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<document id= "1DF22AD6C71A883D20FC061340323B9D" ID-DOI= "10.4314/met.v33i1.15" ID-ISSN= "2307-5031" ID-Zenodo-Dep= "14682409" ID-ZooBank= "7CF9077A-E55B-4290-8EFA-A47448254139" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "felipe" IM.tables_requiresApprovalFor= "GgImagineBatch" IM.taxonomicNames_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1737170591275" checkinUser= "felipe" docAuthor= "Aduse-Poku, Kwaku, Lohman, David J. & Richardson, Ian D." docDate= "2022" docId= "03ED3970FFE1D64DF993F86CD26AFEB1" docLanguage= "en" docName= "Metamorphosis.33.130-163.pdf" docOrigin= "Metamorphosis 33 (1)" docSource= "https://doi.org/10.4314/met.v33i1.15" docStyle= "DocumentStyle:73CBAE06DE6A380ACC54A94377F5FCBA.1:Metamorphosis.2021-.journal_article" docStyleId= "73CBAE06DE6A380ACC54A94377F5FCBA" docStyleName= "Metamorphosis.2021-.journal_article" docStyleVersion= "1" docTitle= "Neptis kupe Aduse-Poku, Lohman & Richardson, 2022, sp. nov." docType= "treatment" docVersion= "2" lastPageNumber= "135" masterDocId= "FFD44108FFE5D648F917FFB1D351FFCF" masterDocTitle= "Revision of the genus Neptis Fabricius, 1807 (Papilionoidea: Lepidoptera: Nymphalidae) in the Afrotropical Region, Part 4: The phylogeny of the Nysiades group with eight new species" masterLastPageNumber= "163" masterPageNumber= "130" pageNumber= "134" updateTime= "1737171773407" updateUser= "ExternalLinkService" zenodo-license-document= "CC-BY-NC-ND-4.0" >
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<mods:title id= "3FC644D3E8AB5142ED562EB46B473588" > Revision of the genus Neptis Fabricius, 1807 (Papilionoidea: Lepidoptera: Nymphalidae) in the Afrotropical Region, Part 4: The phylogeny of the Nysiades group with eight new species</mods:title>
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<mods:namePart id= "7AED57EF7E05629564992C0E584E3585" > Aduse-Poku, Kwaku</mods:namePart>
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<mods:namePart id= "A85F140962D1AA08777D78A2F3732AE8" > Lohman, David J.</mods:namePart>
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<mods:namePart id= "7302CB9B4B5091D4068534D546459188" > Richardson, Ian D.</mods:namePart>
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<treatment id= "03ED3970FFE1D64DF993F86CD26AFEB1" ID-DOI= "http://doi.org/10.5281/zenodo.14682397" ID-Zenodo-Dep= "14682397" LSID= "urn:lsid:plazi:treatment:03ED3970FFE1D64DF993F86CD26AFEB1" httpUri= "http://treatment.plazi.org/id/03ED3970FFE1D64DF993F86CD26AFEB1" lastPageId= "5" lastPageNumber= "135" pageId= "4" pageNumber= "134" >
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<subSubSection id= "C35EDBEDFFE1D64CF993F86CD2EBF83A" box= "[132,442,2013,2037]" pageId= "4" pageNumber= "134" type= "nomenclature" >
<paragraph id= "8BFB8866FFE1D64CF993F86CD2EBF83A" blockId= "4.[132,755,1983,2062]" box= "[132,442,2013,2037]" pageId= "4" pageNumber= "134" >
<emphasis id= "B9305474FFE1D64CF993F86CD232F83A" bold= "true" box= "[132,355,2013,2037]" pageId= "4" pageNumber= "134" >
<taxonomicName id= "4C44F3E5FFE1D64CF993F86CD254F83A" authorityName= "Aduse-Poku & Lohman & Richardson" authorityYear= "2022" box= "[132,261,2013,2037]" class= "Insecta" family= "Nymphalidae" genus= "Neptis" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "kupe" status= "sp. nov." >
<emphasis id= "B9305474FFE1D64CF993F86CD254F83A" bold= "true" box= "[132,261,2013,2037]" italics= "true" pageId= "4" pageNumber= "134" > Neptis kupe</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "A203E90FFFE1D64CF81AF86CD232F83A" box= "[269,355,2013,2037]" pageId= "4" pageNumber= "134" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
(
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<figureCitation id= "137F94E3FFE1D64CF865F86CD2E3F83A" box= "[370,434,2013,2037]" captionStart= "Figure 6" captionStartId= "4.[132,200,1608,1629]" captionTargetBox= "[162,1424,117,1575]" captionTargetId= "figure-269@4.[162,1424,117,1575]" captionTargetPageId= "4" captionText= "Figure 6 – figureDoi= "http://doi.org/10.5281/zenodo.14682421" httpUri= "https://zenodo.org/record/14682421/files/figure.png" pageId= "4" pageNumber= "134" > Fig. 6</figureCitation>
