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<document id="EFCBD222A49FD9424269B9039CD47486" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55574FF90FF4AF979FCE6BB5A" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus mokaensis Bernhauer 1915, comb. nov." docType="treatment" docVersion="7" lastPageNumber="82" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="80" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="6FEE80A6511C88DA1CBFB706156143A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4719A319D7FDC52E7650E6209BC56021">
<mods:title id="014C01B9F3B1B7C52A2BDA7E665493F6">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="C4C604F68B9B473C0DEA626D13310133" type="personal">
<mods:role id="D5097E7192977C69A90352E3F8D1C1E6">
<mods:roleTerm id="1599D1C91F9CA29359C36FD108D86FB6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A252AAA9F4F13DD863A51B315B1256A">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="9CF9F7B25A65D76936A618273C76C4E9" type="personal">
<mods:role id="27F574175222B2D1DE242F860DFE4C8F">
<mods:roleTerm id="575BA429A16714F925B36DCAD33ACEE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2134CB1A260C4C6344CA513EF22B3018">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="77290AA4337ACF79D22439B90A102491">text</mods:typeOfResource>
<mods:relatedItem id="2BFE8F7C74C7F97CFE660B68D6AC0CE1" type="host">
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<mods:title id="03BEE2105FE529AA9D1D54BBA2DB9B09">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ECF1212A84686CF2ED4EC63947C01B38">
<mods:date id="EA175C3373FA0682B8E858BAA7319BD6">2015</mods:date>
<mods:detail id="8FD32587E74AC32E9CED5C80E73D804D" type="volume">
<mods:number id="BBBEEE37B2E1B289B6B43B8177D7F76B">4038</mods:number>
</mods:detail>
<mods:detail id="54E7E999AD0B34E7673B00513C7192D9" type="issue">
<mods:number id="2E4465A8F20F4C99D3B29FAAC88AE335">1</mods:number>
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<mods:start id="5BDA5C81BA00E0D1E9E0BF6346CFCA26">1</mods:start>
<mods:end id="41CCBE19CBC204A6F8F93B3FC0188E37">94</mods:end>
</mods:extent>
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<mods:classification id="0204923A13B4B008A2BF29A0AF7EEFAB">journal article</mods:classification>
<mods:identifier id="9ABD547445270270E416C69608C199B1" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="1845EFC5F14076BE0D4774729CA4066D" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
<mods:identifier id="DEB19E8C2EAC636DEA5950D84688A3AD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A3BAB7631E719C9ACC047584C8B133D5" type="Zenodo-Dep">289876</mods:identifier>
<mods:identifier id="115967FA555302C2463395FE1C5DB317" type="ZooBank">1B62B78C-AA59-4417-A4FC-1CC9CED745E0</mods:identifier>
</mods:mods>
<treatment id="039D87F55574FF90FF4AF979FCE6BB5A" ID-DOI="http://doi.org/10.5281/zenodo.6101703" ID-GBIF-Taxon="127693930" ID-Zenodo-Dep="6101703" LSID="urn:lsid:plazi:treatment:039D87F55574FF90FF4AF979FCE6BB5A" httpUri="http://treatment.plazi.org/id/039D87F55574FF90FF4AF979FCE6BB5A" lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80">
<subSubSection id="C32E65685574FF8EFF4AF979FE46B93E" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="8B8B36E35574FF8EFF4AF979FCA3B8DC" blockId="79.[151,860,1759,1819]" box="[151,860,1759,1785]" pageId="79" pageNumber="80">
<heading id="D0C3818F5574FF8EFF4AF979FCA3B8DC" bold="true" box="[151,860,1759,1785]" fontSize="11" level="1" pageId="79" pageNumber="80" reason="1">
<emphasis id="B940EAF15574FF8EFF4AF979FCA3B8DC" bold="true" box="[151,860,1759,1785]" pageId="79" pageNumber="80">
<taxonomicName id="4C344D605574FF8EFF4AF979FD37B8DC" ID-CoL="7C9FQ" authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[151,712,1759,1785]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mokaensis" status="comb. nov.">
<emphasis id="B940EAF15574FF8EFF4AF979FE20B8DC" bold="true" box="[151,479,1759,1785]" italics="true" pageId="79" pageNumber="80">Thyreocephalus mokaensis</emphasis>
(Bernhauer, 1915)
</taxonomicName>
,
<taxonomicNameLabel id="A273578A5574FF8EFD0BF979FCA3B8DC" box="[726,860,1759,1785]" pageId="79" pageNumber="80">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36E35574FF8EFF4AF8A4FE46B93E" blockId="79.[151,860,1759,1819]" box="[151,441,1794,1819]" pageId="79" pageNumber="80">(Figs. 170, 171, 206–211)</paragraph>
</subSubSection>
<subSubSection id="C32E65685574FF8EFF4AF8ECFC4FB9B8" pageId="79" pageNumber="80" type="reference_group">
<paragraph id="8B8B36E35574FF8EFF4AF8ECFC66B95B" blockId="79.[151,1071,1866,1949]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="AB2411CD5574FF8EFF4AF8ECFC66B95B" pageId="79" pageNumber="80">
<taxonomicName id="4C344D605574FF8EFF4AF8ECFDB4B945" ID-CoL="3CBNL" authority="Bernhauer, 1915a: 299" authorityName="Bernhauer" authorityPageNumber="299" authorityYear="1915" box="[151,587,1866,1888]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF15574FF8EFF4AF8ECFEA4B945" box="[151,347,1866,1888]" italics="true" pageId="79" pageNumber="80">Eulissus mokaensis</emphasis>
