treatments-xml/data/03/AE/87/03AE87EDFF99CA57FC89A5EBFD84A5D2.xml

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<mods:title id="DB2C3790D070E1A822249BD230262142">Five new species of Phyllophaga (Coleoptera: Melolonthidae) of northwestern Mexico</mods:title>
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<mods:namePart id="503BE647809BB7EF0A97F7696F701DE6">Morón, Miguel Ángel</mods:namePart>
<mods:affiliation id="C15DBAF1F14CD93F254BF71BCE4B8D27">Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, México</mods:affiliation>
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(
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) 
<emphasis id="B973EAE9FF99CA55FBC4A5EBFAA8A7D5" box="[1125,1283,1246,1271]" italics="true" pageId="1" pageNumber="1036">balacachiana</emphasis>
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<taxonomicNameLabel id="A2405792FF99CA55FAABA5D5FACCA7D5" box="[1290,1383,1248,1271]" pageId="1" pageNumber="1036" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="133C2A7EFF99CA55FAD5A5EBFC1DA638" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="1" pageNumber="1036">Figs. 1 and 2, 5–10</figureCitation>
)
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Body length 
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; humeral width 
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. Head reddish yellow, body and legs testaceous light yellowish (
<figureCitation id="133C2A7EFF99CA55FB41A4B0FA93A6BB" box="[1248,1336,1413,1433]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="1" pageNumber="1036">Figs. 1–2</figureCitation>
). Clypeus wider than long (2.5:1.0), anterior border slightly curved, with margin clearly elevated, surface slightly convex at center, covered with punctures,glabrous. Frons wider than long (2.4:1.0), slightly concave, with irregularly distributed punctures, glabrous, 3.2 
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wider than each eye’s dorsal diameter. Canthus with 10 setae. Vertex with a transverse carinate ridge. Occipital area without punctures. Antenna 10-segmented; with club 1.1 
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longer than preceding 6 segments combined.
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Pronotum wider than long (1.6:1.0) and 2.4 
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wider than frons, but only 1.5 
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wider than head including eyes. Pronotal disc with shallow, small punctures regularly separated and distributed on disc, but less dense near posterior half of midline; lateral margins obtusely angled, widest just behind middle, lateral marginal bead irregularly crenulate, with long, slender setae; anterior marginal bead without setae; anterior angles obtuse, not prominent; posterior angles widely obtuse, rounded. Scutellum 1.1 
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wider than long, with 12 shallow, minute, round punctures. Elytron 3.0× longer than wide, with shallow, rugose, round punctures along the striae and weakly rugo-punctate interstriae, with scattered minute setae on disc, the only regular line of erect setae occurs along external border. Pterosternum with long, abundant light yellow setae. Protibia shorter than protarsus (1.0:1.5). Metatibia shorter than metatarsus (1.0:1.2). Tarsal claws regularly bi-pectinate (
<figureCitation id="133C2A7EFF9ACA56FED0A6FEFE00A4FD" box="[369,427,1995,2015]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="2" pageNumber="1037">Fig. 5</figureCitation>
).
</paragraph>
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Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, 
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91000, 
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Escuela Superior de Agricultura del Valle del Fuerte,Universidad Autónoma de 
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,Calle 16 y Av.Japaraqui,El Estero,Juan José Rios,Ahome, 
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81110, 
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Centro de Agroecología, Instituto de Ciencias, Benemérita Universidad Autónoma de 
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, 14 Sur 6301, San Manuel, 
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, 
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72570, 
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*Corresponding author; E-mail: 
<docAuthorEmail id="D40857ECFF99CA55FED5A658FDFFA45D" box="[372,596,1901,1919]" pageId="1" pageNumber="1036">miguel.moron@inecol.mx</docAuthorEmail>
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<emphasis id="B973EAE9FF9ACA56FF3DA4CCFF51A529" bold="true" box="[156,250,1529,1547]" pageId="2" pageNumber="1037">Figs. 1–16.</emphasis>
Habitus and morphological characters of 
<taxonomicName id="4C074D78FF9ACA56FDC7A4CCFCE2A529" authorityName="Morón, Lugo-García &amp; Aragón-García" authorityYear="2015" box="[614,841,1529,1547]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="balacachiana" status="sp. nov.">
<emphasis id="B973EAE9FF9ACA56FDC7A4CCFCE2A529" box="[614,841,1529,1547]" italics="true" pageId="2" pageNumber="1037">Phyllophaga balacachiana</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FCF5A4CCFC6BA529" bold="true" box="[852,960,1529,1547]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FCF5A4CCFC6BA529" box="[852,960,1529,1547]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C074D78FF9ACA56FC4AA4CCFB10A529" authorityName="Moron, Lugo-Garcia &amp; Aragon-Garcia" authorityYear="2015" box="[1003,1211,1529,1547]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="tascatensis" status="sp. nov.">
<emphasis id="B973EAE9FF9ACA56FC4AA4CCFB10A529" box="[1003,1211,1529,1547]" italics="true" pageId="2" pageNumber="1037">Phyllophaga tascatensis</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FB67A4CCFA9AA529" bold="true" box="[1222,1329,1529,1547]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FB67A4CCFA9AA529" box="[1222,1329,1529,1547]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
. 
