654 lines
65 KiB
XML
654 lines
65 KiB
XML
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<document ID-DOI="10.1590/S1679-62252011000300006" ID-ISSN="1982-0224" ID-Zenodo-Dep="5422996" approvalRequired="6" approvalRequired_for_textStreams="6" checkinTime="1630382949743" checkinUser="carolina" docAuthor="Birindelli, José L. O., Fayal, Danielle F. & Wosiacki, Wolmar B." docDate="2011" docId="03FE1801FFC4FF82D48BF962DA89FB85" docLanguage="en" docName="NeotropIchthyol.2011.51.pdf" docOrigin="Neotropical Ichthyology 9 (3)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252011000300006&lng=en&tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Hassar gabiru Birindelli & Fayal & Wosiacki 2011, new species" docType="treatment" docVersion="2" lastPageId="10" lastPageNumber="525" masterDocId="FFC76079FFC1FF88D758FFB7D95CFFC1" masterDocTitle="Taxonomic revision of thorny catfish genus Hassar (Siluriformes: Doradidae)" masterLastPageNumber="542" masterPageNumber="515" pageId="5" pageNumber="520" updateTime="1630689920882" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of thorny catfish genus Hassar (Siluriformes: Doradidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Birindelli, José L. O.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fayal, Danielle F.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wosiacki, Wolmar B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2011-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>9</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>515</mods:start>
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<mods:end>542</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252011000300006&lng=en&tlng=en</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S1679-62252011000300006</mods:identifier>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5422996</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85" httpUri="http://treatment.plazi.org/id/03FE1801FFC4FF82D48BF962DA89FB85" lastPageId="10" lastPageNumber="525" pageId="5" pageNumber="520">
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<subSubSection box="[979,1281,1749,1768]" pageId="5" pageNumber="520" type="nomenclature">
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<paragraph blockId="5.[979,1281,1749,1800]" box="[979,1281,1749,1768]" pageId="5" pageNumber="520">
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<heading box="[979,1281,1749,1768]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="520" reason="2">
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<emphasis bold="true" box="[979,1281,1749,1768]" pageId="5" pageNumber="520">
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<taxonomicName authority="Birindelli & Fayal & Wosiacki, 2011" authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[979,1137,1749,1768]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="520" phylum="Chordata" rank="species" species="gabiru" status="new species">
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<emphasis bold="true" box="[979,1137,1749,1768]" italics="true" pageId="5" pageNumber="520">Hassar gabiru</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1150,1281,1749,1768]" pageId="5" pageNumber="520" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="520" type="description">
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<paragraph blockId="5.[979,1281,1749,1800]" box="[1060,1200,1781,1800]" pageId="5" pageNumber="520">
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<emphasis bold="true" box="[1060,1200,1781,1800]" pageId="5" pageNumber="520">
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<figureCitation box="[1060,1133,1781,1800]" captionStart="Fig" captionStartId="9.[129,168,1873,1892]" captionTargetBox="[129,1452,280,1834]" captionTargetId="figure-56@9.[129,1452,280,1834]" captionTargetPageId="9" captionText="Fig. 7. Hassar gabiru: (a) MZUSP 108440, 103.4 mm SL, holotype, rio Curuá, Altamira, PA; (b) INPA 26726, 156.6 mm SL, rio Xingu at Ilha do Babaquara, Altamira, PA; (c) MZUSP 87026, 65.3 mm SL, paratype, rio Curisevo, Gaúcha do Norte, MT." figureDoi="http://doi.org/10.5281/zenodo.5423010" httpUri="https://zenodo.org/record/5423010/files/figure.png" pageId="5" pageNumber="520">Figs. 7</figureCitation>
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and
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<figureCitation box="[1187,1200,1781,1800]" captionStart="Fig" captionStartId="10.[136,175,757,776]" captionTargetBox="[156,1415,217,715]" captionTargetId="figure-515@10.[136,1459,193,715]" captionTargetPageId="10" captionText="Fig. 8. Hassar gabiru, paratype, MZUSP 97219, c.130 mm SL, rio Curuá, Iriri-Xingu drainage, Altamira, PA. Photo by Mark Sabaj Pérez." figureDoi="http://doi.org/10.5281/zenodo.5423012" httpUri="https://zenodo.org/record/5423012/files/figure.png" pageId="5" pageNumber="520">8</figureCitation>
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</emphasis>
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</paragraph>
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<paragraph blockId="5.[809,1452,1845,1960]" pageId="5" pageNumber="520">
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<taxonomicName box="[809,980,1845,1864]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="520" phylum="Chordata" rank="species" species="orestis">
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<emphasis box="[809,980,1845,1864]" italics="true" pageId="5" pageNumber="520">Hassar orestis</emphasis>
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</taxonomicName>
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.—
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<bibRefCitation author="Camargo, M" box="[1028,1344,1845,1864]" pageId="5" pageNumber="520" pagination="123 - 147" refId="ref21541" refString="Camargo, M, T. Giarrizzo & V. Isaac. 2004. Review of the geographic distribution of fish fauna of the Xingu river basin, Brazil. Ecotropica, 10: 123 - 147." type="journal article" year="2004">
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Camargo
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<emphasis box="[1143,1197,1845,1864]" italics="true" pageId="5" pageNumber="520">et al</emphasis>
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., 2004: 143
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</bibRefCitation>
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[in part, distribution, Xingu upstream, middle Xingu, Bacajá river].—Camargo, 2009: 208 [feeding habitat].—Camargo & Giarrizzo, 2009: 221 [biological data].
