291 lines
38 KiB
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291 lines
38 KiB
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<document id="3749AD62D7E9D5E7FE0384D8E1BF14B3" ID-DOI="10.5281/zenodo.170066" ID-GBIF-Dataset="e3737643-9288-451a-b814-4f22256e41a4" ID-ISSN="11755326" ID-Zenodo-Dep="170066" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749874227" checkinUser="plazi" docAuthor="Peters, J. G., Flowers, R. W., Hubbard, M. D., Domínguez, E. & Savage, H. M." docDate="2005" docId="03F787BB0337FFCBF9733DA3F1FAC5A4" docLanguage="en" docName="zt01054p060.pdf" docOrigin="Zootaxa 1054" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hagenulus marshalli Peters, Flowers, Hubbard, Domínguez & Savage, 2005, sp. n." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFCEFFC30332FFCCF87B3B65F310C04F" masterDocTitle="New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="56" updateTime="1698211586579" updateUser="plazi">
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<mods:titleInfo id="6622DA8B8C94F7BA5027FFA8A28ABEFB">
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<mods:title id="0522A5DD6018EA9AC7F3FDA4718A1A37">New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="1F7BC3A905D3880739D5499DD1C0609C" type="personal">
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<mods:namePart id="9C2F6B9DCACF1FCA4B228ACABFF536D2">Peters, J. G.</mods:namePart>
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</mods:name>
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<mods:name id="016D5BFBC8C9EF500ED8B470A2A1A7A7" type="personal">
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<mods:namePart id="CF8C2F5BBDBA2FFC00348E4C3BB52527">Flowers, R. W.</mods:namePart>
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</mods:name>
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<mods:name id="658CBF0FD2C3A680061FBFB728650974" type="personal">
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</mods:role>
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<mods:namePart id="8007A0FD8F5C0962F1C5F7CED88129FD">Hubbard, M. D.</mods:namePart>
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</mods:name>
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<mods:name id="C5F565E7481E9A12EE3FDF7660680228" type="personal">
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</mods:role>
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<mods:namePart id="CD48913B6273C9B0B458B3EFA33A6EFA">Domínguez, E.</mods:namePart>
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</mods:role>
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<mods:namePart id="106B0133F320D4FB3090E2F3F7B265A7">Savage, H. M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8A965E6183CECACEDAD2C906089A1200">text</mods:typeOfResource>
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<mods:title id="2B9962F94113059B126E507C637CD36B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DA1AFDEC2682EFDB174D495A1B9E5759">
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<mods:date id="17702F94A2F16A79118DC95E7D5D023A">2005</mods:date>
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<mods:detail id="604745BB5398573870DE802AD2837590" type="volume">
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<mods:number id="AEA68B2896B4750F5D85AC37B5AF0694">1054</mods:number>
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<mods:classification id="50A5D3786D4D5DDFA63B0390BBB7B456">journal article</mods:classification>
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<mods:identifier id="0B25A085BF40296510322652F54D9F2A" type="DOI">10.5281/zenodo.170066</mods:identifier>
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<mods:identifier id="F35C02FAAF4113A6630731D841B38406" type="GBIF-Dataset">e3737643-9288-451a-b814-4f22256e41a4</mods:identifier>
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<mods:identifier id="DB427DD2C37B27168E1A9D4CCCDD4944" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="3E88EE4BAE35967C6FFDFC632B19FFCE" type="Zenodo-Dep">170066</mods:identifier>
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<treatment id="03F787BB0337FFCBF9733DA3F1FAC5A4" ID-DOI="http://doi.org/10.5281/zenodo.6267048" ID-GBIF-Taxon="119337550" ID-Zenodo-Dep="6267048" LSID="urn:lsid:plazi:treatment:03F787BB0337FFCBF9733DA3F1FAC5A4" httpUri="http://treatment.plazi.org/id/03F787BB0337FFCBF9733DA3F1FAC5A4" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56">