<treatmentCitation id="0A9510F25574FF8EFEBCF8ECFDB4B945" author="Bernhauer" box="[353,587,1866,1888]" page="299" pageId="79" pageNumber="80" year="1915">
<bibRefCitation id="EFA54B125574FF8EFEBCF8ECFDE7B945" author="Bernhauer" box="[353,536,1866,1888]" pageId="79" pageNumber="80" refString="Bernhauer, M. (1915 a) Neue Staphyliniden des tropischen Afrika. Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien, 65, 287 - 321." type="journal article" year="1915" yearSuffix="a">Bernhauer, 1915a</bibRefCitation>
: 299
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25574FF8EFD88F8ECFCB1B945" author="Scheerpeltz" box="[597,846,1866,1888]" page="1317" pageId="79" pageNumber="80" year="1933">
<bibRefCitation id="EFA54B125574FF8EFD88F8ECFCF1B945" author="Scheerpeltz" box="[597,782,1866,1888]" pageId="79" pageNumber="80" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1317
</treatmentCitation>
;
<treatmentCitation id="0A9510F25574FF8EFC85F8EDFBD3B945" author="Herman" box="[856,1068,1866,1888]" page="3628" pageId="79" pageNumber="80" year="2001">
<bibRefCitation id="EFA54B125574FF8EFC85F8EDFC13B945" author="Herman" box="[856,1004,1866,1888]" pageId="79" pageNumber="80" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3628
</treatmentCitation>
.
<taxonomicName id="4C344D605574FF8EFF4AF8CEFDABB95B" ID-CoL="3CBN9" authority="Bernhauer, 1939b: 259" authorityName="Bernhauer" authorityPageNumber="259" authorityYear="1939" box="[151,596,1896,1918]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="flaviventris">
<emphasis id="B940EAF15574FF8EFF4AF8CEFE9BB95B" box="[151,356,1896,1918]" italics="true" pageId="79" pageNumber="80">Eulissus flaviventris</emphasis>
<treatmentCitation id="0A9510F25574FF8EFEB4F8CEFDABB95B" author="Bernhauer" box="[361,596,1896,1918]" page="259" pageId="79" pageNumber="80" year="1939">
<bibRefCitation id="EFA54B125574FF8EFEB4F8CEFDDDB95B" author="Bernhauer" box="[361,546,1896,1918]" pageId="79" pageNumber="80" refString="Bernhauer, M. (1939 b) Zur Staphylinidenfauna von Kamerun. Entomologische Blatter, 35 (5), 252 - 262." type="journal article" year="1939" yearSuffix="b">Bernhauer, 1939b</bibRefCitation>
: 259
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25574FF8EFD83F8CFFCCEB95B" author="Herman" box="[606,817,1896,1918]" page="3627" pageId="79" pageNumber="80" year="2001">
<bibRefCitation id="EFA54B125574FF8EFD83F8CFFD0CB95B" author="Herman" box="[606,755,1896,1918]" pageId="79" pageNumber="80" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3627
</treatmentCitation>
,
<emphasis id="B940EAF15574FF8EFCE1F8CEFC66B95B" bold="true" box="[828,921,1896,1918]" pageId="79" pageNumber="80">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36E35574FF8EFF4AF821FC4FB9B8" blockId="79.[151,1071,1866,1949]" box="[151,944,1927,1949]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="AB2411CD5574FF8EFF4AF821FC4FB9B8" box="[151,944,1927,1949]" pageId="79" pageNumber="80">
<taxonomicName id="4C344D605574FF8EFF4AF821FD94B9B8" ID-CoL="56RYP" authority="Coiffait, 1968: 155" authorityName="Coiffait" authorityPageNumber="155" authorityYear="1968" box="[151,619,1927,1949]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis id="B940EAF15574FF8EFF4AF821FE5EB9B8" box="[151,417,1927,1949]" italics="true" pageId="79" pageNumber="80">Thyreocephalus semirufus</emphasis>
<treatmentCitation id="0A9510F25574FF8EFE7AF821FD94B9B8" author="Coiffait" box="[423,619,1927,1949]" page="155" pageId="79" pageNumber="80" year="1968">
<bibRefCitation id="EFA54B125574FF8EFE7AF821FDC7B9B8" author="Coiffait" box="[423,568,1927,1949]" pageId="79" pageNumber="80" refString="Coiffait, H. (1968) Contribution a la connaissance des Xantholininae (Coleopteres Staphylinidae) du Gabon. Biologia Gabonica, 4 (2), 127 - 161." type="journal article" year="1968">Coiffait, 1968</bibRefCitation>
: 155
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25574FF8EFDA8F82EFCB7B9B8" author="Herman" box="[629,840,1927,1949]" page="3767" pageId="79" pageNumber="80" year="2001">
<bibRefCitation id="EFA54B125574FF8EFDA8F82EFCF6B9B8" author="Herman" box="[629,777,1927,1949]" pageId="79" pageNumber="80" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3767
</treatmentCitation>
,
<emphasis id="B940EAF15574FF8EFC8FF821FC4FB9B8" bold="true" box="[850,944,1927,1949]" pageId="79" pageNumber="80">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E65685574FF90FF4AF86CFE5DBD68" lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="8B8B36E35574FF8EFF4AF86CFDDEB9C6" blockId="79.[151,545,1994,2019]" box="[151,545,1994,2019]" pageId="79" pageNumber="80">
<emphasis id="B940EAF15574FF8EFF4AF86CFECAB9C6" bold="true" box="[151,309,1994,2019]" pageId="79" pageNumber="80">
<typeStatus id="548F88415574FF8EFF4AF86CFF2DB9C6" box="[151,210,1994,2019]" pageId="79" pageNumber="80">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32376735574FF8EFEE1F86CFE32B9C7" box="[316,461,1994,2019]" name="Equatorial Guinea" pageId="79" pageNumber="80">Fernando Po</collectingCountry>
, Moka.