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5423004" ID-Zenodo-Dep="5423004" httpUri="https://zenodo.org/record/5423004/files/figure.png" pageId="6" pageNumber="521" startId="6.[136,174,1781,1800]" targetBox="[136,1459,174,1761]" targetPageId="6">
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<paragraph blockId="6.[136,1459,1781,1960]" pageId="6" pageNumber="521">
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<emphasis bold="true" box="[136,203,1781,1800]" pageId="6" pageNumber="521">Fig. 4.</emphasis>
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Gas bladders in ventral view of
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<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1881" box="[531,676,1781,1800]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="affinis">
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<emphasis box="[531,676,1781,1800]" italics="true" pageId="6" pageNumber="521">Hassar affinis</emphasis>
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</taxonomicName>
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(a-d): (
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<emphasis bold="true" box="[752,765,1781,1800]" pageId="6" pageNumber="521">a</emphasis>
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) MZUSP 90583, 54.4 mm SL; (
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<emphasis bold="true" box="[1095,1110,1781,1800]" pageId="6" pageNumber="521">b</emphasis>
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) MZUSP 74890, 85.9 mm SL; (
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<emphasis bold="true" box="[1438,1450,1781,1800]" pageId="6" pageNumber="521">c</emphasis>
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) MZUSP 43604, 228.0 mm SL; (
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<emphasis bold="true" box="[468,483,1813,1832]" pageId="6" pageNumber="521">d</emphasis>
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) ANSP 69392, 201.6 mm SL (holotype of
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<taxonomicName authorityName="Fowler" authorityYear="1941" box="[923,1069,1813,1832]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="woodi">
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<emphasis box="[923,1069,1813,1832]" italics="true" pageId="6" pageNumber="521">Hassar woodi</emphasis>
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</taxonomicName>
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, photo by Mark Sabaj Pérez);
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<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gabiru">
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<emphasis italics="true" pageId="6" pageNumber="521">Hassar gabiru</emphasis>
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</taxonomicName>
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(e-h): (
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<emphasis bold="true" box="[278,289,1845,1864]" pageId="6" pageNumber="521">e</emphasis>
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) MZUSP 87026, 69.0 mm SL; (
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<emphasis bold="true" box="[611,619,1845,1864]" pageId="6" pageNumber="521">f</emphasis>
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) MZUSP 94142, 137.6 mm SL; (
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<emphasis bold="true" box="[953,966,1845,1864]" pageId="6" pageNumber="521">g</emphasis>
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) INPA 31043, 122.5 mm SL; (
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<emphasis bold="true" box="[1271,1285,1845,1864]" pageId="6" pageNumber="521">h</emphasis>
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) MZUSP 94142, 128.6 mm SL;
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<taxonomicName authorityName="Kindle" authorityYear="1895" box="[285,437,1877,1896]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="wilderi">
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<emphasis box="[285,437,1877,1896]" italics="true" pageId="6" pageNumber="521">Hassar wilderi</emphasis>
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</taxonomicName>
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(i-l): (
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<emphasis bold="true" box="[500,507,1877,1896]" pageId="6" pageNumber="521">i</emphasis>
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) MZUSP 46016, 67.0 mm SL; (
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<emphasis bold="true" box="[831,839,1877,1896]" pageId="6" pageNumber="521">j</emphasis>
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) MZUSP 4857, 103.6 mm SL; (