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<subSubSection id="C34465260337FFC9F9733DA3F2B7C74C" pageId="5" pageNumber="56" type="nomenclature">
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<paragraph id="8BE136AD0337FFC9F9733DA3F142C6AF" blockId="5.[264,594,1734,1795]" box="[264,594,1734,1760]" pageId="5" pageNumber="56">
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<heading id="D0A981C10337FFC9F9733DA3F142C6AF" bold="true" box="[264,594,1734,1760]" fontSize="11" level="1" pageId="5" pageNumber="56" reason="1">
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<emphasis id="B92AEABF0337FFC9F9733DA3F142C6AF" bold="true" box="[264,594,1734,1760]" pageId="5" pageNumber="56">
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<taxonomicName id="4C5E4D2E0337FFC9F9733DA3F118C6AF" box="[264,520,1734,1760]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
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<emphasis id="B92AEABF0337FFC9F9733DA3F118C6AF" bold="true" box="[264,520,1734,1760]" italics="true" pageId="5" pageNumber="56">Hagenulus marshalli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21957C40337FFC9FA6B3DA3F142C6AF" box="[528,594,1734,1760]" pageId="5" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE136AD0337FFC9F9733D8CF2B7C74C" blockId="5.[264,594,1734,1795]" box="[264,423,1769,1795]" pageId="5" pageNumber="56">
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(
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<figureCitation id="13652A280337FFC9F96B3D8CF28EC74C" box="[272,414,1769,1795]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="5" pageNumber="56">Figures 3–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34465260337FFCBF9733C4BF1FAC5A4" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56" type="description">
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<paragraph id="8BE136AD0337FFCAF9733C4BF612C111" blockId="5.[264,1323,1838,1904]" lastBlockId="6.[264,1325,284,1892]" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
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This new species is represented by two unassociated male imagos in good condition from
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<collectingCountry id="F349763D0337FFC9F9733C33F27CC73F" box="[264,364,1878,1904]" name="Ecuador" pageId="5" pageNumber="56">Ecuador</collectingCountry>
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. Because it represents a significant generic record for South
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<collectingCountry id="F349763D0337FFC9FC073C33F7F4C73F" box="[1148,1252,1878,1904]" name="United States of America" pageId="5" pageNumber="56">America</collectingCountry>
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, it is described now even though nymphs and females are unknown. Previously,
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<taxonomicName id="4C5E4D2E0334FFCAFC0C3A79F7E6C17A" box="[1143,1270,284,309]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B92AEABF0334FFCAFC0C3A79F7E6C17A" box="[1143,1270,284,309]" italics="true" pageId="6" pageNumber="57">Hagenulus</emphasis>
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</taxonomicName>
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was believed to be restricted to the greater Antilles (
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<collectingCountry id="F349763D0334FFCAFB4D3A21F068C111" box="[822,888,324,350]" name="Cuba" pageId="6" pageNumber="57">Cuba</collectingCountry>
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,
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<collectingCountry id="F349763D0334FFCAFBFF3A21F0D2C111" box="[900,962,324,350]" name="Haiti" pageId="6" pageNumber="57">Haiti</collectingCountry>
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,
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<collectingCountry id="F349763D0334FFCAFBB53A21F73CC111" box="[974,1068,324,350]" name="Jamaica" pageId="6" pageNumber="57">Jamaica</collectingCountry>
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and
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<collectingCountry id="F349763D0334FFCAFC1E3A21F7E5C111" box="[1125,1269,324,350]" name="Puerto Rico" pageId="6" pageNumber="57">Puerto Rico</collectingCountry>
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).