<emphasis id="B973EAE9FF9ACA56FA9CA4CCFAC8A529" bold="true" box="[1341,1379,1529,1547]" pageId="2" pageNumber="1037">1–4.</emphasis>
Habitus of males: 1–2. 
<taxonomicName id="4C074D78FF9ACA56FF1DA724FEE8A501" authorityName="Morón, Lugo-García &amp; Aragón-García" authorityYear="2015" box="[188,323,1553,1571]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="balacachiana" status="new species">
<emphasis id="B973EAE9FF9ACA56FF1DA724FEE8A501" box="[188,323,1553,1571]" italics="true" pageId="2" pageNumber="1037">P. balacachiana</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FEEFA724FE11A501" bold="true" box="[334,442,1553,1571]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FEEFA724FE11A501" box="[334,442,1553,1571]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
, 1. dorsal view, 2. lateral view; 3–4. 
<taxonomicName id="4C074D78FF9ACA56FD5AA724FCC5A501" authorityName="Moron, Lugo-Garcia &amp; Aragon-Garcia" authorityYear="2015" box="[763,878,1553,1571]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="tascatensis" status="new species">
<emphasis id="B973EAE9FF9ACA56FD5AA724FCC5A501" box="[763,878,1553,1571]" italics="true" pageId="2" pageNumber="1037">P. tascatensis</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FCDBA724FC4DA501" bold="true" box="[890,998,1553,1571]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FCDBA724FC4DA501" box="[890,998,1553,1571]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 
<emphasis id="B973EAE9FF9ACA56FA3EA724FA7BA501" bold="true" box="[1439,1488,1553,1571]" pageId="2" pageNumber="1037">5–10.</emphasis>
<taxonomicName id="4C074D78FF9ACA56FA74A724FEE4A519" authorityName="Morón, Lugo-García &amp; Aragón-García" authorityYear="2015" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="balacachiana" status="sp. nov.">
<emphasis id="B973EAE9FF9ACA56FA74A724FEE4A519" italics="true" pageId="2" pageNumber="1037">Phyllophaga balacachiana</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FEFAA71CFE6DA519" bold="true" box="[347,454,1577,1595]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FEFAA71CFE6DA519" box="[347,454,1577,1595]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 
<emphasis id="B973EAE9FF9ACA56FCF1A774FC27A571" bold="true" box="[848,908,1601,1619]" pageId="2" pageNumber="1037">11–16.</emphasis>
<taxonomicName id="4C074D78FF9ACA56FC31A774FBF4A571" authorityName="Moron, Lugo-Garcia &amp; Aragon-Garcia" authorityYear="2015" box="[912,1119,1601,1619]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1037" phylum="Arthropoda" rank="species" species="tascatensis" status="sp. nov.">
<emphasis id="B973EAE9FF9ACA56FC31A774FBF4A571" box="[912,1119,1601,1619]" italics="true" pageId="2" pageNumber="1037">Phyllophaga tascatensis</emphasis>
</taxonomicName>
, 
<emphasis id="B973EAE9FF9ACA56FBCBA774FB7EA571" bold="true" box="[1130,1237,1601,1619]" pageId="2" pageNumber="1037">
<taxonomicNameLabel id="A2405792FF9ACA56FBCBA774FB7EA571" box="[1130,1237,1601,1619]" pageId="2" pageNumber="1037" rank="species">new species</taxonomicNameLabel>
</emphasis>
: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BB836FBFF9ACA56FCDDA790FA69A4FD" blockId="2.[856,1538,1701,2015]" pageId="2" pageNumber="1037">
Propygidium completely dull,regularly punctate,with scattered short setae. Pygidium strongly convex (
<figureCitation id="133C2A7EFF9ACA56FB32A7F6FB61A5F5" box="[1171,1226,1731,1751]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="2" pageNumber="1037">Fig. 2</figureCitation>
), prominent, glossy, with sparse, round,shallow setiferous punctures mainly at sides; apical margin with 9 medium-size setae. Fifh abdominal sternite slightly convex, weakly furrowed on midline; anal plate short, weakly furrowed on midline, with 20 short setae, posterior margin entire at middle. Parameres large, wide, fused dorsally (
<figureCitation id="133C2A7EFF9ACA56FC4FA660FBE0A44B" box="[1006,1099,1877,1897]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="2" pageNumber="1037">Figs. 7–8</figureCitation>
), clearly dentate before apex, lateral border weakly carinate; tectum large, uniformly convex, with 2 basal rounded prominences; aedeagus much longer, sclerotized support curved downward and widely bifurcate apically (
<figureCitation id="133C2A7EFF9ACA56FB08A698FAA8A4E3" box="[1193,1283,1965,1985]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="2" pageNumber="1037">Figs. 6–8</figureCitation>
). Length of genital capsule from the apex of parameres to the border of basal piece 
<quantity id="4CFF9B1EFF9ACA56FAD3A6FEFA15A4FD" box="[1394,1470,1995,2015]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="1037" unit="mm" value="5.8">5.8 mm</quantity>
.
