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<mods:title id="F092F8553B44163A0584693641B7D8C5">A monograph of the Oligochaete family Alluroididae</mods:title>
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<mods:namePart id="6E55F7D13C64F8ACF3BC93ED19E18705">Jamieson, Barrie G. M.</mods:namePart>
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<mods:affiliation id="EA8CB0BFF293B851A49EE18DBAE1B28F">School of the Environment, University of Queensland and 1 Maxwell Court, Coolum Beach, Queensland, Australia 4573.</mods:affiliation>
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<mods:namePart id="411287A3CD80CC1A99CB7F32443A67B3">Fragoso, Carlos</mods:namePart>
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<mods:affiliation id="2413C7BAB032950231CAA9A9154F5084">Red de Bioversidad y Sistmatica, Instituto de Ecologia A. C, Xalapa, Mexico.</mods:affiliation>
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<mods:title id="43CC8B8C0EAAFC7E412413B05C0F57F1">Zootaxa</mods:title>
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<mods:date id="7878AD96A93580EB3CE6ACB5E2E40ECF">2024</mods:date>
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<subSubSection id="A79665027946FFE3CED9FA71FD47F9BC" box="[151,707,1522,1549]" pageId="16" pageNumber="417" type="nomenclature">
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<paragraph id="EF3336897946FFE3CED9FA71FD47F9BC" blockId="16.[151,707,1522,1549]" box="[151,707,1522,1549]" pageId="16" pageNumber="417">
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<heading id="B47B81E57946FFE3CED9FA71FD47F9BC" bold="true" box="[151,707,1522,1549]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
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<emphasis id="DDF8EA9B7946FFE3CED9FA71FD47F9BC" bold="true" box="[151,707,1522,1549]" pageId="16" pageNumber="417">
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<taxonomicName id="288C4D0A7946FFE3CED9FA71FDB9F9BC" ID-CoL="C2D4" authority="Beddard 1894" authorityName="Beddard" authorityYear="1894" box="[151,573,1522,1549]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="16" pageNumber="417" phylum="Annelida" rank="species" species="pordagei">
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<emphasis id="DDF8EA9B7946FFE3CED9FA71FE02F9BD" bold="true" box="[151,390,1522,1549]" italics="true" pageId="16" pageNumber="417">Alluroides pordagei</emphasis>
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<bibRefCitation id="8B1D4B787946FFE3CFC3FA71FDB9F9BC" author="Beddard, F. E." box="[397,573,1522,1548]" pageId="16" pageNumber="417" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard 1894</bibRefCitation>
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</taxonomicName>
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(see
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<figureCitation id="77B72A0C7946FFE3CC34FA71FD3DF9BC" box="[634,697,1522,1548]" captionStart="FIGURE 8" captionStartId="17.[151,250,964,988]" captionTargetBox="[185,1401,247,940]" captionTargetId="figure-46@17.[151,1436,234,941]" captionTargetPageId="17" captionText="FIGURE 8. Alluroides pordagei Beddard 1894. A. Longitudinal section of the anterior region of the body of Beddard’s paratype. B. Spermatheca. C. Longitudinal section of wall of atrium. D. Transverse section of atrium.Abbreviations: at, atrium; at ch, atrial chamber; br, brain; cil, cilia; ep, epidermis; epi, epithelium; gl duc, glandular ductule(s); gl sh, glandular sheath; int, intestine; lum, lumen; m, mouth; ph m, pharyngeal musculature; per, peritoneum; sep.gl, septal gland; sp, spermatheca; sperm, spermatozoa; sph, sphincter; sp epi, spermathecal epithelium; sp m sh, muscular sheath of spermatheca; v.d, vas deferens.. Slightly modified from Jamieson (1968)." figureDoi="http://doi.org/10.5281/zenodo.14022647" httpUri="https://zenodo.org/record/14022647/files/figure.png" pageId="16" pageNumber="417">Fig 8</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A79665027946FFE3CED9F9B4FD8AF958" pageId="16" pageNumber="417" type="reference_group">
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<paragraph id="EF3336897946FFE3CED9F9B4FD13F9FF" blockId="16.[151,1437,1591,1768]" box="[151,663,1591,1615]" pageId="16" pageNumber="417">
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<treatmentCitationGroup id="CF9C11A77946FFE3CED9F9B4FD13F9FF" box="[151,663,1591,1615]" pageId="16" pageNumber="417">
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<treatmentCitation id="6E2D10987946FFE3CED9F9B4FDB6F9FF" author="Beddard, F. E." box="[151,562,1591,1615]" page="244" pageId="16" pageNumber="417" year="1894">
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<taxonomicName id="288C4D0A7946FFE3CED9F9B4FDB6F9FF" ID-CoL="C2D4" authority="Beddard, 1894: 244" authorityName="Beddard" authorityPageNumber="244" authorityYear="1894" box="[151,562,1591,1615]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="16" pageNumber="417" phylum="Annelida" rank="species" species="pordagei">