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<emphasis bold="true" box="[1163,1177,1877,1896]" pageId="6" pageNumber="521">k</emphasis>
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) MZUSP 63148, 148.8 mm SL; (
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<emphasis bold="true" box="[186,193,1909,1928]" pageId="6" pageNumber="521">l</emphasis>
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) MZUSP 62998, 161.6 mm SL;
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<taxonomicName box="[522,670,1909,1928]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="orestis">
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<emphasis box="[522,670,1909,1928]" italics="true" pageId="6" pageNumber="521">Hassar orestis</emphasis>
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</taxonomicName>
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(m-p): (
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<emphasis bold="true" box="[753,775,1909,1928]" pageId="6" pageNumber="521">m</emphasis>
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) MZUSP 6991, 71.0 mm SL; (
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<emphasis bold="true" box="[1086,1100,1909,1928]" pageId="6" pageNumber="521">n</emphasis>
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) MZUSP 74680, 123.5 mm SL; (
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<emphasis bold="true" box="[1437,1450,1909,1928]" pageId="6" pageNumber="521">o</emphasis>
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) MZUSP 15512, 132.0 mm SL; (
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<emphasis bold="true" box="[458,472,1941,1960]" pageId="6" pageNumber="521">p</emphasis>
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) MZUSP 32542, 220.0 mm SL. Scale bars = 5 mm.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5423006" ID-Zenodo-Dep="5423006" httpUri="https://zenodo.org/record/5423006/files/figure.png" pageId="7" pageNumber="522" startId="7.[129,167,575,594]" targetBox="[160,759,197,518]" targetPageId="7">
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<paragraph blockId="7.[129,772,575,690]" pageId="7" pageNumber="522">
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<emphasis bold="true" box="[129,195,575,594]" pageId="7" pageNumber="522">Fig. 5.</emphasis>
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Schematic illustration of midlateral scutes of (
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<emphasis bold="true" box="[669,682,575,594]" pageId="7" pageNumber="522">a</emphasis>
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)
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<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1881" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="affinis">
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<emphasis italics="true" pageId="7" pageNumber="522">Hassar affinis</emphasis>
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</taxonomicName>
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, based on MZUSP 89935, 110.7 mm SL, and (
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<emphasis bold="true" box="[668,682,607,626]" pageId="7" pageNumber="522">b</emphasis>
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)
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<taxonomicName class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="orestis">
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<emphasis italics="true" pageId="7" pageNumber="522">Hassar orestis</emphasis>
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</taxonomicName>
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, based on MZUSP 6991, 65.7 mm SL, in left lateral view. Arrow indicates anteriormost scute with median thorn.
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</paragraph>
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</caption>
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<subSubSection pageId="7" pageNumber="522" type="materials_examined">
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<paragraph blockId="7.[129,772,782,886]" pageId="7" pageNumber="522">
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<materialsCitation collectingDate="2010-10-22" collectionCode="MZUSP" collectorName="Vila Castelo dos Sonhos & Altamira, PA & J. L. Birindelli & L. M. Sousa & A. L. Netto-Ferreira & M. H. Sabaj Perez & N. Lujan." country="Brazil" latitude="-8.318611" location="rio Curua" longLatPrecision="21" longitude="-55.08972" municipality="Iriri" pageId="7" pageNumber="522" specimenCode="MZUSP 108440" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[129,231,782,799]" pageId="7" pageNumber="522">