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</paragraph>
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<paragraph id="8BE136AD0334FFCAF9433A09F7C9C5B1" blockId="6.[264,1325,284,1892]" pageId="6" pageNumber="57">
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Male imago (in alcohol). Body: 6.0–
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<quantity id="4CA69B480334FFCAFA8E3A09F044C1C9" box="[757,852,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="57" unit="mm" value="6.5">6.5 mm</quantity>
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, forewings
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<quantity id="4CA69B480334FFCAFB993A09F766C1C9" box="[994,1142,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.2" metricValueMin="6.8" pageId="6" pageNumber="57" unit="mm" value="7.0" valueMax="7.2" valueMin="6.8">6.8–7.2 mm</quantity>
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, hind wing
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<quantity id="4CA69B480334FFCAFD733A09F229C1E2" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="57" unit="mm" value="0.7">0.7 mm</quantity>
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; foreleg
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<quantity id="4CA69B480334FFCAF9D93AF6F124C1E2" box="[418,564,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.3" metricValueMin="6.5" pageId="6" pageNumber="57" unit="mm" value="6.9" valueMax="7.3" valueMin="6.5">6.5–7.3 mm</quantity>
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; cerci
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<quantity id="4CA69B480334FFCAFAF93AF6F1FFC1E2" box="[642,751,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="57" unit="mm" value="12.5">12.5 mm</quantity>
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. Head brown, dorsal portion of eyes meeting on meson of head, ocelli large. Pronotum light brown, heavily washed with blackish brown on median carina and laterally; meso and metanotum yellowish brown, pleural area extending anteriorly from forewing to pronotum washed with darker brown. Pro and mesosternum brown, mesobasisternum a little lighter, median margins of mesofurcasternal protuberances darker; metasternum heavily washed with blackish brown. Forewings: Sc brown, R a lighter brown, other longitudinal veins pale yellowbrown to hyaline, cross veins dark brown, surrounded by small blackish brown clouds in basal half of wing, clouds heaviest medially (
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<figureCitation id="13652A280334FFCAF99C39AAF13FC2A6" box="[487,559,719,745]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 3</figureCitation>
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), membrane of costal and subcostal area pale golden brown, the color fading to hyaline throughout remainder of wing membrane.
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<collectingCountry id="F349763D0334FFCAFC783992F72EC35E" box="[1027,1086,759,785]" name="India" pageId="6" pageNumber="57">Hind</collectingCountry>
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wings with reduced venation, veins and membrane pale yellowbrown to hyaline (
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<figureCitation id="13652A280334FFCAFBAC387BF72DC377" box="[983,1085,798,824]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 4–5</figureCitation>
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); hind wings curved dorsally over scutellum (
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<figureCitation id="13652A280334FFCAFA563823F166C32F" box="[557,630,838,864]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 6</figureCitation>
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). Legs: ratios of foreleg segments to tibia (length of tibiae
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<quantity id="4CA69B480334FFCAF9733808F28EC3C8" box="[264,414,877,903]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="6" pageNumber="57" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
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): femur 0.46–0.49: tarsal segments 0.03; 0.32–0.34; 0.22–0.29; 0.13–0.14; 0.06–0.07; femur of foreleg brown, faded to pale brown basally and a little darker medially; remainder of foreleg segments with tibia pale yellowish brown and tarsal segments paler, a narrow dark brown band near apex of tibia, apical third to apical half of tarsal segments 2–4 darker brown, tarsal segments 1 and 5 without color markings; other legs with light yellowish brown femora, tibiae and tarsi paler. Abdominal tergum 1 light brown with narrow blackish brown posterior margin and lateral area; terga 2–5 hyaline with short, anterior sublateral dark streaks and blackish brown posterior band extended laterally and anteriorly as in
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<figureCitation id="13652A280334FFCAF9BB3FCCF117C48C" box="[448,519,1193,1219]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 9</figureCitation>
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; terga 6 and 7 with similar lateral patterns, although posterior bands reduced or absent; terga 8–10 opaque, light brown, with anterior blackish brown wash on terga 8–9, tergum 10 with only anterior submedian streaks. Abdominal sterna 1 and 2 heavily washed with blackish brown, wash continuing anterolaterally and anterosubmedially to sternum 3; sterna 4–8 with light blackish brown markings anterolaterally, marks progressively reduced in size on posterior sterna; sternum 9 light yellowish brown. Genitalia: styliger plate light brown; forceps pale yellowbrown; penes light brown, divided to near base, divergent, without spines, penes somewhat twisted as in
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<figureCitation id="13652A280334FFCAFC673ED8F791C598" box="[1052,1153,1469,1495]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 7–8</figureCitation>
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. Cerci hyaline with blackish brown rings at articulations; median filament broken off and missing.