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<emphasis id="DDF8EA9B7946FFE3CED9F9B4FEE5F9FF" box="[151,353,1591,1615]" italics="true" pageId="16" pageNumber="417">Alluroides pordagei</emphasis>
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<bibRefCitation id="8B1D4B787946FFE3CF29F9B4FDB6F9FF" author="Beddard, F. E." box="[359,562,1591,1615]" pageId="16" pageNumber="417" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard, 1894: 244</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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,
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<figureCitation id="77B72A0C7946FFE3CC72F9B4FDFEF9FF" box="[572,634,1591,1615]" captionStart="FIGURE 4" captionStartId="12.[151,250,1906,1930]" captionTargetBox="[242,1363,1542,1866]" captionTargetId="figure-568@12.[215,1371,1530,1885]" captionTargetPageId="12" captionText="FIGURE 4. Alluroides brinkhursti brinkhursti Jamieson (1968). Longitudinal section of the holotype. Abbreviations: br, brain; bv, blood vessel; m, mouth; msh, muscular sheath of atrium; oe, oesophagus; oo, oocyte: os, ovisac; ph m, pharyngeal musculature; pr c, prostatic cells; sep, septum; sep gl, septal gland; sp p, spermathecal pore; s. ves, seminal vesicle; sp, spermatheca; sp f, sperm funnel; tes, testis; vd, vas deferens; v n, ventral nerve cord. Adapted from Jamieson (1968)." figureDoi="http://doi.org/10.5281/zenodo.14022635" httpUri="https://zenodo.org/record/14022635/files/figure.png" pageId="16" pageNumber="417">Fig. 4</figureCitation>
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,
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<figureCitation id="77B72A0C7946FFE3CCCAF9B4FD17F9FF" box="[644,659,1591,1615]" captionStart="FIGURE 5" captionStartId="13.[151,250,1798,1822]" captionTargetBox="[205,1383,202,1774]" captionTargetId="figure-19@13.[205,1383,201,1775]" captionTargetPageId="13" captionText="FIGURE 5. Alluroides brinkhursti brinkhursti Jamieson (1968). A. Transverse section (TS) of clitellum, showing single cell layer; the cells with conspicuous secretory granules and each with a basal nucleus. B. TS through the wall of the atrium, showing a group of atrial gland cells with ductule penetrating the muscular sheath of the atrium. C. Longitudinal section through the male pore, showing the ectal end of the atrium, which forms a penis with muscular sheath, ciliated epithelium and rope of spermatozoa in the lumen, forming in the ectal chamber a sperm mass. D. Alluroides pordagei. Oblique section through the atrial bulb, containing a large sperm mass, and the associated atrium. From Jamieson (2006)." figureDoi="http://doi.org/10.5281/zenodo.14022639" httpUri="https://zenodo.org/record/14022639/files/figure.png" pageId="16" pageNumber="417">5</figureCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="EF3336897946FFE3CED9F9D6FDE5F93C" blockId="16.[151,1437,1591,1768]" pageId="16" pageNumber="417">
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<treatmentCitationGroup id="CF9C11A77946FFE3CED9F9D6FDE5F93C" pageId="16" pageNumber="417">
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<treatmentCitation id="6E2D10987946FFE3CED9F9D6FD64F9DD" author="Michaelsen, W." box="[151,736,1621,1646]" page="107" pageId="16" pageNumber="417" year="1900">
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<taxonomicName id="288C4D0A7946FFE3CED9F9D6FD64F9DD" ID-CoL="C2D4" authority="Beddard, Michaelsen 1900: 106 - 107" authorityName="Beddard, Michaelsen" authorityPageNumber="106 - 107" authorityYear="1900" box="[151,736,1621,1646]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="16" pageNumber="417" phylum="Annelida" rank="species" species="pordagei">
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<emphasis id="DDF8EA9B7946FFE3CED9F9D6FEE5F9DD" box="[151,353,1621,1645]" italics="true" pageId="16" pageNumber="417">Alluroides pordagei</emphasis>
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Beddard,
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<bibRefCitation id="8B1D4B787946FFE3CF87F9D6FD64F9DD" author="Michaelsen, W." box="[457,736,1621,1646]" pageId="16" pageNumber="417" refId="ref23961" refString="Michaelsen, W. (1900) Oligochaeta. Das Tierreich. Vol. 10. Lief. Friedlander und Sohn, Berlin, 575 pp. https: // doi. org / 10.5962 / bhl. title. 11605" type="book" year="1900">Michaelsen 1900: 106–107</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="6E2D10987946FFE3CCA5F9D6FC42F9DD" author="Brinkhurst, R. O." box="[747,966,1621,1645]" page="527" pageId="16" pageNumber="417" year="1964">