</paragraph>
<paragraph id="8BB836FBFF9BCA57FFC1A1F6FE82A3FB" blockId="3.[96,297,195,217]" box="[96,297,195,217]" pageId="3" pageNumber="1038">
<heading id="D0F08197FF9BCA57FFC1A1F6FE82A3FB" box="[96,297,195,217]" fontSize="10" level="3" pageId="3" pageNumber="1038" reason="9">
FEMALE 
<typeStatus id="54BC8859FF9BCA57FF1FA1F6FE82A3FB" box="[190,297,195,217]" pageId="3" pageNumber="1038" type="allotype">ALLOTYPE</typeStatus>
</heading>
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF29A1DBFD15A140" blockId="3.[96,778,238,610]" pageId="3" pageNumber="1038">
Similar to male,except:Body length 
<quantity id="4CFF9B1EFF9BCA57FE72A1DBFD83A220" box="[467,552,238,258]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="1038" unit="mm" value="15.2">15.2 mm</quantity>
.Humeral width 
<quantity id="4CFF9B1EFF9BCA57FD1DA1DBFCAEA220" box="[700,773,238,258]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="1038" unit="mm" value="5.9">5.9 mm</quantity>
. Antennal club nearly as long as preceding 5 segments combined (1.1:1.0). Anterior border of clypeus nearly straight at middle. Punctation on clypeus, frons, pronotum, and elytra denser and deeper. Pygidium flattened on basal half, shallowly depressed on distal half, with scattered setiferous small punctures; marginal bead widened near apex, with 14 short setae. Fifh sternite widely convex with few lateral short setae. Anal plate convex, with numerous short and medium-size setae. Tarsal claws irregularly serrate-dentate along external ventral borders, and irregularly pectinate along internal ventral borders (
<figureCitation id="133C2A7EFF9BCA57FE27A0C3FE61A128" box="[390,458,502,522]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="3" pageNumber="1038">Fig. 10</figureCitation>
). Ventral genital plates irregularly concave, with apical border membranous, widely sinuate. Dorsal genital plates not fused, irregularly excavate, with rounded and irregularly reflexed apical border, with few short setae near the border (
<figureCitation id="133C2A7EFF9BCA57FDDEA37BFD1FA140" box="[639,692,590,610]" captionStart="Figs" captionStartId="2.[156,193,1529,1547]" captionTargetBox="[144,1536,192,1476]" captionTargetId="figure-226@2.[144,1536,192,1498]" captionTargetPageId="2" captionText="Figs. 1–16. Habitus and morphological characters of Phyllophaga balacachiana, new species and Phyllophaga tascatensis, new species. 1–4. Habitus of males: 1–2. P. balacachiana, new species, 1. dorsal view, 2. lateral view; 3–4. P. tascatensis, new species, 3. lateral view, 4. dorsal view; scale bar = 2 mm. 5–10. Phyllophaga balacachiana, new species: 5. male tarsal claw; 6–8. male genital capsule: 6. lateral view, 7. distal view, 8. dorsal view; 9. female genital plates, ventral view; 10. female tarsal claw; scale bar = 1 mm, except in Figs. 5 and 10 = 0.5 mm. 11–16. Phyllophaga tascatensis, new species: 11. male tarsal claw; 12–14. male genital capsule: 12. lateral view, 13. distal view, 14. dorsal view; 15. female tarsal claw; 16. female genital plates, ventral view; scale bar = 1 mm, except in Figs. 11 and 15 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11548349" httpUri="https://zenodo.org/record/11548349/files/figure.png" pageId="3" pageNumber="1038">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BB836FBFF9BCA57FFC1A34EFF7DA1B3" blockId="3.[96,776,635,923]" box="[96,214,635,657]" pageId="3" pageNumber="1038">VARIATION</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A3A8FD39A004" blockId="3.[96,776,635,923]" pageId="3" pageNumber="1038">
Males. Total body length 16.2–17.0 mm. Humeral width 
<quantity id="4CFF9B1EFF9BCA57FD1EA3A8FF21A1EC" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.4" metricValueMin="5.6" pageId="3" pageNumber="1038" unit="mm" value="6.0" valueMax="6.4" valueMin="5.6">5.6–6.4 mm</quantity>
. Convexity grade of pygidial plate varies slightly between specimens, as well as number of setae. The pygidium in the specimen from Totoliboqui, 
<collectingRegion id="49C3F819FF9BCA57FF78A3C0FE8AA02B" box="[217,289,757,777]" country="Mexico" name="Sonora" pageId="3" pageNumber="1038">Sonora</collectingRegion>
, is more convex, and the setae on pygidial disc are longer; this specimen also has minute setae on the elytra.