</paragraph>
<caption id="DF4B666B556BFF91FF4AF814FE30B9C2" httpUri="https://zenodo.org/record/289925/files/figure.png" pageId="80" pageNumber="81" targetBox="[214,1378,201,1943]" targetPageId="80">
<paragraph id="8B8B36E3556BFF91FF4AF814FE30B9C2" blockId="80.[151,1436,1970,2023]" pageId="80" pageNumber="81">
<emphasis id="B940EAF1556BFF91FF4AF814FE9AB9ED" bold="true" box="[151,357,1970,1992]" pageId="80" pageNumber="81">FIGURES 207–211</emphasis>
.
<taxonomicName id="4C344D60556BFF91FEACF814FCFAB9ED" authority="Bernhauer" authorityName="Bernhauer" box="[369,773,1970,1992]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556BFF91FEACF814FD7EB9ED" box="[369,641,1970,1992]" italics="true" pageId="80" pageNumber="81">Thyreocephalus mokaensis</emphasis>
(Bernhauer)
</taxonomicName>
; 207, head and pronotum; 208, labrum; 209, male tergite 10; 210, male sternite 9; 211, aedeagus.
</paragraph>
</caption>
<paragraph id="8B8B36E3556AFF90FF1AFF31FD74BFC8" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFF31FE0FBE95" bold="true" box="[199,496,151,176]" pageId="81" pageNumber="82">
<typeStatus id="548F8841556AFF90FF1AFF31FEFDBE95" box="[199,258,151,176]" pageId="81" pageNumber="82">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="4C344D60556AFF90FE2AFF31FD2FBE95" box="[503,720,151,176]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FE2AFF31FD2FBE95" box="[503,720,151,176]" italics="true" pageId="81" pageNumber="82">Eulissus mokaensis</emphasis>
</taxonomicName>
.
<typeStatus id="548F8841556AFF90FD02FF31FCAABE95" box="[735,853,151,176]" pageId="81" pageNumber="82" type="lectotype">
<emphasis id="B940EAF1556AFF90FD02FF31FCAABE95" bold="true" box="[735,853,151,176]" pageId="81" pageNumber="82">Lectotype</emphasis>
</typeStatus>
(by present designation) ♂:
<collectingCountry id="F3237673556AFF90FB45FF31FA9ABE95" box="[1176,1381,151,176]" name="Equatorial Guinea" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FB45FF31FA9ABE95" bold="true" box="[1176,1381,151,176]" pageId="81" pageNumber="82">FERNANDO PO</emphasis>
</collectingCountry>
: “Is.
<collectingCountry id="F3237673556AFF90FF4AFF1DFEC6BEF6" box="[151,313,187,212]" name="Equatorial Guinea" pageId="81" pageNumber="82">Fernando Poo</collectingCountry>
, Moka,
<date id="FF8A1023556AFF90FE46FF1DFE12BEF6" box="[411,493,187,211]" pageId="81" pageNumber="82" value="1902-02">II.1902</date>
L. Fea,
<quantity id="4CCC9B06556AFF90FD89FF1DFD09BEF6" box="[596,758,187,212]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="81" pageNumber="82" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</quantity>
”, “
<typeStatus id="548F8841556AFF90FCC6FF1DFC9ABEF6" box="[795,869,187,211]" pageId="81" pageNumber="82">Typus</typeStatus>
”, “
<taxonomicName id="4C344D60556AFF90FC56FF1DFB9EBEF1" authority="Bernh." authorityName="Bernh." box="[907,1121,187,212]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FC56FF1DFC00BEF1" box="[907,1023,187,212]" italics="true" pageId="81" pageNumber="82">mokaensis</emphasis>
, Bernh.
</taxonomicName>
”, “
<taxonomicName id="4C344D60556AFF90FB5AFF1DFAC0BEF1" authority="Brh." authorityName="Brh." box="[1159,1343,187,212]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis" status="sp. nov.">
<emphasis id="B940EAF1556AFF90FB5AFF1DFB03BEF1" box="[1159,1276,187,212]" italics="true" pageId="81" pageNumber="82">mokaensis</emphasis>
, Brh.