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<bibRefCitation id="8B1D4B787946FFE3CCA5F9D6FC42F9DD" author="Brinkhurst, R. O." box="[747,966,1621,1645]" pageId="16" pageNumber="417" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst 1964: 527</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="6E2D10987946FFE3CD9EF9D6FB14F9DD" author="Jamieson, B. G. M." box="[976,1168,1621,1645]" page="75" pageId="16" pageNumber="417" year="1968">
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<bibRefCitation id="8B1D4B787946FFE3CD9EF9D6FB14F9DD" author="Jamieson, B. G. M." box="[976,1168,1621,1645]" pageId="16" pageNumber="417" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson 1968: 75</bibRefCitation>
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</treatmentCitation>
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,
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<figureCitation id="77B72A0C7946FFE3CAD5F9D6FB66F9DD" box="[1179,1250,1621,1645]" captionStart="FIGURE 15" captionStartId="28.[151,250,697,721]" captionTargetBox="[176,1399,189,662]" captionTargetId="figure-21@28.[176,1412,181,673]" captionTargetPageId="28" captionText="FIGURE 15. Righiella jamiesoni Omodeo and Coates 2001. A. Scheme showing arrangement of genital organs and vascular commissures. B. Cross section through spermathecal duct; C. Cross section through prostate. Abbreviations: a, spermathecal ampulla, at, male atrium, cv, commissural vessels, d, spermathecal duct, md, male duct, os, egg sac, pr, prostate, sps, sperm sac. From Omodeo and Coates (2001), reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.14022669" httpUri="https://zenodo.org/record/14022669/files/figure.png" pageId="16" pageNumber="417">Fig. 15</figureCitation>
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–18;
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<treatmentCitation id="6E2D10987946FFE3CB58F9D6FE43F93C" author="Brinkhurst, R. O. & Jamieson, B. G. M." page="713" pageId="16" pageNumber="417" year="1971">
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<bibRefCitation id="8B1D4B787946FFE3CB58F9D6FE43F93C" author="Brinkhurst, R. O. & Jamieson, B. G. M." pageId="16" pageNumber="417" pagination="708 - 722" refId="ref22671" refString="Brinkhurst, R. O. & Jamieson, B. G. M. (1971) 14. Family Alluroididae. In: Brinkhurst, R. O. & Jamieson, B. G. M., Aquatic Oligochaeta of the World. Oliver and Boyd, Edinburgh, pp. 708 - 722." type="book chapter" year="1971">Brinkhurst & Jamieson 1971: 713–714</bibRefCitation>
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</treatmentCitation>
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,
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<figureCitation id="77B72A0C7946FFE3CF9FF9F7FDD9F93C" box="[465,605,1652,1676]" captionStart="FIGURE 14" captionStartId="26.[151,250,1837,1861]" captionTargetBox="[206,1382,421,1809]" captionTargetId="figure-158@26.[201,1386,417,1813]" captionTargetPageId="26" captionText="FIGURE 14. Lacandodrilus paludosus Fragoso & Rojas 2023. A. External view of the anterior ventral region of a clitellate adult. B. Internal view of the anterior region of an adult. C. Spermatheca. Abbreviations: a, ampulla: clit, clitellum; d, duct; dv, dorsal vessel; gg, genital glands (prostates?); img, internal male gonoducts; mf, male funnels; n, nephridia; oe, oesophagus; sg, septal glands. Modified from Fragoso & Rojas (2023)." figureDoi="http://doi.org/10.5281/zenodo.14022665" httpUri="https://zenodo.org/record/14022665/files/figure.png" pageId="16" pageNumber="417">Fig. 14.3D, E</figureCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="EF3336897946FFE3CED9F910FE08F97A" blockId="16.[151,1437,1591,1768]" pageId="16" pageNumber="417">
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<treatmentCitationGroup id="CF9C11A77946FFE3CED9F910FE08F97A" pageId="16" pageNumber="417">
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<emphasis id="DDF8EA9B7946FFE3CED9F910FF20F91A" box="[151,164,1683,1706]" italics="true" pageId="16" pageNumber="417">?</emphasis>
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<treatmentCitation id="6E2D10987946FFE3CEE7F910FD40F91A" author="Michaelsen, W." box="[169,708,1683,1707]" page="7" pageId="16" pageNumber="417" year="1913">
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<taxonomicName id="288C4D0A7946FFE3CEE7F910FD40F91A" ID-CoL="C2D6" authority="Beddard. Michaelsen 1913: 7" authorityName="Beddard. Michaelsen" authorityPageNumber="7" authorityYear="1913" box="[169,708,1683,1707]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="16" pageNumber="417" phylum="Annelida" rank="species" species="tanganyikae">
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<emphasis id="DDF8EA9B7946FFE3CEE7F910FE10F91B" box="[169,404,1683,1707]" italics="true" pageId="16" pageNumber="417">Alluroides tanganyikae</emphasis>
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Beddard.