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)
</paragraph>
</subSubSection>
<subSubSection id= "C35EDBEDFFE1D64CF993F84BD624F7F7" pageId= "4" pageNumber= "134" type= "description" >
<paragraph id= "8BFB8866FFE1D64CF993F84BD1A2F7C1" blockId= "4.[132,755,1983,2062]" box= "[132,755,2042,2062]" pageId= "4" pageNumber= "134" >
<uri id= "FFD58464FFE1D64CF993F84BD1A2F7C1" box= "[132,755,2042,2062]" pageId= "4" pageNumber= "134" >
urn:lsid:zoobank.org:act:
<uuid id= "FFE2B2B3FFE1D64CF84FF84BD1A2F7C1" box= "[344,755,2042,2062]" pageId= "4" pageNumber= "134" > 89E58A84-519B-4E2D-AC5B-026DFB151772</uuid>
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<paragraph id= "8BFB8866FFE1D64CFA28F961D624F7F7" blockId= "4.[831,1456,1743,2104]" pageId= "4" pageNumber= "134" >
In
<date id= "FFFAAEA6FFE1D64CFA75F961D0A9F928" box= "[866,1016,1743,1768]" pageId= "4" pageNumber= "134" value= "2012-01" > January 2012</date>
, Robert Tropek undertook a collecting expedition to Mount Kupe (Koupé) in West
<collectingCountry id= "F353C8F6FFE1D64CFC04F95CD6D3F8CA" box= "[1299,1410,1773,1797]" name= "Cameroon" pageId= "4" pageNumber= "134" > Cameroon</collectingCountry>
and captured a specimen with facies similar to those of
<taxonomicName id= "4C44F3E5FFE1D64CFC7CF8BCD770F88C" authority= "Rebel 1914" authorityName= "Rebel" authorityYear= "1914" class= "Insecta" family= "Nymphalidae" genus= "Neptis" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "lugubris" >
<emphasis id= "B9305474FFE1D64CFC7CF8BCD0C9F88C" italics= "true" pageId= "4" pageNumber= "134" > Neptis lugubris</emphasis>
<bibRefCitation id= "EFD5F597FFE1D64CFAB6F89AD770F88C" author= "REBEL, H." box= "[929,1057,1835,1859]" pageId= "4" pageNumber= "134" pagination= "219 - 294" refId= "ref25125" refString= "REBEL, H. 1914. Wissenschaftliche Ergebnisse der Expedition R. Grauer nach Zentralafrika, Dezember 1909 bis 1911. Lepidopteren. Annalen des (K. K.) Naturhistorischen Hofmuseums. Wien 28: 219 - 294." type= "journal article" year= "1914" > Rebel 1914</bibRefCitation>
</taxonomicName>
. This expedition was followed up in April the next year by Szabolcs Sáfián, Robert Tropek, and Marianne Espeland who captured further specimens of this
<taxonomicName id= "4C44F3E5FFE1D64CFA28F839D0E9F850" authorityName= "Rebel" authorityYear= "1914" box= "[831,952,1927,1951]" class= "Insecta" family= "Nymphalidae" genus= "Neptis" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "lugubris" >
<emphasis id= "B9305474FFE1D64CFA28F839D0E9F850" box= "[831,952,1927,1951]" italics= "true" pageId= "4" pageNumber= "134" > N. lugubris</emphasis>
</taxonomicName>
-like species. An ABRI-commissioned collector captured a further specimen the following year. Subsequent molecular analysis showed that the Mt. Kupe specimens are distinct from
<taxonomicName id= "4C44F3E5FFE1D64CFD9EF855D650F834" authorityName= "Rebel" authorityYear= "1914" box= "[1161,1281,2019,2043]" class= "Insecta" family= "Nymphalidae" genus= "Neptis" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "lugubris" >
<emphasis id= "B9305474FFE1D64CFD9EF855D650F834" box= "[1161,1281,2019,2043]" italics= "true" pageId= "4" pageNumber= "134" > N. lugubris</emphasis>
</taxonomicName>
, and these are described here as
<taxonomicName id= "4C44F3E5FFE1D64CFAEBF7B3D729F7D6" authorityName= "Aduse-Poku & Lohman & Richardson" authorityYear= "2022" box= "[1020,1144,2049,2073]" class= "Insecta" family= "Nymphalidae" genus= "Neptis" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "kupe" status= "sp. nov." >
<emphasis id= "B9305474FFE1D64CFAEBF7B3D729F7D6" box= "[1020,1144,2049,2073]" italics= "true" pageId= "4" pageNumber= "134" > Neptis kupe</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "A203E90FFFE1D64CFD69F7B3D780F7D6" box= "[1150,1233,2050,2073]" pageId= "4" pageNumber= "134" rank= "species" > sp. nov.</taxonomicNameLabel>
It will be shown that these two species can also be separated by the facies.