</taxonomicName>
<taxonomicNameLabel id="A273578A556AFF90FA94FF1AFA75BEF6" box="[1353,1418,187,211]" pageId="81" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
”, “
<typeStatus id="548F8841556AFF90FF7CFF79FEF1BED2" box="[161,270,223,247]" pageId="81" pageNumber="82" type="syntype">Syntypus</typeStatus>
,
<emphasis id="B940EAF1556AFF90FEC6FF78FE03BED2" box="[283,508,222,247]" italics="true" pageId="81" pageNumber="82">
<taxonomicName id="4C344D60556AFF90FEC6FF78FE84BED2" box="[283,379,222,247]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">Eulissus</taxonomicName>
,
<taxonomicName id="4C344D60556AFF90FE55FF78FE03BED2" authority="Bernhauer" authorityName="Bernhauer" box="[392,508,222,247]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">mokaensis</taxonomicName>
</emphasis>
, Bernhauer” (
<collectionCode id="ED25AE26556AFF90FD43FF79FCF7BED2" box="[670,776,223,247]" pageId="81" pageNumber="82">MCSNG</collectionCode>
). This specimen is chosen as
<typeStatus id="548F8841556AFF90FB87FF78FB3BBED2" box="[1114,1220,222,247]" pageId="81" pageNumber="82" type="lectotype">lectotype</typeStatus>
. The specimen has additional label “
<typeStatus id="548F8841556AFF90FE8AFEA4FE29BF3F" box="[343,470,258,282]" pageId="81" pageNumber="82" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="4C344D60556AFF90FE00FEA7FCAEBF3F" authority="Bh., Bordoni" authorityName="Bh., Bordoni" box="[477,849,257,282]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FE00FEA7FD48BF3F" box="[477,695,257,282]" italics="true" pageId="81" pageNumber="82">Eulissus mokaensis</emphasis>
Bh., Bordoni
</taxonomicName>
des. 2006”.
<typeStatus id="548F8841556AFF90FC3DFEA7FB6FBF3F" box="[992,1168,257,282]" pageId="81" pageNumber="82" type="paralectotype">
<emphasis id="B940EAF1556AFF90FC3DFEA7FB6FBF3F" bold="true" box="[992,1168,257,282]" pageId="81" pageNumber="82">Paralectotypes</emphasis>
</typeStatus>
: 7 spec.:
<collectingCountry id="F3237673556AFF90FB20FEA7FF41BF18" name="Equatorial Guinea" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FB20FEA7FF41BF18" bold="true" pageId="81" pageNumber="82">FERNANDO PO</emphasis>
</collectingCountry>
: “Is.
<collectingCountry id="F3237673556AFF90FED5FE82FE52BF19" box="[264,429,292,317]" name="Equatorial Guinea" pageId="81" pageNumber="82">Fernando Poo</collectingCountry>
, Moka,
<date id="FF8A1023556AFF90FDC9FE83FD99BF18" box="[532,614,293,317]" pageId="81" pageNumber="82" value="1902-02">II.1902</date>
L. Fea,
<quantity id="4CCC9B06556AFF90FD0BFE82FC85BF18" box="[726,890,292,317]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="81" pageNumber="82" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</quantity>
”, “
<typeStatus id="548F8841556AFF90FC7FFE83FBF0BF18" box="[930,1039,293,317]" pageId="81" pageNumber="82" type="syntype">Syntypus</typeStatus>
,
<taxonomicName id="4C344D60556AFF90FBFDFE82FB82BF18" box="[1056,1149,292,317]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FBFDFE82FB82BF18" box="[1056,1149,292,317]" italics="true" pageId="81" pageNumber="82">Eulissus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D60556AFF90FB4FFE82FA6DBF18" authority="Bernhauer" authorityName="Bernhauer" box="[1170,1426,292,317]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FB4FFE82FAF9BF18" box="[1170,1286,292,317]" italics="true" pageId="81" pageNumber="82">mokaensis</emphasis>
, Bernhauer
</taxonomicName>
” (
<collectionCode id="ED25AE26556AFF90FF43FEEEFEF7BF45" box="[158,264,328,352]" pageId="81" pageNumber="82">MCSNG</collectionCode>
). These specimens are labelled “
<typeStatus id="548F8841556AFF90FD45FEE1FCC0BF45" box="[664,831,327,352]" pageId="81" pageNumber="82" type="paralectotype">Paralectotypus</typeStatus>
<taxonomicName id="4C344D60556AFF90FC93FEE1FB29BF45" authority="Bh., Bordoni" authorityName="Bh., Bordoni" box="[846,1238,327,352]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FC93FEE1FBD1BF45" box="[846,1070,327,352]" italics="true" pageId="81" pageNumber="82">Eulissus mokaensis</emphasis>
Bh., Bordoni
</taxonomicName>
des. 2006”. All precendent
<typeStatus id="548F8841556AFF90FEC4FECAFEB6BFA1" box="[281,329,364,388]" pageId="81" pageNumber="82">type</typeStatus>
specimens are labelled “
<taxonomicName id="4C344D60556AFF90FDB9FECDFC2ABFA1" authority="Bh." authorityName="Bh." box="[612,981,363,388]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FDB9FECDFC6DBFA1" box="[612,914,363,388]" italics="true" pageId="81" pageNumber="82">Thyreocephalus mokaensis</emphasis>
(Bh.)
</taxonomicName>
, Bordoni det. 2006”. 2 ♀: “Is.