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<bibRefCitation id="8B1D4B787946FFE3CFB5F910FD40F91A" author="Michaelsen, W." box="[507,708,1683,1707]" pageId="16" pageNumber="417" pagination="1 - 32" refId="ref24002" refString="Michaelsen, W. (1913) Oligochaten vom tropischen und sudlichsubtropischen Afrika. Zoologica, Stuttgart 68, 1 - 32." type="journal article" year="1913">Michaelsen 1913: 7</bibRefCitation>
|
|||
|
</taxonomicName>
|
|||
|
</treatmentCitation>
|
|||
|
, Pl. XIX,
|
|||
|
<figureCitation id="77B72A0C7946FFE3CD67F910FCE2F91B" box="[809,870,1683,1707]" captionStart="FIGURE 9" captionStartId="18.[151,250,1338,1362]" captionTargetBox="[584,994,1126,1314]" captionTargetId="figure-367@18.[567,1021,1095,1316]" captionTargetPageId="18" captionText="FIGURE 9. Alluroides ruwenzoriensis Brinkhurst 1964. Atrium and vas deferens entering at midlength. From Brinkhurst (1964)." figureDoi="http://doi.org/10.5281/zenodo.14022649" httpUri="https://zenodo.org/record/14022649/files/figure.png" pageId="16" pageNumber="417">Fig. 9</figureCitation>
|
|||
|
; 1914 a: 89; 1914 b: 165, Pl. V.
|
|||
|
<figureCitation id="77B72A0C7946FFE3CAE6F910FB76F91B" box="[1192,1266,1683,1707]" captionStart="FIGURE 13" captionStartId="24.[151,250,1144,1168]" captionTargetBox="[455,1137,189,1112]" captionTargetId="figure-21@24.[443,1145,181,1120]" captionTargetPageId="24" captionText="FIGURE 13. Kathrynella guyanae Omodeo 1996.A. Ventral view of anterior segments.Note the pale lateral line.B. arrangement of genital organs. Abbreviations: a, atrium; gp, genital papilla; ♂, male pore; sp, spematheca; t, testis; ts, thickened septum. The penial setae are seen projecting from the male pore. Modified from Omodeo (1996)." figureDoi="http://doi.org/10.5281/zenodo.14022661" httpUri="https://zenodo.org/record/14022661/files/figure.png" pageId="16" pageNumber="417">Fig. 13</figureCitation>
|
|||
|
; 1915: 29; 1935: 36; 1936: 37 (part.)
|
|||
|
</treatmentCitationGroup>
|
|||
|
</paragraph>
|
|||
|
<paragraph id="EF3336897946FFE3CED9F953FD8AF958" blockId="16.[151,1437,1591,1768]" box="[151,526,1744,1768]" pageId="16" pageNumber="417">
|
|||
|
(Non)
|
|||
|
<taxonomicName id="288C4D0A7946FFE3CE94F953FD8EF958" ID-CoL="C2D6" authority="Beddard 1906" authorityName=", Beddard" authorityYear="1906" box="[218,522,1744,1768]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="16" pageNumber="417" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CE94F953FEF6F958" box="[218,370,1744,1768]" italics="true" pageId="16" pageNumber="417">A. tanganyikae</emphasis>
|
|||
|
<bibRefCitation id="8B1D4B787946FFE3CF36F953FD8EF958" author="Beddard, F. E. & Cunnington, W. A. & Report on the Oligochaeta & Proceedings of the Zoological Society of London" box="[376,522,1744,1768]" pageId="16" pageNumber="417" pagination="206 - 218" refId="ref22511" refString="Beddard, F. E. (190 a 6) Zoological results of the Third Tanganyika Expedition, conducted by Cunnington, W. A. (1904 - 1905) Report on the Oligochaeta. Proceedings of the Zoological Society of London, 1906, 206 - 218. https: // doi. org / 10.1111 / j. 1469 - 7998.1906. tb 08432. x" type="book chapter" year="1906">Beddard 1906</bibRefCitation>
|
|||
|
</taxonomicName>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="A79665027946FFE2CED9F895FB6EF961" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="description">
|
|||
|
<paragraph id="EF3336897946FFE2CED9F895FDEEFF65" blockId="16.[151,1436,1814,2021]" lastBlockId="17.[151,1436,151,213]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
|||
|
<materialsCitation id="5FE43CD47946FFE2CED9F895FDE2FF65" collectionCode="VIII" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" specimenCount="1">
|
|||
|
Length
|
|||
|
<quantity id="28749B6C7946FFE3CEA0F895FEEEF881" box="[238,362,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.8" metricValueMin="2.5" pageId="16" pageNumber="417" unit="mm" value="36.5" valueMax="48.0" valueMin="25.0">25–48 mm</quantity>
|
|||
|
, width 1.0 mm, segments 202. Prolobous. Form approximately cylindrical throughout, anterior segments at least biannulate with the transverse furrow shortly postsetal. Lateral line in
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CAD6F8B9FB36F8E4" box="[1176,1202,1850,1876]" italics="true" pageId="16" pageNumber="417">bc</emphasis>
|
|||
|
, nearer to
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CB7AF8B9FAC6F8E4" box="[1332,1346,1850,1876]" italics="true" pageId="16" pageNumber="417">b</emphasis>
|
|||
|
than to
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CED9F8DCFF27F8C8" box="[151,163,1887,1912]" italics="true" pageId="16" pageNumber="417">c</emphasis>
|
|||
|
. Anus terminal. Ventral setal couples of XIII absent at maturity; all setae sigmoid, with approximately central, well–developed node; none modified as genital setae. Outer setal couples dorsal,
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CA76F801FBEFF82C" box="[1080,1131,1922,1948]" italics="true" pageId="16" pageNumber="417">dd:u</emphasis>
|
|||
|
=ca 0.2. Clitellum annular, XII–XVI. Male pores a pair precisely lateral, or slightly ventrolateral, nearer to
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CA53F825FBAFF870" box="[1053,1067,1958,1984]" italics="true" pageId="16" pageNumber="417">b</emphasis>
|
|||
|
than to
|
|||
|
<emphasis id="DDF8EA9B7946FFE3CAC6F824FB10F870" box="[1160,1172,1959,1984]" italics="true" pageId="16" pageNumber="417">c</emphasis>
|
|||
|