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</paragraph>
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|
|
<paragraph id="8BE136AD0334FFCAF9433D69F0ADC602" blockId="6.[264,1325,284,1892]" pageId="6" pageNumber="57">
|
|||
|
|
Etymology: species is named for G. B.
|
|||
|
|
<collectingCountry id="F349763D0334FFCAFB7A3D69F079C669" box="[769,873,1548,1574]" name="Marshall Islands" pageId="6" pageNumber="57">Marshall</collectingCountry>
|
|||
|
|
who assisted C. W. and L. B. O’Brien in the collection of many aquatic insects in South
|
|||
|
|
<collectingCountry id="F349763D0334FFCAFB2B3D56F0A9C602" box="[848,953,1587,1613]" name="United States of America" pageId="6" pageNumber="57">America</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE136AD0334FFCAF9433D3EF183C68B" blockId="6.[264,1325,284,1892]" pageId="6" pageNumber="57">
|
|||
|
|
Material:
|
|||
|
|
<typeStatus id="54E5880F0334FFCAF9D73D3EF10CC63A" box="[428,540,1627,1653]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
|
|||
|
|
, one male imago,
|
|||
|
|
<collectingCountry id="F349763D0334FFCAFA833D3EF04EC63A" box="[760,862,1627,1653]" name="Ecuador" pageId="6" pageNumber="57">Ecuador</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<quantity id="4CA69B480334FFCAFB103D3EF0A7C63A" box="[875,951,1627,1653]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="57" unit="km" value="30.0">30 km</quantity>
|
|||
|
|
W of Puyo,
|
|||
|
|
<date id="FFE0106D0334FFCAFC293D3EF7F9C63A" box="[1106,1257,1627,1653]" pageId="6" pageNumber="57" value="1978-04-27">27–IV–1978</date>
|
|||
|
|
, L.B. & C. W. O’Brien;
|
|||
|
|
<typeStatus id="54E5880F0334FFCAF99F3DE7F15CC6D3" box="[484,588,1666,1692]" pageId="6" pageNumber="57" type="paratype">paratype</typeStatus>
|
|||
|
|
, one male imago, same data as
|
|||
|
|
<typeStatus id="54E5880F0334FFCAFBBF3DE7F73EC6D3" box="[964,1070,1666,1692]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
|
|||
|
|
. Deposited in collections of Florida A&M University.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE136AD0334FFCAF9433DB4F72BC72B" blockId="6.[264,1325,284,1892]" pageId="6" pageNumber="57">
|
|||
|
|
Diagnosis:
|
|||
|
|
<taxonomicName id="4C5E4D2E0334FFCAF9C43DB4F1A9C6A5" box="[447,697,1745,1770]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
|
|||
|
|
<emphasis id="B92AEABF0334FFCAF9C43DB4F1A9C6A5" box="[447,697,1745,1770]" italics="true" pageId="6" pageNumber="57">Hagenulus marshalli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A21957C40334FFCAFAB93DB4F013C6A4" box="[706,771,1745,1771]" pageId="6" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
can be distinguished from all other described species of
|
|||
|
|
<taxonomicName id="4C5E4D2E0334FFCAF9FD3D9CF115C75D" box="[390,517,1785,1810]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0334FFCAF9FD3D9CF115C75D" box="[390,517,1785,1810]" italics="true" pageId="6" pageNumber="57">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the following characters: 1) fork of vein MP in the forewing not symmetrical (MP2 attached to MP1 at a cross vein as in
|
|||
|
|
<figureCitation id="13652A280334FFCAFBF13C45F0DFC775" box="[906,975,1824,1850]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 3</figureCitation>
|
|||
|
|
); 2) penes without spines and slightly twisted (
|
|||
|
|
<figureCitation id="13652A280334FFCAF9B73C2FF122C72B" box="[460,562,1866,1892]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 7–8</figureCitation>
|
|||
|
|
); and 3) color pattern as illustrated in
|
|||
|
|
<figureCitation id="13652A280334FFCAFB8B3C2FF727C72B" box="[1008,1079,1866,1892]" captionStart="FIGURES 1 – 9" captionStartId="3.[264,383,1180,1204]" captionTargetBox="[276,1291,297,1120]" captionTargetId="figure@3.[266,1306,284,1144]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Figs. 1 – 2: Hind wings from two female paratypes of Ulmeritoides fidalgoi. Figs. 3 – 9: Hagenulus marshalli sp. n., male: 3, forewing; 4, hind wing (proportional); 5, hind wing enlarged; 6, lateral outline of head and thorax showing position of hind wing; 7, genitalia, ventral view; 8, detail of penis lobe; 9, lateral view of abdominal segments 4 – 6. Concave longitudinal veins stippled in Figs. 1 – 5." httpUri="https://zenodo.org/record/170067/files/figure.png" pageId="6" pageNumber="57">Fig. 9</figureCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE136AD0335FFCBF9433A79F7EBC1B1" blockId="7.[264,1324,284,1515]" pageId="7" pageNumber="58">
|
|||
|
|