<collectingCountry id="F3237673556AFF90FAEFFECDFF3ABF83" name="Equatorial Guinea" pageId="81" pageNumber="82">Fernando Poo</collectingCountry>
, Moka,
<date id="FF8A1023556AFF90FEF8FE29FE86BF82" box="[293,377,399,423]" pageId="81" pageNumber="82" value="1902-02">II.1902</date>
,
<quantity id="4CCC9B06556AFF90FE55FE28FDD7BF83" box="[392,552,398,423]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="81" pageNumber="82" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</quantity>
, L. Fea”, “
<taxonomicName id="4C344D60556AFF90FD76FE28FBCDBF82" authority="Brh., Cotypus" authorityName="Brh., Cotypus" box="[683,1074,398,423]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FD76FE28FC77BF82" box="[683,904,398,423]" italics="true" pageId="81" pageNumber="82">Eulissus mokaensis</emphasis>
Brh., Cotypus
</taxonomicName>
” (
<collectionCode id="ED25AE26556AFF90FB93FE28FB5CBF82" LSID="urn:lsid:biocol.org:col:34795" box="[1102,1187,398,423]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="81" pageNumber="82">FMNH</collectionCode>
).These two specimen were labelled “
<typeStatus id="548F8841556AFF90FE92FE17FE09BFEF" box="[335,502,433,458]" pageId="81" pageNumber="82" type="paralectotype">Paralectotypus</typeStatus>
<taxonomicName id="4C344D60556AFF90FDDBFE17FBF9BFEF" authority="Bh., Janak &amp; Bordoni" authorityName="Bh., Janak &amp; Bordoni" box="[518,1030,433,458]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FDDBFE17FD17BFEF" box="[518,744,433,458]" italics="true" pageId="81" pageNumber="82">Eulissus mokaensis</emphasis>
Bh., Janák &amp; Bordoni
</taxonomicName>
des. 2014” and “
<taxonomicName id="4C344D60556AFF90FB37FE17FDF7BFC8" authority="Bh., Janak &amp; Bordoni" authorityName="Bh., Janak &amp; Bordoni" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FB37FE17FEF3BFC8" italics="true" pageId="81" pageNumber="82">Thyreocephalus mokaensis</emphasis>
Bh., Janák &amp; Bordoni
</taxonomicName>
det. 2014”.
</paragraph>
<paragraph id="8B8B36E3556AFF90FF1AFE51FAF1BC72" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<taxonomicName id="4C344D60556AFF90FF1AFE51FE4FBC35" box="[199,432,503,528]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="flaviventris">
<emphasis id="B940EAF1556AFF90FF1AFE51FE4FBC35" box="[199,432,503,528]" italics="true" pageId="81" pageNumber="82">Eulissus flaviventris</emphasis>
</taxonomicName>
.
<typeStatus id="548F8841556AFF90FE19FE51FDCEBC35" box="[452,561,503,528]" pageId="81" pageNumber="82" type="holotype">
<emphasis id="B940EAF1556AFF90FE19FE51FDCEBC35" bold="true" box="[452,561,503,528]" pageId="81" pageNumber="82">Holotype</emphasis>
</typeStatus>
♀:
<collectingCountry id="F3237673556AFF90FDB5FE51FCEEBC35" box="[616,785,503,528]" name="Cameroon" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FDB5FE51FCEEBC35" bold="true" box="[616,785,503,528]" pageId="81" pageNumber="82">CAMEROON</emphasis>
</collectingCountry>
: “Umg. Kamerunberg, Ekona, 5.–
<date id="FF8A1023556AFF90FB74FE51FAD1BC35" box="[1193,1326,503,528]" pageId="81" pageNumber="82" value="1935-11-10">11.10.1935</date>
”, “Dr. F. Zumpt leg., Eing. Nr. 7, 1936”, “
<taxonomicName id="4C344D60556AFF90FDD5FDBDFD2BBC11" authority="Brnh." authorityName="Brnh." box="[520,724,539,564]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="flaviventris">
<emphasis id="B940EAF1556AFF90FDD5FDBDFD79BC11" box="[520,646,539,564]" italics="true" pageId="81" pageNumber="82">flaviventris</emphasis>
, Brnh.
</taxonomicName>
<typeStatus id="548F8841556AFF90FD06FDBAFCDDBC11" box="[731,802,540,564]" pageId="81" pageNumber="82">Typus</typeStatus>
, unic.
<taxonomicName id="4C344D60556AFF90FCB6FDBDFC37BC11" box="[875,968,539,564]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FCB6FDBDFC37BC11" box="[875,968,539,564]" italics="true" pageId="81" pageNumber="82">Eulissus</emphasis>
</taxonomicName>
” (
<collectionCode id="ED25AE26556AFF90FC39FDBDFBC4BC16" LSID="urn:lsid:biocol.org:col:34795" box="[996,1083,539,563]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="81" pageNumber="82">FMNH</collectionCode>
). It is identical to
<taxonomicName id="4C344D60556AFF90FAD9FDBAFA6BBC11" box="[1284,1428,539,564]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FAD9FDBAFA6BBC11" box="[1284,1428,539,564]" italics="true" pageId="81" pageNumber="82">T. mokaensis</emphasis>
</taxonomicName>
. The label “
<taxonomicName id="4C344D60556AFF90FECEFD98FCC2BC72" authority="Bh., Janak &amp; Bordoni" authorityName="Bh., Janak &amp; Bordoni" box="[275,829,574,599]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FECEFD98FDBEBC72" box="[275,577,574,599]" italics="true" pageId="81" pageNumber="82">Thyreocephalus mokaensis</emphasis>
Bh., Janák &amp; Bordoni
</taxonomicName>
det. 2014” was attached to the specimen.