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<typeStatus box="[129,227,782,799]" pageId="7" pageNumber="522">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[241,406,782,799]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 108440</specimenCode>
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(
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<quantity box="[426,556,782,799]" metricMagnitude="0" metricUnit="m" metricValue="1.1304" pageId="7" pageNumber="522" unit="mm" value="1130.4">1, 130.4 mm</quantity>
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SL),
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<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD533FCB9DB93FCDE" box="[619,719,782,799]" country="Brazil" latitude="-8.318611" longLatPrecision="21" longitude="-55.08972" municipality="Iriri" name="rio Curua" pageId="7" pageNumber="522">rio Curuá</location>
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,
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<collectingMunicipality box="[732,771,782,799]" pageId="7" pageNumber="522">Iriri</collectingMunicipality>
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drainage, at
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<collectorName box="[261,522,811,828]" pageId="7" pageNumber="522">Vila Castelo dos Sonhos</collectorName>
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,
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<geoCoordinate box="[537,635,811,828]" degrees="8" direction="south" minutes="19" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="7" value="-8.318611">8°19’7’’S</geoCoordinate>
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<geoCoordinate box="[646,767,811,828]" degrees="55" direction="west" minutes="5" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="23" value="-55.08972">55°5’23’’W</geoCoordinate>
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,
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|
<collectorName box="[129,258,840,857]" pageId="7" pageNumber="522">Altamira, PA</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectingCountry box="[267,329,840,857]" name="Brazil" pageId="7" pageNumber="522">Brazil</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<date box="[338,459,840,857]" pageId="7" pageNumber="522" value="2010-10-22">
|
|||
|
|
<collectingDate box="[338,459,840,857]" pageId="7" pageNumber="522" value="2010-10-22">22 Oct 2010</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[468,611,840,857]" pageId="7" pageNumber="522">J. L. Birindelli</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[620,740,840,857]" pageId="7" pageNumber="522">L. M. Sousa</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName pageId="7" pageNumber="522">A. L. Netto-Ferreira</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[308,488,869,886]" pageId="7" pageNumber="522">M. H. Sabaj Pérez</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[498,592,869,886]" pageId="7" pageNumber="522">N. Lujan.</collectorName>
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="7.[129,772,929,1609]" pageId="7" pageNumber="522">
|
|||
|
|
<materialsCitation collectingDate="1981-08" collectingDateMax="2008-08" collectingDateMin="1981-08" collectionCode="ANSP, INPA, MPEG, MZUSP" collectorName="Altamira, PA & L. Rapp & J. Zuanon. & H. Lopez-Fernandez & M. Goulding. & C. R. Moreira & J. L. Birindelli & da Mirim & Tarik Yegun & M. C. Lopes" country="Brazil" county="rio Jatoba" latitude="-12.696944" location="Xingu" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" pageId="7" pageNumber="522" specimenCode="ANSP 187335, INPA 26726, INPA 31043, MPEG 847, MPEG 6736, MZUSP 86888, MZUSP 86991, MZUSP 87026, MZUSP 94142, MZUSP 94308, MZUSP 94374, MZUSP 97219, MZUSP 100328" specimenCount="2" typeStatus="paratype">
|
|||
|
|
<emphasis bold="true" box="[129,466,929,946]" pageId="7" pageNumber="522">
|
|||
|
|
<typeStatus box="[129,235,929,946]" pageId="7" pageNumber="522">Paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7AEFC16D869FC73" box="[246,309,929,946]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Xingu" pageId="7" pageNumber="522">Xingu</location>
|
|||
|
|
basin (
|
|||
|
|
<collectingCountry box="[385,454,929,946]" name="Brazil" pageId="7" pageNumber="522">Brazil</collectingCountry>
|
|||
|
|
):
|
|||
|
|
</emphasis>
|
|||
|
|
<specimenCode box="[473,613,929,946]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="522">ANSP 187335</specimenCode>
|
|||
|
|
(3, 125.0-
|
|||
|
|
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.517" pageId="7" pageNumber="522" unit="mm" value="151.7">151.7 mm</quantity>
|
|||
|
|
SL), same data as holotype.