</paragraph>
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<materialsCitation id= "3B2C823BFFE0D64DF993FFC6D184FF62" collectingDate= "2014-04" collectionCode= "ABRI" country= "Cameroon" elevation= "1700" latitude= "4.8" location= "Mt. Kupe" longLatPrecision= "1306" longitude= "9.7" pageId= "5" pageNumber= "135" specimenCount= "1" specimenCount-male= "1" typeStatus= "holotype" >
<emphasis id= "B9305474FFE0D64DF993FFC6D3BDFF40" bold= "true" box= "[132,236,119,143]" pageId= "5" pageNumber= "135" >
<typeStatus id= "54FF36C4FFE0D64DF993FFC6D3BDFF40" box= "[132,236,119,143]" pageId= "5" pageNumber= "135" type= "holotype" > Holotype</typeStatus>
</emphasis>
:
<specimenCount id= "9D4243EFFFE0D64DF9E9FFC9D243FF40" box= "[254,274,120,143]" count= "1" pageId= "5" pageNumber= "135" type= "male" > ♂</specimenCount>
<location id= "8E9BDEBDFFE0D64DF80AFFC9D2D8FF40" LSID= "urn:lsid:plazi:treatment:03ED3970FFE1D64DF993F86CD26AFEB1:8E9BDEBDFFE0D64DF80AFFC9D2D8FF40" box= "[285,393,120,143]" country= "Cameroon" latitude= "4.8" longLatPrecision= "1306" longitude= "9.7" name= "Mt. Kupe" pageId= "5" pageNumber= "135" > Mt. Kupe</location>
,
<collectingCountry id= "F353C8F6FFE0D64DF88FFFC6D15BFF40" box= "[408,522,119,143]" name= "Cameroon" pageId= "5" pageNumber= "135" > Cameroon</collectingCountry>
,
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,
<geoCoordinate id= "EE70EEA1FFE0D64DFB9BFFC6D1BEFF40" box= "[652,751,119,143]" degrees= "09" direction= "east" minutes= "42" orientation= "longitude" pageId= "5" pageNumber= "135" precision= "925" value= "9.7" > 09°42ʹ </geoCoordinate>
,
<quantity id= "4CBC2583FFE0D64DF993FF27D24BFF62" box= "[132,282,150,174]" metricMagnitude= "3" metricUnit= "m" metricValue= "1.7" metricValueMax= "1.9" metricValueMin= "1.5" pageId= "5" pageNumber= "135" unit= "m" value= "1700.0" valueMax= "1900.0" valueMin= "1500.0" >
<elevation id= "00696F55FFE0D64DF993FF27D24BFF62" box= "[132,282,150,174]" metricMagnitude= "3" metricUnit= "m" metricValue= "1.7" metricValueMax= "1.9" metricValueMin= "1.5" pageId= "5" pageNumber= "135" unit= "m" value= "1700.0" valueMax= "1900.0" valueMin= "1500.0" > 1500– </elevation>
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,
<date id= "FFFAAEA6FFE0D64DF832FF27D224FF61" box= "[293,373,150,174]" pageId= "5" pageNumber= "135" value= "2014-04" >
<collectingDate id= "EFBE574EFFE0D64DF832FF27D224FF61" box= "[293,373,150,174]" pageId= "5" pageNumber= "135" value= "2014-04" > iv.2014</collectingDate>
</date>
; ABRI-152668,
<collectionCode id= "ED5510A3FFE0D64DFB3DFF27D138FF61" box= "[554,617,150,174]" pageId= "5" pageNumber= "135" > ABRI</collectionCode>
collector.