lines in the setal arc of XIII, each surrounded by a circular tumescence almost filling the segment longitudinally. Female pores in 13/14 directly in line with the male pores, inconspicuous. Spermathecal pore paired or single, dorsal and anterior in
|
|||
|
<collectionCode id="899DAE4C7947FFE2CB2BFF14FA1CFF01" box="[1381,1432,151,177]" pageId="17" pageNumber="418">VIII</collectionCode>
|
|||
|
, in the
|
|||
|
<typeStatus id="3037882B7947FFE2CE93FF38FE89FF65" box="[221,269,187,213]" pageId="17" pageNumber="418">type</typeStatus>
|
|||
|
paired near the dorsal midline
|
|||
|
</materialsCitation>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
<caption id="BBF366017947FFE2CED9FC47FDB2FB20" ID-DOI="http://doi.org/10.5281/zenodo.14022647" ID-Zenodo-Dep="14022647" httpUri="https://zenodo.org/record/14022647/files/figure.png" pageId="17" pageNumber="418" startId="17.[151,250,964,988]" targetBox="[185,1401,247,940]" targetPageId="17" targetType="figure">
|
|||
|
<paragraph id="EF3336897947FFE2CED9FC47FDB2FB20" blockId="17.[151,1437,963,1168]" pageId="17" pageNumber="418">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CED9FC47FE93FC6C" bold="true" box="[151,279,964,988]" pageId="17" pageNumber="418">FIGURE 8.</emphasis>
|
|||
|
<taxonomicName id="288C4D0A7947FFE2CF6FFC40FD10FC6C" authority="Beddard 1894" authorityName="Beddard" authorityYear="1894" box="[289,660,963,988]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="17" pageNumber="418" phylum="Annelida" rank="species" species="pordagei">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CF6FFC40FE77FC6B" bold="true" box="[289,499,963,987]" italics="true" pageId="17" pageNumber="418">Alluroides pordagei</emphasis>
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CFB0FC47FD10FC6C" author="Beddard, F. E." box="[510,660,964,988]" pageId="17" pageNumber="418" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard 1894</bibRefCitation>
|
|||
|
</taxonomicName>
|
|||
|
.
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CCEFFC47FD37FC6C" bold="true" box="[673,691,964,988]" pageId="17" pageNumber="418">A</emphasis>
|
|||
|
. Longitudinal section of the anterior region of the body of Beddard’s paratype.
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CEB5FC6BFE88FBB0" bold="true" box="[251,268,1000,1024]" pageId="17" pageNumber="418">B</emphasis>
|
|||
|
. Spermatheca.
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CFE9FC6BFE3DFBB0" bold="true" box="[423,441,1000,1024]" pageId="17" pageNumber="418">C</emphasis>
|
|||
|
. Longitudinal section of wall of atrium.
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CD1FFC6BFCE7FBB0" bold="true" box="[849,867,1000,1024]" pageId="17" pageNumber="418">D</emphasis>
|
|||
|
. Transverse section of atrium. Abbreviations: at, atrium; at ch, atrial chamber; br, brain; cil, cilia; ep, epidermis; epi, epithelium; gl duc, glandular ductule(s); gl sh, glandular sheath; int, intestine; lum, lumen; m, mouth; ph m, pharyngeal musculature; per, peritoneum; sep.gl, septal gland; sp, spermatheca; sperm, spermatozoa; sph, sphincter; sp epi, spermathecal epithelium; sp m sh, muscular sheath of spermatheca; v.d, vas deferens.. Slightly modified from
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CFC9FBFBFDB7FB20" author="Jamieson, B. G. M." box="[391,563,1144,1168]" pageId="17" pageNumber="418" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson (1968)</bibRefCitation>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<paragraph id="EF3336897947FFE2CE89FB61FB6EF961" blockId="17.[151,1437,1250,2032]" pageId="17" pageNumber="418">Septa 4/5 (the first clearly developed), 9/10 exceptionally strongly thickened, 10/11 thin and attenuated, 11/12 and (though less so) 12/13 moderately thickened, the succeeding septa thin. Septal glands posterior in V– IX? Nephridia commencing in XII–XVI. Pharynx anterior to septum 4/5. Intestine commencing in XIII, internal epithelium with tall cilia; oesophagus similarly ciliated in XII in front of the oesophageal valve; ciliation not certainly present further anteriorly. Brown–granuled chloragogen cells commencing (always?) at 1/2 IX. Dorsal blood vessel large and adherent to the gut in XIII posteriorly; slender and freer anteriorly to this. Hearts (posterior commissural) very tortuous extending posteriorly into XI(XII?). Septum 10/11 bulging posteriorly far into XI to form a wide–mouthed (unpaired?) seminal vesicle. Muscular tubes of the atria forming slender somewhat coiled tubes extending from XIII into XIV or as far as XVI and widening immediately before opening at the male pores. With the exception of the terminal ectal chamber each tubular atrium surrounded by a sheath of unicellular prostate cells, which approximately doubled the width of the apparatus, the cells communicating with the atrial lumen by minute ductules. Vasa deferentia running in the glandular prostatic duct and presumably entering the atrium entally. Ovaries and funnels in XIII. Oviduct penetrating the body wall immediately behind septum 13/14. Female gonadial cells, at maturity, in XIII–XXI, probably in ovisacs. Spermatheca single or paired, dorsally in VIII.</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="A79665027947FFE2CE89F959FD60F834" pageId="17" pageNumber="418" type="distribution">
|
|||
|
<paragraph id="EF3336897947FFE2CE89F959FD60F834" blockId="17.[151,1437,1250,2032]" pageId="17" pageNumber="418">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CE89F959FEDDF944" bold="true" box="[199,345,1754,1780]" pageId="17" pageNumber="418">Distribution</emphasis>
|
|||
|
.