|
|||
|
|
<specimenCode box="[440,559,958,975]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="7" pageNumber="522">INPA 26726</specimenCode>
|
|||
|
|
(
|
|||
|
|
<quantity box="[573,692,958,975]" metricMagnitude="0" metricUnit="m" metricValue="1.1565999999999999" pageId="7" pageNumber="522" unit="mm" value="1156.6">1, 156.6 mm</quantity>
|
|||
|
|
SL),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD5B0FC09D99CFC2D" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Xingu" pageId="7" pageNumber="522">rio Xingu</location>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD794FC6CD8D9FC2D" box="[204,389,987,1004]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Ilha do Babaquara" pageId="7" pageNumber="522">Ilha do Babaquara</location>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[399,531,987,1004]" pageId="7" pageNumber="522">Altamira, PA</collectorName>
|
|||
|
|
,
|
|||
|
|
<date box="[543,637,987,1004]" pageId="7" pageNumber="522" value="1990-10">
|
|||
|
|
<collectingDate box="[543,637,987,1004]" pageId="7" pageNumber="522" value="1990-10">Oct 1990</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[649,729,987,1004]" pageId="7" pageNumber="522">L. Rapp</collectorName>
|
|||
|
|
Py-
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7D9FC4FD99FFBC8" box="[129,195,1016,1033]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Daniel" pageId="7" pageNumber="522">Daniel</location>
|
|||
|
|
&
|
|||
|
|
<collectorName box="[233,342,1016,1033]" pageId="7" pageNumber="522">J. Zuanon.</collectorName>
|
|||
|
|
<specimenCode box="[351,476,1016,1033]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="7" pageNumber="522">INPA 31043</specimenCode>
|
|||
|
|
(
|
|||
|
|
<quantity box="[495,626,1016,1033]" metricMagnitude="0" metricUnit="m" metricValue="3.1225" pageId="7" pageNumber="522" unit="mm" value="3122.5">3, 122.5 mm</quantity>
|
|||
|
|
SL [only one measured]),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7A1FBA2D866FBE7" box="[249,314,1045,1062]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Mouth" pageId="7" pageNumber="522">Mouth</location>
|
|||
|
|
of
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD600FBA2D8ECFBE7" box="[344,432,1045,1062]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Novo" pageId="7" pageNumber="522">rio Novo</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[441,548,1045,1062]" degrees="4" direction="south" minutes="28" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="15" value="-4.4708333">4°28’15’’S</geoCoordinate>
|
|||
|
|
53°40’6’W,
|
|||
|
|
<date box="[670,767,1045,1062]" pageId="7" pageNumber="522" value="2008-08">
|
|||
|
|
<collectingDate box="[670,767,1045,1062]" pageId="7" pageNumber="522" value="2008-08">Aug 2008</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[129,332,1073,1090]" pageId="7" pageNumber="522">H. López-Fernández</collectorName>
|
|||
|
|
<emphasis box="[340,385,1073,1090]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[400,512,1073,1090]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="7" pageNumber="522">MPEG 847</specimenCode>
|
|||
|
|
(3, 142.4-
|
|||
|
|
<quantity box="[618,720,1073,1090]" metricMagnitude="-1" metricUnit="m" metricValue="1.722" pageId="7" pageNumber="522" unit="mm" value="172.2">172.2 mm</quantity>
|
|||
|
|
SL), confluence of
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD650FBF9D82BFB9E" box="[264,375,1102,1119]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Culuene" pageId="7" pageNumber="522">rio Culuene</location>
|
|||
|
|
and
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD6FBFBF9DB37FB9E" box="[419,619,1102,1119]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Sete de Setembro" pageId="7" pageNumber="522">rio Sete de Setembro</location>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[671,767,1102,1119]" pageId="7" pageNumber="522" value="1981-08">
|
|||
|
|
<collectingDate box="[671,767,1102,1119]" pageId="7" pageNumber="522" value="1981-08">Aug 1981</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[129,264,1131,1148]" pageId="7" pageNumber="522">M. Goulding.</collectorName>
|
|||
|
|
<specimenCode box="[271,396,1131,1148]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="7" pageNumber="522">MPEG 6736</specimenCode>
|
|||
|
|
(3, 108.7-
|
|||
|
|
<quantity box="[503,585,1131,1148]" metricMagnitude="-1" metricUnit="m" metricValue="1.31" pageId="7" pageNumber="522" unit="mm" value="131.0">131 mm</quantity>
|
|||
|
|
SL), Xingu basin,
|
|||
|
|
<collectorName box="[129,256,1160,1177]" pageId="7" pageNumber="522">Altamira, PA</collectorName>
|
|||
|
|
,
|
|||
|
|
<date box="[265,353,1160,1177]" pageId="7" pageNumber="522" value="2001-01">
|
|||
|
|
<collectingDate box="[265,353,1160,1177]" pageId="7" pageNumber="522" value="2001-01">Jan 2001</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, Camargo.