Discussion: The variation in foreleg ratios is mostly due to a relatively short third tarsal segment on one of the
|
|||
|
|
<typeStatus id="54E5880F0335FFCBFA453A21F1B3C111" box="[574,675,324,350]" pageId="7" pageNumber="58" type="paratype">paratype</typeStatus>
|
|||
|
|
forelegs. All forelegs were present, but only two other legs of a potential eight were found loose in the vial. The color pattern was the same on both, but it is not known if these were meso or metathoracic legs or to which specimen each belonged. Both are placed with the
|
|||
|
|
<typeStatus id="54E5880F0335FFCBFA873AD9F076C199" box="[764,870,444,470]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
|
|||
|
|
. The intensity of color on abdominal segments 8–10 and on the mesosternum is lighter in the
|
|||
|
|
<typeStatus id="54E5880F0335FFCBFBE73A81F713C1B1" box="[924,1027,484,510]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
|
|||
|
|
than in the
|
|||
|
|
<typeStatus id="54E5880F0335FFCBFCF53A81F7E7C1B1" box="[1166,1271,484,510]" pageId="7" pageNumber="58" type="paratype">paratype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE136AD0335FFCBF9433969F098C2EF" blockId="7.[264,1324,284,1515]" pageId="7" pageNumber="58">
|
|||
|
|
In the forewings, MP2 is closely tied to MP1 by a short cross vein, not joined symmetrically as in other species of
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFA273952F1CBC21F" box="[604,731,567,592]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFA273952F1CBC21F" box="[604,731,567,592]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Nevertheless, this species is clearly a member of the
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF94E393AF2A4C237" box="[309,436,607,632]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF94E393AF2A4C237" box="[309,436,607,632]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group of genera because of the unique shape of the hind wing, the attachment of ICu1 to CuP, and forceps typical for the genus.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE136AD0335FFCBF94339D4F1FAC5A4" blockId="7.[264,1324,284,1515]" pageId="7" pageNumber="58">
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBF94339D4F2C8C284" author="Kluge" box="[312,472,689,715]" pageId="7" pageNumber="59" refString="Kluge, N. Ju. (1994) A revision of Leptophlebiidae of Cuba (Insecta, Ephemeroptera). Zoosystematica Rossica, 1993, 2 (2), 247 - 285." type="journal article" year="1994">Kluge (1994)</bibRefCitation>
|
|||
|
|
treated as subgenera all members of the
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFBAA39D4F740C285" box="[977,1104,689,714]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFBAA39D4F740C285" box="[977,1104,689,714]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group from
|
|||
|
|
<collectingCountry id="F349763D0335FFCBFC9739D4F63AC284" box="[1260,1322,689,715]" name="Cuba" pageId="7" pageNumber="58">Cuba</collectingCountry>
|
|||
|
|
previously treated as genera by
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFAE839BCF028C2BC" author="Peters" box="[659,824,729,755]" pageId="7" pageNumber="59" refString="Peters, W. L. (1971) A revision of the Leptophlebiidae of the West Indies (Ephemeroptera). Smithsonian Contributions to Zoology, 62, 1 - 48." type="journal article" year="1971">Peters (1971)</bibRefCitation>
|
|||
|
|
, including
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFBB839BCF273C355" italics="true" pageId="7" pageNumber="58">
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFBB839BCF755C2BD" box="[963,1093,729,754]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Hagenulus</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC2D39BCF7F4C2BD" box="[1110,1252,729,754]" class="Insecta" family="Leptophlebiidae" genus="Borinquena" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Borinquena</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC8F39BCF273C355" class="Insecta" family="Leptophlebiidae" genus="Careospina" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Careospina</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF9C53864F122C355" box="[446,562,769,794]" class="Insecta" family="Leptophlebiidae" genus="Traverina" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF9C53864F122C355" box="[446,562,769,794]" italics="true" pageId="7" pageNumber="58">Traverina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and established two new subgenera,