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A21AFF4CA0B9" blockId="3.[96,776,635,923]" pageId="3" pageNumber="1038">
Females. Total body length 16.5–17.0 mm. Humeral width 
<quantity id="4CFF9B1EFF9BCA57FD1EA21AFF21A043" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="6.9" metricValueMin="6.6" pageId="3" pageNumber="1038" unit="mm" value="6.75" valueMax="6.9" valueMin="6.6">6.6–6.9 mm</quantity>
. 
<specimenCount id="9D01FD72FF9BCA57FF35A278FE8AA043" box="[148,289,845,865]" count="1" pageId="3" pageNumber="1038" type="generic">One specimen</specimenCount>
from Las Praderas has a feeble longitudinal keel near the basal border of pygidium and medium-size setae on disc of the pygidium.
</paragraph>
</subSubSection>
<subSubSection id="C31D6570FF9BCA57FFC1A281FCA2A7F6" pageId="3" pageNumber="1038" type="materials_examined">
<paragraph id="8BB836FBFF9BCA57FFC1A281FEA2A0E8" blockId="3.[96,265,948,970]" box="[96,265,948,970]" pageId="3" pageNumber="1038">
<heading id="D0F08197FF9BCA57FFC1A281FEA2A0E8" box="[96,265,948,970]" fontSize="10" level="3" pageId="3" pageNumber="1038" reason="9">
<collectionCode id="ED16AE3EFF9BCA57FFC1A281FF3EA0E8" box="[96,149,948,970]" pageId="3" pageNumber="1038">TYPE</collectionCode>
<collectionCode id="ED16AE3EFF9BCA57FF3AA281FEA2A0E8" box="[155,265,948,970]" pageId="3" pageNumber="1038">MATERIAL</collectionCode>
</heading>
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A2EAFD41A7A4" blockId="3.[96,778,991,1158]" pageId="3" pageNumber="1038">
<materialsCitation id="3B6F3CA6FF9BCA57FF25A2EAFE6EA733" collectingDate="2010-08-09" collectionCode="IEXA" collectorName="G. A. Lugo" country="Mexico" elevation="0" latitude="26.125805" location="El Fuerte" longLatPrecision="1" longitude="-108.88172" pageId="3" pageNumber="1038" specimenCount="1" stateProvince="Sinaloa" typeStatus="holotype">
<typeStatus id="54BC8859FF9BCA57FF25A2EAFF45A0D1" box="[132,238,991,1011]" pageId="3" pageNumber="1038" type="holotype">HOLOTYPE</typeStatus>
: 
<specimenCount id="9D01FD72FF9BCA57FF56A2EAFE8FA0D1" box="[247,292,991,1011]" pageId="3" pageNumber="1038" type="male">male</specimenCount>
— 
<collectingCountry id="F310766BFF9BCA57FE99A2EAFE26A0D1" box="[312,397,991,1011]" name="Mexico" pageId="3" pageNumber="1038">MEXICO</collectingCountry>
: 
<collectingRegion id="49C3F819FF9BCA57FE36A2EAFE74A0D1" box="[407,479,991,1011]" country="Mexico" name="Sinaloa" pageId="3" pageNumber="1038">Sinaloa</collectingRegion>
, 
<location id="8ED86020FF9BCA57FE49A2EAFDE9A0D1" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FE49A2EAFDE9A0D1" box="[488,578,991,1011]" country="Mexico" municipality="El Fuerte" name="El Fuerte" pageId="3" pageNumber="1038" stateProvince="Sinaloa">El Fuerte</location>
, 
<location id="8ED86020FF9BCA57FDEAA2EAFCADA0D1" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FDEAA2EAFCADA0D1" box="[587,774,991,1011]" country="Mexico" municipality="El Fuerte" name="Sierra de Balacachi" pageId="3" pageNumber="1038" stateProvince="Sinaloa">Sierra de Balacachi</location>
, 
<date id="FFB9103BFF9BCA57FFC1A2C8FF73A733" box="[96,216,1021,1041]" pageId="3" pageNumber="1038" value="2010-08-09">
<collectingDate id="EFFDE9D3FF9BCA57FFC1A2C8FF73A733" box="[96,216,1021,1041]" pageId="3" pageNumber="1038" value="2010-08-09">09-VIII-2010</collectingDate>
</date>
, 
<elevation id="002AD1C8FF9BCA57FF7EA2C8FEB6A733" box="[223,285,1021,1041]" pageId="3" pageNumber="1038">
<quantity id="4CFF9B1EFF9BCA57FF7EA2C8FEB6A733" box="[223,285,1021,1041]" metricUnit="m" pageId="3" pageNumber="1038" value="104.0">104 m</quantity>
</elevation>
, 
<collectorName id="26F2532DFF9BCA57FE84A2C8FE2EA733" box="[293,389,1021,1041]" pageId="3" pageNumber="1038">G. A. Lugo</collectorName>
(