</materialsCitation>
</paragraph>
<paragraph id= "8BFB8866FFE0D64DF993FF7CD26AFEB1" blockId= "5.[132,757,205,382]" pageId= "5" pageNumber= "135" >
<materialsCitation id= "3B2C823BFFE0D64DF993FF7CD197FECC" collectingDate= "2014-04-07" collectingDateMax= "2014-04-10" collectingDateMin= "2014-04-07" collectionCode= "ANHRT" pageId= "5" pageNumber= "135" specimenCount= "1" specimenCount-male= "1" typeStatus= "paratype" >
<emphasis id= "B9305474FFE0D64DF993FF7CD3A7FF2A" bold= "true" box= "[132,246,205,229]" pageId= "5" pageNumber= "135" >
<typeStatus id= "54FF36C4FFE0D64DF993FF7CD3A7FF2A" box= "[132,246,205,229]" pageId= "5" pageNumber= "135" type= "paratype" > Paratypes</typeStatus>
</emphasis>
:
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same locality,
<date id= "FFFAAEA6FFE0D64DF8A1FF7FD112FF2A" box= "[438,579,205,229]" pageId= "5" pageNumber= "135" value= "2014-04-07" valueMax= "2014-04-10" valueMin= "2014-04-07" >
<collectingDate id= "EFBE574EFFE0D64DF8A1FF7FD112FF2A" box= "[438,579,205,229]" pageId= "5" pageNumber= "135" value= "2014-04-07" valueMax= "2014-04-10" valueMin= "2014-04-07" > 7– </collectingDate>
</date>
, Sáfián, Tropek, Espeland, ANHRTUK00214937,
<collectionCode id= "ED5510A3FFE0D64DF8E0FF5AD102FECC" box= "[503,595,235,259]" pageId= "5" pageNumber= "135" > ANHRT</collectionCode>
collection
</materialsCitation>
;
<materialsCitation id= "3B2C823BFFE0D64DFBCFFF5DD266FEB1" collectingDate= "2012-01" collectingDateMax= "2014-04-10" collectingDateMin= "2012-01" collectionCode= "ANHRT, ABRI" pageId= "5" pageNumber= "135" specimenCount= "1" specimenCount-female= "1" typeStatus= "paratype" >
<specimenCount id= "9D4243EFFFE0D64DFBCFFF5DD1BFFECC" box= "[728,750,236,259]" count= "1" pageId= "5" pageNumber= "135" type= "female" > ♀</specimenCount>
: same locality,
<date id= "FFFAAEA6FFE0D64DF82DFEBBD22CFEED" box= "[314,381,266,290]" pageId= "5" pageNumber= "135" value= "2012-01" >
<collectingDate id= "EFBE574EFFE0D64DF82DFEBBD22CFEED" box= "[314,381,266,290]" pageId= "5" pageNumber= "135" value= "2012-01" > i.2012</collectingDate>
</date>
, Tropek, ANHRTUK00214938,
<collectionCode id= "ED5510A3FFE0D64DF993FE98D3B1FE8E" box= "[132,224,297,321]" pageId= "5" pageNumber= "135" > ANHRT</collectionCode>
collection; 1 unknown sex: same locality,
<date id= "FFFAAEA6FFE0D64DFBCEFE9BD3A7FEAF" pageId= "5" pageNumber= "135" value= "2014-04-07" valueMax= "2014-04-10" valueMin= "2014-04-07" >
<collectingDate id= "EFBE574EFFE0D64DFBCEFE9BD3A7FEAF" pageId= "5" pageNumber= "135" value= "2014-04-07" valueMax= "2014-04-10" valueMin= "2014-04-07" > 7– </collectingDate>
</date>
, Sáfián, Tropek, Espeland, ABRI-152667,
<collectionCode id= "ED5510A3FFE0D64DF993FED7D392FEB1" box= "[132,195,358,382]" pageId= "5" pageNumber= "135" > ABRI</collectionCode>
collection
</materialsCitation>
.
</paragraph>
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</treatment>
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