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC413864F7FBC355" box="[1082,1259,769,794]" class="Insecta" family="Leptophlebiidae" genus="Poecilophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFC413864F7FBC355" box="[1082,1259,769,794]" italics="true" pageId="7" pageNumber="58">Poecilophlebia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF973384CF2DAC30D" box="[264,458,809,834]" class="Insecta" family="Leptophlebiidae" genus="Turquinophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF973384CF2DAC30D" box="[264,458,809,834]" italics="true" pageId="7" pageNumber="58">Turquinophlebia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. One species
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFA0E384CF7D7C30C" authority="Kluge, 1994" authorityName="Kluge" authorityYear="1994" box="[629,1223,809,835]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="evanescens" subGenus="Careospina">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFA0E384CF1E4C30D" box="[629,756,809,834]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFB73384CF080C30D" box="[776,912,809,834]" italics="true" pageId="7" pageNumber="58">Careospina</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFBD8384CF737C30D" box="[931,1063,809,834]" italics="true" pageId="7" pageNumber="58">evanescens</emphasis>
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFC4A384CF7D7C30C" author="Kluge" box="[1073,1223,809,835]" pageId="7" pageNumber="59" refString="Kluge, N. Ju. (1994) A revision of Leptophlebiidae of Cuba (Insecta, Ephemeroptera). Zoosystematica Rossica, 1993, 2 (2), 247 - 285." type="journal article" year="1994">Kluge, 1994</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, has the characteristic hind wing of
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFA2E3834F1C4C325" box="[597,724,849,874]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFA2E3834F1C4C325" box="[597,724,849,874]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, penes somewhat similar to
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC4A3834F63AC325" box="[1073,1322,849,874]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFC4A3834F63AC325" box="[1073,1322,849,874]" italics="true" pageId="7" pageNumber="58">Hagenulus marshalli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A21957C40335FFCBF973381CF257C3DC" box="[264,327,889,915]" pageId="7" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, a female with a short ovipositor, and nymphs without filterfeeding adaptations. In addition to wing characters, nymphs of the
|
|||
|
|
<typeStatus id="54E5880F0335FFCBFB7838C4F026C3F4" box="[771,822,929,955]" pageId="7" pageNumber="58">type</typeStatus>
|
|||
|
|
species of
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFBC338C4F758C3F5" box="[952,1096,929,954]" italics="true" pageId="7" pageNumber="58">
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFBC338C4F727C3F5" box="[952,1079,929,954]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Hagenulus</taxonomicName>
|
|||
|
|
(
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC3338C4F254C3AC" authority="Eaton, 1882" authorityName="Eaton" authorityYear="1882" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="caligatus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFC3338C4F7C8C3F5" box="[1096,1240,929,954]" italics="true" pageId="7" pageNumber="58">H. caligatus</emphasis>
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFCA438C4F254C3AC" author="Eaton" pageId="7" pageNumber="59" refString="Eaton, A. E. (1882) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 18, 207 - 208." type="journal article" year="1882">Eaton, 1882</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) and a second Cuban species (
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFAD138ACF0FEC3AC" authority="Peters, 1971" authorityName="Peters" authorityYear="1971" box="[682,1006,969,995]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="morrisonae">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFAD138ACF045C3AD" box="[682,853,969,994]" italics="true" pageId="7" pageNumber="58">H. morrisonae</emphasis>
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFB2738ACF0FEC3AC" author="Peters" box="[860,1006,969,995]" pageId="7" pageNumber="59" refString="Peters, W. L. (1971) A revision of the Leptophlebiidae of the West Indies (Ephemeroptera). Smithsonian Contributions to Zoology, 62, 1 - 48." type="journal article" year="1971">Peters, 1971</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) have unique nymphs with filterfeeding adaptations. For this reason,