<collectionCode id="ED16AE3EFF9BCA57FE2FA2C8FE6BA733" box="[398,448,1021,1041]" pageId="3" pageNumber="1038">IEXA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B6F3CA6FF9BCA57FE6CA2C8FF5CA70C" collectingDate="2010-08-09" collectionCode="IEXA" collectorName="G. A. Lugo" country="Mexico" elevation="0" latitude="26.125805" location="Sierra de Balacachi" longLatPrecision="1" longitude="-108.88172" municipality="El Fuerte" pageId="3" pageNumber="1038" specimenCount="1" stateProvince="Sinaloa" typeStatus="allotype">
<typeStatus id="54BC8859FF9BCA57FE6CA2C8FD9BA733" box="[461,560,1021,1041]" pageId="3" pageNumber="1038" type="allotype">ALLOTYPE</typeStatus>
: 
<specimenCount id="9D01FD72FF9BCA57FD97A2C8FDDCA733" box="[566,631,1021,1041]" pageId="3" pageNumber="1038" type="female">female</specimenCount>
—same data as holotype (
<collectionCode id="ED16AE3EFF9BCA57FF61A52FFF59A70C" box="[192,242,1050,1070]" pageId="3" pageNumber="1038">IEXA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B6F3CA6FF9BCA57FF5EA52FFEBFA769" collectingDate="2010-08-09" collectionCode="IEXA, MXAL" collectorName="G. A. Lugo" country="Mexico" elevation="0" latitude="26.125805" location="Sierra de Balacachi" longLatPrecision="1" longitude="-108.88172" municipality="El Fuerte" pageId="3" pageNumber="1038" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="Sinaloa" typeStatus="paratype">
<typeStatus id="54BC8859FF9BCA57FF5EA52FFEC6A70C" box="[255,365,1050,1070]" pageId="3" pageNumber="1038" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="9D01FD72FF9BCA57FED5A52FFE15A70C" box="[372,446,1050,1070]" count="6" pageId="3" pageNumber="1038" type="male">6 males</specimenCount>
and 
<specimenCount id="9D01FD72FF9BCA57FE4DA52FFD94A70C" box="[492,575,1050,1070]" count="1" pageId="3" pageNumber="1038" type="female">1 female</specimenCount>
—same data as holotype (
<collectionCode id="ED16AE3EFF9BCA57FF36A502FF63A769" box="[151,200,1079,1099]" pageId="3" pageNumber="1038">IEXA</collectionCode>
, 
<collectionCode id="ED16AE3EFF9BCA57FF70A502FEA4A769" box="[209,271,1079,1099]" pageId="3" pageNumber="1038">MXAL</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B6F3CA6FF9BCA57FEBCA502FD2DA74B" collectingDate="2010-07-29" collectionCode="MasAgro" collectorName="G. A. Lugo" country="Mexico" elevation="0" location="El Fuerte" municipality="El Fuerte" pageId="3" pageNumber="1038" specimenCount="1" specimenCount-female="1" stateProvince="Sinaloa" typeStatus="paratype">
<typeStatus id="54BC8859FF9BCA57FEBCA502FE20A769" box="[285,395,1079,1099]" pageId="3" pageNumber="1038" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="9D01FD72FF9BCA57FE34A502FE43A769" box="[405,488,1079,1099]" count="1" pageId="3" pageNumber="1038" type="female">1 female</specimenCount>
— 
<collectingCountry id="F310766BFF9BCA57FE5DA502FDFBA769" box="[508,592,1079,1099]" name="Mexico" pageId="3" pageNumber="1038">MEXICO</collectingCountry>
: 
<collectingRegion id="49C3F819FF9BCA57FDFBA502FD09A769" box="[602,674,1079,1099]" country="Mexico" name="Sinaloa" pageId="3" pageNumber="1038">Sinaloa</collectingRegion>
, 
<location id="8ED86020FF9BCA57FD0DA502FCADA769" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FD0DA502FCADA769" box="[684,774,1079,1099]" country="Mexico" municipality="El Fuerte" name="El Fuerte" pageId="3" pageNumber="1038" stateProvince="Sinaloa">El Fuerte</location>
, 
<location id="8ED86020FF9BCA57FFC1A560FF76A74B" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FFC1A560FF76A74B" box="[96,221,1109,1129]" country="Mexico" municipality="El Fuerte" name="Las Praderas" pageId="3" pageNumber="1038" stateProvince="Sinaloa">Las Praderas</location>
, 
<date id="FFB9103BFF9BCA57FF46A560FEF1A74B" box="[231,346,1109,1129]" pageId="3" pageNumber="1038" value="2010-07-29">
<collectingDate id="EFFDE9D3FF9BCA57FF46A560FEF1A74B" box="[231,346,1109,1129]" pageId="3" pageNumber="1038" value="2010-07-29">29-VII-2010</collectingDate>
</date>
, 
<elevation id="002AD1C8FF9BCA57FEC5A560FE0DA74B" box="[356,422,1109,1129]" pageId="3" pageNumber="1038">