|
|||
|
<collectingCountry id="979B76197947FFE2CF29F958FE30F945" box="[359,436,1755,1781]" name="Kenya" pageId="17" pageNumber="418">Kenya</collectingCountry>
|
|||
|
: swamp
|
|||
|
<quantity id="28749B6C7947FFE2CC5BF959FDE2F944" box="[533,614,1754,1780]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="17" pageNumber="418" unit="mi" value="4.0">4 miles</quantity>
|
|||
|
inland from
|
|||
|
<collectingCountry id="979B76197947FFE2CCB8F959FCE0F944" box="[758,868,1754,1780]" name="Kenya" pageId="17" pageNumber="418">Mombasa</collectingCountry>
|
|||
|
(
|
|||
|
<typeStatus id="3037882B7947FFE2CD3DF959FC28F944" box="[883,940,1754,1780]" pageId="17" pageNumber="418">Type</typeStatus>
|
|||
|
locality,
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CA5DF959FB30F944" author="Beddard, F. E." box="[1043,1204,1754,1780]" pageId="17" pageNumber="418" pagination="201 - 269" refId="ref22462" refString="Beddard, F. E. (1894) A contribution to our knowledge of the Oligochaeta of tropical eastern Africa. Quarterly Journal of Microscopical Science, 36, 201 - 269. https: // doi. org / 10.1242 / jcs. s 2 - 36.142.201" type="journal article" year="1894">Beddard 1894</bibRefCitation>
|
|||
|
); lower forest region of Mt.
|
|||
|
<collectingCountry id="979B76197947FFE2CEA8F97CFEABF8A9" box="[230,303,1791,1817]" name="Kenya" pageId="17" pageNumber="418">Kenya</collectingCountry>
|
|||
|
at
|
|||
|
<quantity id="28749B6C7947FFE2CF1CF97DFE2CF8A9" box="[338,424,1790,1817]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="418" unit="m" value="2400.0">2400 m</quantity>
|
|||
|
(
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CFF9F97DFD0BF8A9" author="Michaelsen, W." box="[439,655,1790,1817]" pageId="17" pageNumber="418" pagination="81 - 127" refId="ref24025" refString="Michaelsen, W. (1914 a) Oligochaten vom tropischen Afrika. Mitteilungen aus dem Zoologischen Museum in Hamburg, 21, 81 - 127." type="journal article" year="1914">Michaelsen 1914 a</bibRefCitation>
|
|||
|
, b;
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CCF4F97DFCF9F8A9" author="Michaelsen, W." box="[698,893,1790,1817]" pageId="17" pageNumber="418" pagination="23 - 42" refId="ref24095" refString="Michaelsen, W. (1915) Oligochaeta. In: Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale. 1911 - 1912. Resultats scientifiques. Vermes, 2, pp. 23 - 42." type="journal article" year="1915">Michaelsen 1915</bibRefCitation>
|
|||
|
, as
|
|||
|
<taxonomicName id="288C4D0A7947FFE2CDE6F97DFBD5F8A8" authorityName=", Beddard" authorityYear="1906" box="[936,1105,1790,1816]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="17" pageNumber="418" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CDE6F97DFBD5F8A8" box="[936,1105,1790,1816]" italics="true" pageId="17" pageNumber="418">A. tanganyikae</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
); Kahawa stream, Athi River tributary, near Nairobi, collector B.G.M. Jamieson,
|
|||
|
<date id="9B3210497947FFE2CCB9F8A1FC1CF88C" box="[759,920,1826,1853]" pageId="17" pageNumber="418" value="1967-12-11">11 Dec. 1967</date>
|
|||
|
.