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFA833894F084C444" author="Kluge" box="[760,916,1009,1035]" pageId="7" pageNumber="59" refString="Kluge, N. Ju. (1994) A revision of Leptophlebiidae of Cuba (Insecta, Ephemeroptera). Zoosystematica Rossica, 1993, 2 (2), 247 - 285." type="journal article" year="1994">Kluge (1994)</bibRefCitation>
|
|||
|
|
removed all nonCuban species of
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF9733F7CF297C47D" box="[264,391,1049,1074]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF9733F7CF297C47D" box="[264,391,1049,1074]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the subgenus
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFA0A3F7CF1E0C47D" box="[625,752,1049,1074]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFA0A3F7CF1E0C47D" box="[625,752,1049,1074]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as their nymphs were not known with certainty, but he did not give them any new taxonomic placement. The same problem applies to
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF9733F0CF113C4CD" box="[264,515,1129,1154]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF9733F0CF113C4CD" box="[264,515,1129,1154]" italics="true" pageId="7" pageNumber="58">Hagenulus marshalli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A21957C40335FFCBFA6B3F0CF145C4CC" box="[528,597,1129,1155]" pageId="7" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, which also might represent a species of
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFC263F0CF7F4C4CD" box="[1117,1252,1129,1154]" class="Insecta" family="Leptophlebiidae" genus="Careospina" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFC263F0CF7F4C4CD" box="[1117,1252,1129,1154]" italics="true" pageId="7" pageNumber="58">Careospina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from which it would then be distinguished by the heavy coloration of the wing, the asymmetrical MP fork, and male eyes which meet dorsally.
|
|||
|
|
<bibRefCitation id="EFCF4B5C0335FFCBFB0A3FDCF72CC49C" author="Staniczek" box="[881,1084,1209,1235]" pageId="7" pageNumber="60" refString="Staniczek, A. H. (2003) New fossil mayflies from Dominican amber (Insecta: Ephemeroptera: Leptophlebiidae: Atalophlebiinae). Stuttgarter Beitrage zur Naturkunde, B, 341, 1 - 22." type="journal article" year="2003">Staniczek (2003)</bibRefCitation>
|
|||
|
|
described one new genus (
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBF9183F84F2E0C4B5" box="[355,496,1249,1274]" class="Insecta" family="Leptophlebiidae" genus="Hagenulites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBF9183F84F2E0C4B5" box="[355,496,1249,1274]" italics="true" pageId="7" pageNumber="58">Hagenulites</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) and three new species of the genus
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFBB73F84F746C4B5" box="[972,1110,1249,1274]" class="Insecta" family="Leptophlebiidae" genus="Borinquena" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFBB73F84F746C4B5" box="[972,1110,1249,1274]" italics="true" pageId="7" pageNumber="58">Borinquena</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from Dominican amber (Tertiary: Eocene to Miocene); genera and species described by Staniczek all possess a symmetrical MP fork. The asymmetrical MP fork in
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFB983E54F7CDC505" box="[995,1245,1329,1354]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFB983E54F7CDC505" box="[995,1245,1329,1354]" italics="true" pageId="7" pageNumber="58">Hagenulus marshalli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A21957C40335FFCBFC933E54F63BC504" box="[1256,1323,1329,1355]" pageId="7" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
eventually may indicate a new genus, but it seems best to delay such action until females and nymphs are known. At present, we prefer to retain described species of
|
|||
|
|
<taxonomicName id="4C5E4D2E0335FFCBFCF73EE4F61BC5D5" box="[1164,1291,1409,1434]" class="Insecta" family="Leptophlebiidae" genus="Hagenulus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92AEABF0335FFCBFCF73EE4F61BC5D5" box="[1164,1291,1409,1434]" italics="true" pageId="7" pageNumber="58">Hagenulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in this, the oldest genus, and to treat all of Kluge’s (1994) subgenera as genera until more extensive revisionary work is completed.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|