Verona (serie 2, Monografie Naturalistiche)" pageId="24" pageNumber="25" pagination="65 - 88" title="Biological investigation of the Museo Civico di Storia Naturale of Verona in South China caves. Research in South China Karst." volume="3" year="2008">Latella and Hu (2008)</bibRefCitation>
and in 
<bibRefCitation author="Latella, L" journalOrPublisher="Memorie del Museo Civico di Storia Naturale di Verona (serie 2, Monografie Naturalistiche)" pageId="24" pageNumber="25" pagination="25 - 64" title="Caves explored during the scientific expeditions to China of the Museo Civico di Storia Naturale of Verona. Research in South China Karst." volume="3" year="2008">Latella and Zorzin (2008)</bibRefCitation>
.
</paragraph>
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</treatment>
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