<quantity id="4CFF9B1EFF9BCA57FEC5A560FE0DA74B" box="[356,422,1109,1129]" metricUnit="m" pageId="3" pageNumber="1038" value="160.0">160 m</quantity>
</elevation>
, 
<collectorName id="26F2532DFF9BCA57FE10A560FDBCA74B" box="[433,535,1109,1129]" pageId="3" pageNumber="1038">G. A. Lugo</collectorName>
(
<collectionCode id="ED16AE3EFF9BCA57FD85A560FD2AA74B" box="[548,641,1109,1129]" pageId="3" pageNumber="1038">MasAgro</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6F3CA6FF9BCA57FD30A560FD4DA7A4" collectingDate="2010-07-30" collectionCode="UAS" collectorName="G. A. Lugo" country="Mexico" elevation="78" location="Totoliboqui" municipality="Huatabampo" pageId="3" pageNumber="1038" specimenCount="1" specimenCount-male="1" stateProvince="Sonora" typeStatus="paratype">
<specimenCount id="9D01FD72FF9BCA57FD30A560FD78A74B" box="[657,723,1109,1129]" count="1" pageId="3" pageNumber="1038" type="male">1 male</specimenCount>
— 
<collectingRegion id="49C3F819FF9BCA57FD46A560FF24A7A4" country="Mexico" name="Sonora" pageId="3" pageNumber="1038">Sonora</collectingRegion>
, 
<collectingMunicipality id="6BDCAC81FF9BCA57FF36A547FEBDA7A4" box="[151,278,1138,1158]" pageId="3" pageNumber="1038">Huatabampo</collectingMunicipality>
, 
<location id="8ED86020FF9BCA57FEBCA547FE20A7A4" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FEBCA547FE20A7A4" box="[285,395,1138,1158]" country="Mexico" municipality="Huatabampo" name="Totoliboqui" pageId="3" pageNumber="1038" stateProvince="Sonora">Totoliboqui</location>
, 
<date id="FFB9103BFF9BCA57FE32A547FDAEA7A4" box="[403,517,1138,1158]" pageId="3" pageNumber="1038" value="2010-07-30">
<collectingDate id="EFFDE9D3FF9BCA57FE32A547FDAEA7A4" box="[403,517,1138,1158]" pageId="3" pageNumber="1038" value="2010-07-30">30-VII-2010</collectingDate>
</date>
, 
<quantity id="4CFF9B1EFF9BCA57FDACA547FDEBA7A4" box="[525,576,1138,1158]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="1038" unit="m" value="78.0">
<elevation id="002AD1C8FF9BCA57FDACA547FDEBA7A4" box="[525,576,1138,1158]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="1038" unit="m" value="78.0">78 m</elevation>
</quantity>
, 
<collectorName id="26F2532DFF9BCA57FDE9A547FD03A7A4" box="[584,680,1138,1158]" pageId="3" pageNumber="1038">G. A. Lugo</collectorName>
(
<collectionCode id="ED16AE3EFF9BCA57FD13A547FD4BA7A4" box="[690,736,1138,1158]" country="Mexico" httpUri="http://biocol.org/cool/swmz-m648" name="Unidad Académica Sisal, Universidad Nacional Autónoma de México" pageId="3" pageNumber="1038" type="University or college">UAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<materialsCitation id="3B6F3CA6FF9BCA57FFC1A5AAFCAEA7F6" collectionCode="TYPE, LOCALITY" collectorName="de Balacachi" country="Mexico" latitude="26.125805" location="Sierra de Balacachi" longLatPrecision="1" longitude="-108.88172" pageId="3" pageNumber="1038" specimenCount="1" stateProvince="Sinaloa">
<paragraph id="8BB836FBFF9BCA57FFC1A5AAFEAAA797" blockId="3.[96,777,1183,1237]" box="[96,257,1183,1205]" pageId="3" pageNumber="1038">TYPE LOCALITY</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A5F5FCA2A7F6" blockId="3.[96,777,1183,1237]" box="[132,777,1215,1237]" pageId="3" pageNumber="1038">
<location id="8ED86020FF9BCA57FF25A5F5FE96A7F6" LSID="urn:lsid:plazi:treatment:03AE87EDFF99CA57FC89A5EBFD84A5D2:8ED86020FF9BCA57FF25A5F5FE96A7F6" box="[132,317,1216,1236]" country="Mexico" latitude="26.125805" longLatPrecision="1" longitude="-108.88172" name="Sierra de Balacachi" pageId="3" pageNumber="1038" stateProvince="Sinaloa">Sierra de Balacachi</location>
, 
<collectingRegion id="49C3F819FF9BCA57FEE5A5F5FE25A7F6" box="[324,398,1216,1236]" country="Mexico" name="Sinaloa" pageId="3" pageNumber="1038">Sinaloa</collectingRegion>
, 
<collectingCountry id="F310766BFF9BCA57FE34A5F5FE77A7F6" box="[405,476,1216,1236]" name="Mexico" pageId="3" pageNumber="1038">México</collectingCountry>
(