|
|||
|
<collectingCountry id="979B76197947FFE2CDE6F8A1FB3DF88C" box="[936,1209,1826,1852]" name="Republic of the Congo" pageId="17" pageNumber="418">Republic of the Congo</collectingCountry>
|
|||
|
: Mulongo, Niunzu; Albertville (
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CF6CF8C5FE68F8D1" author="Michaelsen, W." box="[290,492,1862,1889]" pageId="17" pageNumber="418" pagination="33 - 95" refId="ref24136" refString="Michaelsen, W. (1935) Oligochaten von Belgisch-Kongo. Revue de Zoologie et de Botanie Africaines, 27, 33 - 95." type="journal article" year="1935">Michaelsen 1935</bibRefCitation>
|
|||
|
); Leopoldville (
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CCEBF8C5FCEAF8D0" author="Michaelsen, W." box="[677,878,1862,1888]" pageId="17" pageNumber="418" pagination="37 - 71" refId="ref24161" refString="Michaelsen, W. (1936) Oligochaten von Belgisch-Kongo III. Revue de Zoologie et de Botanie Africaines, 29, 37 - 71." type="journal article" year="1936">Michaelsen 1936</bibRefCitation>
|
|||
|
).
|
|||
|
<collectingCountry id="979B76197947FFE2CDCBF8C5FC76F8D0" box="[901,1010,1862,1888]" name="Zimbabwe" pageId="17" pageNumber="418">Rhodesia</collectingCountry>
|
|||
|
: swampy earth, Zambezi River, near Victoria Falls (as
|
|||
|
<taxonomicName id="288C4D0A7947FFE2CF13F8E9FD53F834" authority=", Michaelsen 1913" authorityName=", Michaelsen" authorityYear="1913" box="[349,727,1898,1924]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="17" pageNumber="418" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CF13F8E9FD88F834" box="[349,524,1898,1924]" italics="true" pageId="17" pageNumber="418">A. tanganyikae,</emphasis>
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CC5DF8E9FD53F834" author="Michaelsen, W." box="[531,727,1898,1924]" pageId="17" pageNumber="418" pagination="1 - 32" refId="ref24002" refString="Michaelsen, W. (1913) Oligochaten vom tropischen und sudlichsubtropischen Afrika. Zoologica, Stuttgart 68, 1 - 32." type="journal article" year="1913">Michaelsen 1913</bibRefCitation>
|
|||
|
</taxonomicName>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="A79665027947FFE1CE89F80DFD08FD8D" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418" type="discussion">
|
|||
|
<paragraph id="EF3336897947FFE2CE89F80DFD7EF840" blockId="17.[151,1437,1250,2032]" pageId="17" pageNumber="418">
|
|||
|
<emphasis id="DDF8EA9B7947FFE2CE89F80DFEB0F818" bold="true" box="[199,308,1934,1960]" pageId="17" pageNumber="418">Remarks</emphasis>
|
|||
|
. Because of the uncertain status of Michaelsen’s material this account (from
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CAFAF80DFADBF818" author="Jamieson, B. G. M." box="[1204,1375,1934,1960]" pageId="17" pageNumber="418" pagination="55 - 86" refId="ref23501" refString="Jamieson, B. G. M. (1968) A taxonometric investigation of the Alluroididae (Oligochaeta). Journal of Zoology, 155, 55 - 86. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 03029. x" type="journal article" year="1968">Jamieson 1968</bibRefCitation>
|
|||
|
) was derived solely from Beddard’s
|
|||
|
<typeStatus id="3037882B7947FFE2CC45F831FDB8F87C" box="[523,572,1970,1996]" pageId="17" pageNumber="418">type</typeStatus>
|
|||
|
–series, and his description, and incorporated results of examinations by
|
|||
|
<bibRefCitation id="8B1D4B787947FFE2CED9F855FEE5F840" author="Brinkhurst, R. O." box="[151,353,2006,2032]" pageId="17" pageNumber="418" pagination="527 - 536" refId="ref22581" refString="Brinkhurst, R. O. (1964) A taxonomic revision of the Alluroididae (Oligochaeta). Proceedings of the Zoological Society of London, 142, 527 - 536. https: // doi. org / 10.1111 / j. 1469 - 7998.1964. tb 04512. x" type="journal article" year="1964">Brinkhurst (1964)</bibRefCitation>
|
|||
|
and Jamieson (Athi River material).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="EF3336897944FFE1CE89FF14FE86FEF1" blockId="18.[151,1437,151,573]" pageId="18" pageNumber="419">
|
|||
|
The extreme septal thickening, and its distribution, now emerge as diagnostic features of
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CAEFFF14FAA7FF01" authorityName="Beddard" authorityYear="1894" box="[1185,1315,151,177]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="pordagei">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CAEFFF14FAA7FF01" box="[1185,1315,151,177]" italics="true" pageId="18" pageNumber="419">A. pordagei</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
(condition still unknown, however, in
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CF80FF38FD0EFF64" box="[462,650,187,213]" italics="true" pageId="18" pageNumber="419">
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CF80FF38FDF3FF65" authorityName=", Michaelsen" authorityYear="1913" box="[462,631,187,213]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="tanganyikae">A. tanganyikae</taxonomicName>
|
|||
|
s
|
|||
|
</emphasis>
|
|||
|
. Beddard and s. Michaelsen). Intraspecific variation from a paired to a single spermatheca (in VIII) is confirmed and reinforces Brinkhurst’s conclusion (1964) that
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CA99FF5CFA11FF48" box="[1239,1429,223,249]" italics="true" pageId="18" pageNumber="419">