</paragraph>
<paragraph id="8B8B36E3556AFF90FF28FDC7FB64BD22" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<taxonomicName id="4C344D60556AFF90FF28FDC7FDD8BC5F" box="[245,551,609,634]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis id="B940EAF1556AFF90FF28FDC7FDD8BC5F" box="[245,551,609,634]" italics="true" pageId="81" pageNumber="82">Thyreocephalus semirufus</emphasis>
</taxonomicName>
.
<typeStatus id="548F8841556AFF90FD9DFDC7FD52BC5F" box="[576,685,609,634]" pageId="81" pageNumber="82" type="holotype">
<emphasis id="B940EAF1556AFF90FD9DFDC7FD52BC5F" bold="true" box="[576,685,609,634]" pageId="81" pageNumber="82">Holotype</emphasis>
</typeStatus>
♂:
<collectingCountry id="F3237673556AFF90FD32FDC7FCA9BC5F" box="[751,854,609,634]" name="Gabon" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FD32FDC7FCA9BC5F" bold="true" box="[751,854,609,634]" pageId="81" pageNumber="82">GABON</emphasis>
</collectingCountry>
: “Belinga 29,
<date id="FF8A1023556AFF90FBC0FDC7FB8BBC5F" box="[1053,1140,609,634]" pageId="81" pageNumber="82" value="63-01-23">23.1.63</date>
, H. Coiffait”, “Mission biologique, au
<collectingCountry id="F3237673556AFF90FE9CFD22FE71BCB8" box="[321,398,644,669]" name="Gabon" pageId="81" pageNumber="82">Gabon</collectingCountry>
, P. P. Grassé Directeur”, “
<typeStatus id="548F8841556AFF90FD64FD22FCD7BCB8" box="[697,808,644,669]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
”, “
<taxonomicName id="4C344D60556AFF90FC91FD22FC01BCB8" box="[844,1022,644,669]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FC91FD22FC01BCB8" box="[844,1022,644,669]" italics="true" pageId="81" pageNumber="82">Thyreocephalus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D60556AFF90FBD0FD23FA6EBCB8" authority="Coiff., H. Coiffait 1967" authorityName="Coiff., H. Coiffait" authorityYear="1967" box="[1037,1425,644,669]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis id="B940EAF1556AFF90FBD0FD23FB85BCB8" box="[1037,1146,645,669]" italics="true" pageId="81" pageNumber="82">semirufus</emphasis>
Coiff., H. Coiffait 1967
</taxonomicName>
” (
<collectionCode id="ED25AE26556AFF90FF43FD0EFEF5BCE5" box="[158,266,680,704]" pageId="81" pageNumber="82">MNHNP</collectionCode>
).
<typeStatus id="548F8841556AFF90FEC0FD01FE6BBCE5" box="[285,404,679,704]" pageId="81" pageNumber="82" type="paratype">
<emphasis id="B940EAF1556AFF90FEC0FD01FE6BBCE5" bold="true" box="[285,404,679,704]" pageId="81" pageNumber="82">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D32FD6A556AFF90FE79FD0EFE30BC9A" box="[420,463,680,704]" pageId="81" pageNumber="82" type="generic">3 ♂</specimenCount>
, 1 ♀:
<collectingCountry id="F3237673556AFF90FDCAFD01FD81BCE5" box="[535,638,679,704]" name="Gabon" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FDCAFD01FD81BCE5" bold="true" box="[535,638,679,704]" pageId="81" pageNumber="82">GABON</emphasis>
</collectingCountry>
: “Belinga 29,
<date id="FF8A1023556AFF90FCFFFD01FC86BCE5" box="[802,889,679,704]" pageId="81" pageNumber="82" value="63-01-23">23.1.63</date>
, H. Coiffait”, “Mission biologique, au
<collectingCountry id="F3237673556AFF90FAEDFD01FA82BCE5" box="[1328,1405,679,704]" name="Gabon" pageId="81" pageNumber="82">Gabon</collectingCountry>
, P. P. Grassé Directeur”, “
<typeStatus id="548F8841556AFF90FE79FD6DFDF2BCC6" box="[420,525,715,739]" pageId="81" pageNumber="82" type="paratype">Paratype</typeStatus>
”, “
<taxonomicName id="4C344D60556AFF90FDE8FD6DFD18BCC1" box="[565,743,715,740]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FDE8FD6DFD18BCC1" box="[565,743,715,740]" italics="true" pageId="81" pageNumber="82">Thyreocephalus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D60556AFF90FD26FD6DFB70BCC1" authority="Coiff., H. Coiffait 1967" authorityName="Coiff., H. Coiffait" authorityYear="1967" box="[763,1167,715,740]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis id="B940EAF1556AFF90FD26FD6DFC97BCC6" box="[763,872,715,739]" italics="true" pageId="81" pageNumber="82">semirufus</emphasis>
Coiff., H. Coiffait 1967
</taxonomicName>
” (
<collectionCode id="ED25AE26556AFF90FB73FD6DFAE5BCC6" box="[1198,1306,715,739]" pageId="81" pageNumber="82">MNHNP</collectionCode>
). All these specimens are labelled also “
<taxonomicName id="4C344D60556AFF90FE01FD48FCB2BD22" authority="Bh." authorityName="Bh." box="[476,845,750,775]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FE01FD48FCF5BD22" box="[476,778,750,775]" italics="true" pageId="81" pageNumber="82">Thyreocephalus mokaensis</emphasis>
(Bh.)