<geoCoordinate id="EE33503CFF9BCA57FE47A5F5FDF4A7F6" box="[486,607,1215,1237]" degrees="26" direction="north" minutes="7" orientation="latitude" pageId="3" pageNumber="1038" precision="1" seconds="32.9" value="26.125805">
26°7'32.9 
<emphasis id="B973EAE9FF9BCA57FDE2A58AFDE6A7F7" bold="true" box="[579,589,1215,1237]" pageId="3" pageNumber="1038">&quot;</emphasis>
N
</geoCoordinate>
, 
<geoCoordinate id="EE33503CFF9BCA57FDC6A5F5FD54A7F6" box="[615,767,1215,1237]" degrees="108" direction="west" minutes="52" orientation="longitude" pageId="3" pageNumber="1038" precision="1" seconds="54.2" value="-108.88172">
108°52'54.2 
<emphasis id="B973EAE9FF9BCA57FD7DA58AFD4DA7F7" bold="true" box="[732,742,1215,1237]" pageId="3" pageNumber="1038">&quot;</emphasis>
W
</geoCoordinate>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C31D6570FF9BCA57FFC1A5D8FEB8A680" pageId="3" pageNumber="1038" type="biology_ecology">
<paragraph id="8BB836FBFF9BCA57FFC1A5D8FE89A621" blockId="3.[96,290,1261,1283]" box="[96,290,1261,1283]" pageId="3" pageNumber="1038">
<heading id="D0F08197FF9BCA57FFC1A5D8FE89A621" box="[96,290,1261,1283]" fontSize="10" level="3" pageId="3" pageNumber="1038" reason="9">
<collectionCode id="ED16AE3EFF9BCA57FFC1A5D8FF4FA621" box="[96,228,1261,1283]" pageId="3" pageNumber="1038">BIOLOGICAL</collectionCode>
<collectionCode id="ED16AE3EFF9BCA57FF4BA5D8FE89A621" box="[234,290,1261,1283]" pageId="3" pageNumber="1038">DATA</collectionCode>
</heading>
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A42DFEB8A680" blockId="3.[96,780,1304,1442]" pageId="3" pageNumber="1038">
Specimens of 
<taxonomicName id="4C074D78FF9BCA57FEAAA42DFE0AA60E" authorityName="Morón, Lugo-García &amp; Aragón-García" authorityYear="2015" box="[267,417,1304,1324]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="balacachiana">
<emphasis id="B973EAE9FF9BCA57FEAAA42DFE0AA60E" box="[267,417,1304,1324]" italics="true" pageId="3" pageNumber="1038">P. balacachiana</emphasis>
</taxonomicName>
were collected in tropical thorny forest, at 
<quantity id="4CFF9B1EFF9BCA57FF00A403FEAAA668" box="[161,257,1334,1354]" metricMagnitude="2" metricUnit="m" metricValue="1.19" metricValueMax="1.6" metricValueMin="0.78" pageId="3" pageNumber="1038" unit="m" value="119.0" valueMax="160.0" valueMin="78.0">78–160 m</quantity>
elevation in Jul (
<specimenCount id="9D01FD72FF9BCA57FE00A403FDB6A668" box="[417,541,1334,1354]" count="2" pageId="3" pageNumber="1038" type="generic">2 specimens</specimenCount>
) and Aug (
<specimenCount id="9D01FD72FF9BCA57FD23A403FCABA668" box="[642,768,1334,1354]" count="9" pageId="3" pageNumber="1038" type="generic">9 specimens</specimenCount>
). Other species of 
<taxonomicName id="4C074D78FF9BCA57FEA3A466FED1A645" authorityName="Harris" authorityYear="1826" box="[258,378,1363,1383]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="genus">
<emphasis id="B973EAE9FF9BCA57FEA3A466FED1A645" box="[258,378,1363,1383]" italics="true" pageId="3" pageNumber="1038">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were 
<taxonomicName id="4C074D78FF9BCA57FD59A466FE30A6A6" authority="(Arrow)" baseAuthorityName="Arrow" baseAuthorityYear="1933" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="cristagalli" subGenus="Listrochelus">
<emphasis id="B973EAE9FF9BCA57FD59A466FCA7A645" box="[760,780,1363,1383]" italics="true" pageId="3" pageNumber="1038">P.</emphasis>
(
<emphasis id="B973EAE9FF9BCA57FFC6A445FF72A6A6" box="[103,217,1392,1412]" italics="true" pageId="3" pageNumber="1038">Listrochelus</emphasis>
) 
<emphasis id="B973EAE9FF9BCA57FF49A445FEEDA6A6" box="[232,326,1392,1412]" italics="true" pageId="3" pageNumber="1038">cristagalli</emphasis>
(Arrow)
</taxonomicName>
, 
<taxonomicName id="4C074D78FF9BCA57FE05A445FDF5A6A6" authority="Saylor" authorityName="Saylor" authorityYear="1940" box="[420,606,1392,1412]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="granti" subGenus="Listrochelus">
<emphasis id="B973EAE9FF9BCA57FE05A445FE1CA6A6" box="[420,439,1392,1412]" italics="true" pageId="3" pageNumber="1038">P.</emphasis>