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CA99FF5CFA05FF49" authorityName=", Michaelsen" authorityYear="1913" box="[1239,1409,223,249]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="tanganyikae">A. tanganyikae</taxonomicName>
|
|||
|
s
|
|||
|
</emphasis>
|
|||
|
. Michaelsen (spermatheca single in VIII), but not
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CC86FE80FD57FEAC" box="[712,723,259,284]" italics="true" pageId="18" pageNumber="419">s</emphasis>
|
|||
|
. Beddard (spermatheca single in IX), should be included in
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CBCAFE80FF7FFEF1" authorityName="Beddard" authorityYear="1894" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="pordagei">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CBCAFE80FF7FFEF1" italics="true" pageId="18" pageNumber="419">A. pordagei</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="EF3336897944FFE1CE89FEC8FD08FD8D" blockId="18.[151,1437,151,573]" pageId="18" pageNumber="419">
|
|||
|
The decision by
|
|||
|
<bibRefCitation id="8B1D4B787944FFE1CFCEFEC8FDD6FED5" author="Michaelsen, W." box="[384,594,331,357]" pageId="18" pageNumber="419" pagination="37 - 71" refId="ref24161" refString="Michaelsen, W. (1936) Oligochaten von Belgisch-Kongo III. Revue de Zoologie et de Botanie Africaines, 29, 37 - 71." type="journal article" year="1936">Michaelsen (1936)</bibRefCitation>
|
|||
|
to recognize three subspecies,
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CDE2FEC8FB94FED5" authorityName="Beddard" authorityYear="1894" box="[940,1040,331,357]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="pordagei">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CDE2FEC8FB94FED5" box="[940,1040,331,357]" italics="true" pageId="18" pageNumber="419">pordagei</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
,
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CA53FEC8FB22FED5" authorityName=", Michaelsen" authorityYear="1913" box="[1053,1190,331,357]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CA53FEC8FB22FED5" box="[1053,1190,331,357]" italics="true" pageId="18" pageNumber="419">tanganyikae</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CA92FEC8FAC4FED4" box="[1244,1344,331,356]" italics="true" pageId="18" pageNumber="419">congicus</emphasis>
|
|||
|
, has not been followed here though it still deserves consideration.
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CD53FEECFC41FE39" authorityName=", Michaelsen" authorityYear="1913" box="[797,965,367,393]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CD53FEECFCAAFE39" box="[797,814,367,393]" italics="true" pageId="18" pageNumber="419">A</emphasis>
|
|||
|
.
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CD72FEECFC41FE39" box="[828,965,367,393]" italics="true" pageId="18" pageNumber="419">tanganyikae</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
has been retained as a distinct species but recognition of
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CF75FE10FE1BFE1C" box="[315,415,403,428]" italics="true" pageId="18" pageNumber="419">congicus</emphasis>
|
|||
|
, from Leopoldsville, as a subspecies appears reasonable. Michaelsen distinguished it by the spermatheca with a thin–walled, sac–like ampulla and sharply demarcated muscular duct. He added that
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CB77FE34FA19FE60" box="[1337,1437,439,464]" italics="true" pageId="18" pageNumber="419">congicus</emphasis>
|
|||
|
was
|
|||
|
<quantity id="28749B6C7944FFE1CE84FE58FED4FE45" box="[202,336,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="18" pageNumber="419" unit="mm" value="95.0" valueMax="100.0" valueMin="90.0">90–100 mm</quantity>
|
|||
|
long, with
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CF81FE58FE6DFE44" box="[463,489,475,500]" italics="true" pageId="18" pageNumber="419">ca</emphasis>
|
|||
|
210 segments, the largest form, whereas the typical subspecies was
|
|||
|
<quantity id="28749B6C7944FFE1CAABFE58FAB0FE45" box="[1253,1332,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="419" unit="mm" value="22.0">22 mm</quantity>
|
|||
|
long, the smallest form, while
|
|||
|
<taxonomicName id="288C4D0A7944FFE1CFCFFE7CFD8EFDA9" authorityName=", Michaelsen" authorityYear="1913" box="[385,522,511,537]" class="Clitellata" family="Alluroididae" genus="Alluroides" kingdom="Animalia" order="Alluroidida" pageId="18" pageNumber="419" phylum="Annelida" rank="species" species="tanganyikae">
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CFCFFE7CFD8EFDA9" box="[385,522,511,537]" italics="true" pageId="18" pageNumber="419">tanganyikae</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
had a length of
|
|||
|
<quantity id="28749B6C7944FFE1CCF5FE7DFC8FFDA9" box="[699,779,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="419" unit="mm" value="45.0">45 mm</quantity>
|
|||
|
, with
|
|||
|
<emphasis id="DDF8EA9B7944FFE1CD00FE7CFCECFDA8" box="[846,872,511,536]" italics="true" pageId="18" pageNumber="419">ca</emphasis>
|
|||
|
150 segments, intermediate between the two other forms. Some of these figures no longer hold.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|