</taxonomicName>
, Janák &amp; Bordoni det, 2014”.
</paragraph>
<paragraph id="8B8B36E3556AFF90FF1AFCB7FE5DBD68" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFCB7FDC7BD0F" bold="true" box="[199,568,785,810]" pageId="81" pageNumber="82">Additional material examined</emphasis>
.
<collectingCountry id="F3237673556AFF90FD96FCB7FCE1BD0F" box="[587,798,785,810]" name="Equatorial Guinea" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FD96FCB7FCE1BD0F" bold="true" box="[587,798,785,810]" pageId="81" pageNumber="82">FERNANDO PO</emphasis>
</collectingCountry>
: 1 spec.:
<collectingCountry id="F3237673556AFF90FC47FCB7FBC3BD0C" box="[922,1084,785,810]" name="Equatorial Guinea" pageId="81" pageNumber="82">Fernando Poo</collectingCountry>
Is., Basilé,
<quantity id="4CCC9B06556AFF90FB0CFCB7FAA8BD0C" box="[1233,1367,785,810]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="81" pageNumber="82" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400–600 m</quantity>
,
<date id="FF8A1023556AFF90FAB4FCB7FF11BD68" pageId="81" pageNumber="82" value="1901-08" valueMax="1901-09-31" valueMin="1901-08-00">viii– ix.1901</date>
, L. Fea (
<collectionCode id="ED25AE26556AFF90FE8DFC93FE69BD68" box="[336,406,821,845]" pageId="81" pageNumber="82">ABFI</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6568556AFF90FF1AFCF1FCAEBAD1" pageId="81" pageNumber="82" type="description">
<paragraph id="8B8B36E3556AFF90FF1AFCF1FE9DBA61" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFCF1FE8BBD55" bold="true" box="[199,372,855,880]" pageId="81" pageNumber="82">Redescription.</emphasis>
Body length
<quantity id="4CCC9B06556AFF90FDD7FCFEFD7BBD4A" box="[522,644,856,880]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="81" pageNumber="82" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B06556AFF90FAB6FCFEFF10BDB6" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="81" pageNumber="82" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.8– 5.8 mm</quantity>
. Head black, pronotum reddish brown, elytra yellowish red, abdomen reddish brown, antennae dark brown; legs light brown (Fig. 170). Body shiny. Head and pronotum with micro-punctation. Head and pronotum and related punctation as in Figs. 171, 207; labrum as in
<figureCitation id="130F2A66556AFF90FD28FC67FCA7BDFF" box="[757,856,961,986]" captionStart="FIGURES 207 – 211" captionStartId="80.[151,264,1970,1992]" captionTargetBox="[214,1378,201,1943]" captionTargetId="figure@80.[198,1388,193,1949]" captionTargetPageId="80" captionText="FIGURES 207 – 211. Thyreocephalus mokaensis (Bernhauer); 207, head and pronotum; 208, labrum; 209, male tergite 10; 210, male sternite 9; 211, aedeagus." httpUri="https://zenodo.org/record/289925/files/figure.png" pageId="81" pageNumber="82">Fig. 208</figureCitation>
. Elytra narrow, dilated posteriad, longer and wider than pronotum, with less marked humeral angles. Surface with fine and sparse punctation, arranged in three series, one near suture, one median and one lateral. Abdomen with fine, transverse micro-striation and fine, deep, not sparse punctation.
</paragraph>
<paragraph id="8B8B36E3556AFF90FF1AFBE8FDAABA88" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<collectingCountry id="F3237673556AFF90FF1AFBE8FEFBBA42" box="[199,260,1102,1127]" name="Malta" pageId="81" pageNumber="82">Male</collectingCountry>
. Tergite 10 and sternite 9 of male genital segment as in
<figureCitation id="130F2A66556AFF90FCA3FBE8FBDCBA42" box="[894,1059,1102,1127]" captionStart="FIGURES 207 – 211" captionStartId="80.[151,264,1970,1992]" captionTargetBox="[214,1378,201,1943]" captionTargetId="figure@80.[198,1388,193,1949]" captionTargetPageId="80" captionText="FIGURES 207 – 211. Thyreocephalus mokaensis (Bernhauer); 207, head and pronotum; 208, labrum; 209, male tergite 10; 210, male sternite 9; 211, aedeagus." httpUri="https://zenodo.org/record/289925/files/figure.png" pageId="81" pageNumber="82">Figs. 209, 210</figureCitation>
. Aedeagus (
<figureCitation id="130F2A66556AFF90FB70FBE8FAF2BA42" box="[1197,1293,1102,1127]" captionStart="FIGURES 207 – 211" captionStartId="80.[151,264,1970,1992]" captionTargetBox="[214,1378,201,1943]" captionTargetId="figure@80.[198,1388,193,1949]" captionTargetPageId="80" captionText="FIGURES 207 – 211. Thyreocephalus mokaensis (Bernhauer); 207, head and pronotum; 208, labrum; 209, male tergite 10; 210, male sternite 9; 211, aedeagus." httpUri="https://zenodo.org/record/289925/files/figure.png" pageId="81" pageNumber="82">Fig. 211</figureCitation>
) ovoid,
<quantity id="4CCC9B06556AFF90FAB6FBE8FF3CBAAC" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="81" pageNumber="82" unit="mm" value="1.95">1.95 mm</quantity>
long, with sub-triangular median lobe; parameres symmetrical; inner sac very narrow and long, folded on itself several times, covered with little scales.