(
<emphasis id="B973EAE9FF9BCA57FE63A445FE79A6A6" box="[450,466,1392,1412]" italics="true" pageId="3" pageNumber="1038">L.</emphasis>
) 
<emphasis id="B973EAE9FF9BCA57FE41A445FDB1A6A6" box="[480,538,1392,1412]" italics="true" pageId="3" pageNumber="1038">granti</emphasis>
Saylor
</taxonomicName>
, and another undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="C31D6570FF9BCA57FFC1A4F2FE8DA559" pageId="3" pageNumber="1038" type="discussion">
<paragraph id="8BB836FBFF9BCA57FFC1A4F2FEF3A6FF" blockId="3.[96,344,1479,1501]" box="[96,344,1479,1501]" pageId="3" pageNumber="1038">
<heading id="D0F08197FF9BCA57FFC1A4F2FEF3A6FF" box="[96,344,1479,1501]" fontSize="10" level="3" pageId="3" pageNumber="1038" reason="9">TAXONOMIC REMARKS</heading>
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A4C7FE8DA559" blockId="3.[96,776,1522,1659]" pageId="3" pageNumber="1038">
<taxonomicName id="4C074D78FF9BCA57FF25A4C7FE2FA524" authorityName="Morón, Lugo-García &amp; Aragón-García" authorityYear="2015" box="[132,388,1522,1542]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="balacachiana">
<emphasis id="B973EAE9FF9BCA57FF25A4C7FE2FA524" box="[132,388,1522,1542]" italics="true" pageId="3" pageNumber="1038">Phyllophaga balacachiana</emphasis>
</taxonomicName>
is very similar to 
<taxonomicName id="4C074D78FF9BCA57FD93A4C7FD6DA524" authority="Saylor" authorityName="Saylor" authorityYear="1940" box="[562,710,1522,1542]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="granti">
<emphasis id="B973EAE9FF9BCA57FD93A4C7FD29A524" box="[562,642,1522,1542]" italics="true" pageId="3" pageNumber="1038">P. granti</emphasis>
Saylor
</taxonomicName>
, but in the latter the parameres have an acute projection dorsally, and the surface of elytra in both sexes is clothed with scattered minute setae. See modification of the 
<bibRefCitation id="EF964B0AFF9BCA57FE85A77FFE0DA57C" author="Saylor LW" box="[292,422,1610,1630]" pageId="3" pageNumber="1038" pagination="59 - 130" refId="ref7725" refString="Saylor LW. 1940. Revision of the scarabaeid beetles of the Phyllophaga subgenus Listrochelus of the U. S. with discussion of related subgenera. Proceed- ings of the United States National Museum 89: 59 - 130." type="journal article" year="1940">Saylor (1940)</bibRefCitation>
key at the end of the 
<taxonomicName id="4C074D78FF9BCA57FD27A77FFCA3A57C" authorityName="Moron, Lugo-Garcia &amp; Aragon-Garcia" authorityYear="2015" box="[646,776,1610,1630]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1038" phylum="Arthropoda" rank="species" species="tascatensis" status="sp. nov.">
<emphasis id="B973EAE9FF9BCA57FD27A77FFCA3A57C" box="[646,776,1610,1630]" italics="true" pageId="3" pageNumber="1038">P. tascatensis</emphasis>
</taxonomicName>
<emphasis id="B973EAE9FF9BCA57FFC1A752FF06A559" bold="true" box="[96,173,1639,1659]" pageId="3" pageNumber="1038">
<taxonomicNameLabel id="A2405792FF9BCA57FFC1A752FF06A559" box="[96,173,1639,1659]" pageId="3" pageNumber="1038" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
description.
</paragraph>
</subSubSection>
<subSubSection id="C31D6570FF9BCA57FFC1A7A1FD84A5D2" pageId="3" pageNumber="1038" type="etymology">
<paragraph id="8BB836FBFF9BCA57FFC1A7A1FF42A588" blockId="3.[96,233,1684,1706]" box="[96,233,1684,1706]" pageId="3" pageNumber="1038">
<heading id="D0F08197FF9BCA57FFC1A7A1FF42A588" box="[96,233,1684,1706]" fontSize="10" level="3" pageId="3" pageNumber="1038" reason="9">ETYMOLOGY</heading>
</paragraph>
<paragraph id="8BB836FBFF9BCA57FF25A78AFD84A5D2" blockId="3.[96,776,1727,1776]" pageId="3" pageNumber="1038">
The species epithet was derived from the indigenous language name “yoreme” of the 
<typeStatus id="54BC8859FF9BCA57FEE1A7E9FEC0A5D2" box="[320,363,1756,1776]" pageId="3" pageNumber="1038">type</typeStatus>
locality “Balacachi.”
</paragraph>
</subSubSection>
</treatment>
</document>