</paragraph>
<paragraph id="8B8B36E3556AFF90FF1AFB11FCAEBAD1" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFB11FE29BAF5" bold="true" box="[199,470,1207,1232]" pageId="81" pageNumber="82">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D60556AFF90FE38FB11FCE4BAF5" box="[485,795,1207,1232]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mokaensis">
<emphasis id="B940EAF1556AFF90FE38FB11FCE4BAF5" box="[485,795,1207,1232]" italics="true" pageId="81" pageNumber="82">Thyreocephalus mokaensis</emphasis>
</taxonomicName>
differs from the similar
<taxonomicName id="4C344D60556AFF90FB85FB1FFB21BAF5" box="[1112,1246,1207,1232]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="burgeoni">
<emphasis id="B940EAF1556AFF90FB85FB1FFB21BAF5" box="[1112,1246,1207,1232]" italics="true" pageId="81" pageNumber="82">T. burgeoni</emphasis>
</taxonomicName>
in the densely punctured disc of head and short median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C32E6568556AFF90FF1AFB58FAFCBB32" box="[199,1283,1278,1303]" pageId="81" pageNumber="82" type="distribution">
<paragraph id="8B8B36E3556AFF90FF1AFB58FAFCBB32" blockId="81.[151,1437,151,1408]" box="[199,1283,1278,1303]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFB58FE9FBB32" bold="true" box="[199,352,1278,1303]" pageId="81" pageNumber="82">Distribution.</emphasis>
The species is distributed in
<collectingCountry id="F3237673556AFF90FD7AFB59FCE0BB32" box="[679,799,1279,1303]" name="Cameroon" pageId="81" pageNumber="82">Cameroon</collectingCountry>
,
<collectingCountry id="F3237673556AFF90FCF4FB58FC47BB33" box="[809,952,1278,1303]" name="Equatorial Guinea" pageId="81" pageNumber="82">Fernando Po</collectingCountry>
<collectingCountry id="F3237673556AFF90FC62FB58FBFCBB32" box="[959,1027,1278,1303]" name="Iceland" pageId="81" pageNumber="82">Island</collectingCountry>
and
<collectingCountry id="F3237673556AFF90FBE4FB58FB7BBB32" box="[1081,1156,1278,1303]" name="Gabon" pageId="81" pageNumber="82">Gabon</collectingCountry>
(
<figureCitation id="130F2A66556AFF90FB4EFB58FB09BB32" box="[1171,1270,1278,1303]" captionStart="FIGURE 206" captionStartId="79.[151,250,1606,1628]" captionTargetBox="[158,1431,199,1578]" captionTargetId="figure@79.[151,1436,193,1584]" captionTargetPageId="79" captionText="FIGURE 206. Distribution of species in Thyreocephalus mocquerysi group (★ = T. atlanticus (Bernhauer), ▲ = T. burgeoni (Bernhauer), ■ = T. camerunensis sp. nov., Δ = T. marginipennis sp. nov., ● = T. mokaensis (Bernhauer), ○ = T. strinatii (Bernhauer)." httpUri="https://zenodo.org/record/289924/files/figure.png" pageId="81" pageNumber="82">Fig. 206</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6568556AFF90FF1AFA87FCE6BB5A" pageId="81" pageNumber="82" type="discussion">
<paragraph id="8B8B36E3556AFF90FF1AFA87FCE6BB5A" blockId="81.[151,1437,151,1408]" pageId="81" pageNumber="82">
<emphasis id="B940EAF1556AFF90FF1AFA87FEC4BB1F" bold="true" box="[199,315,1313,1338]" pageId="81" pageNumber="82">Remarks.</emphasis>
<collectingCountry id="F3237673556AFF90FE99FA84FE9CBB1F" box="[324,355,1314,1338]" name="American Samoa" pageId="81" pageNumber="82">As</collectingCountry>
the superior line of pronotal hypomeron is turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto anterior margin of pronotum, the species was transferred from
<taxonomicName id="4C344D60556AFF90FE02FAC1FDC3BBA5" box="[479,572,1383,1408]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FE02FAC1FDC3BBA5" box="[479,572,1383,1408]" italics="true" pageId="81" pageNumber="82">Eulissus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C344D60556AFF90FDBDFAC1FCEDBBA5" box="[608,786,1383,1408]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1556AFF90FDBDFAC1FCEDBBA5" box="[608,786,1383,1408]" italics="true" pageId="81" pageNumber="82">Thyreocephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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