|
|||
|
|
<specimenCode box="[460,604,1160,1177]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 86888</specimenCode>
|
|||
|
|
(1,
|
|||
|
|
<quantity box="[640,724,1160,1177]" metricMagnitude="-2" metricUnit="m" metricValue="5.62" pageId="7" pageNumber="522" unit="mm" value="56.2">56.2 mm</quantity>
|
|||
|
|
SL),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7D9FB12D80AFB77" box="[129,342,1189,1206]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Sete de Setembro" pageId="7" pageNumber="522">rio Sete de Setembro</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[355,474,1189,1206]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="19" value="-13.505278">13°30’19’’S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[483,610,1189,1206]" degrees="52" direction="west" minutes="24" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="57" value="-52.415836">52°24’57’’W</geoCoordinate>
|
|||
|
|
, Canarana, MT,
|
|||
|
|
<date box="[129,222,1217,1234]" pageId="7" pageNumber="522" value="2004-10">
|
|||
|
|
<collectingDate box="[129,222,1217,1234]" pageId="7" pageNumber="522" value="2004-10">Oct 2004</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[232,368,1217,1234]" pageId="7" pageNumber="522">C. R. Moreira</collectorName>
|
|||
|
|
<emphasis box="[376,420,1217,1234]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[432,581,1217,1234]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 86991</specimenCode>
|
|||
|
|
(12,
|
|||
|
|
<quantity box="[633,771,1217,1234]" metricMagnitude="-2" metricUnit="m" metricValue="7.895" metricValueMax="9.14" metricValueMin="6.65" pageId="7" pageNumber="522" unit="mm" value="78.95" valueMax="91.4" valueMin="66.5">66.5-91.4 mm</quantity>
|
|||
|
|
SL),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7EAFB69D874FB2E" box="[178,296,1246,1263]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Culuene" pageId="7" pageNumber="522">rio Culuene</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[306,421,1246,1263]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="52" value="-13.514444">13°30’52”S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[427,540,1246,1263]" degrees="53" direction="west" minutes="5" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="34" value="-53.092777">53°5’34”W</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD57FFB69DB8CFB2E" box="[551,720,1246,1263]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Gaucha do Norte" pageId="7" pageNumber="522">Gaúcha do Norte</location>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[129,222,1275,1292]" pageId="7" pageNumber="522" value="2004-10">
|
|||
|
|
<collectingDate box="[129,222,1275,1292]" pageId="7" pageNumber="522" value="2004-10">Oct 2004</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[233,378,1275,1292]" pageId="7" pageNumber="522">J. L. Birindelli</collectorName>
|
|||
|
|
<emphasis box="[385,429,1275,1292]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[443,592,1275,1292]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 87026</specimenCode>
|
|||
|
|
(2,
|
|||
|
|
<quantity box="[633,772,1275,1292]" metricMagnitude="-2" metricUnit="m" metricValue="7.225" metricValueMax="7.92" metricValueMin="6.529999999999999" pageId="7" pageNumber="522" unit="mm" value="72.25" valueMax="79.2" valueMin="65.3">65.3-79.2 mm</quantity>
|
|||
|
|
SL),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7E8FAAFD875FAE8" box="[176,297,1304,1321]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Curisevo" pageId="7" pageNumber="522">rio Curisevo</location>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD669FAAFD892FAE8" box="[305,462,1304,1321]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Porto do Vitorio" pageId="7" pageNumber="522">Porto do Vitório</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[470,560,1304,1321]" degrees="13" direction="south" minutes="2" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="5" value="-13.034722">13°2’5”S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[565,688,1304,1321]" degrees="53" direction="west" minutes="25" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="19" value="-53.421947">53°25’19”W</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD5E1FAAFD987FA87" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Gaucha do Norte" pageId="7" pageNumber="522">Gaúcha do Norte</location>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[279,373,1333,1350]" pageId="7" pageNumber="522" value="2004-10">
|
|||
|
|
<collectingDate box="[279,373,1333,1350]" pageId="7" pageNumber="522" value="2004-10">Oct 2004</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, Moreira
|
|||
|
|
<emphasis box="[471,515,1333,1350]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[529,678,1333,1350]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 94142</specimenCode>
|
|||
|
|
(6, 35.9- 136.0 mm SL),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD644FAE6D8CEFAA3" box="[284,402,1361,1378]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Culuene" pageId="7" pageNumber="522">rio Culuene</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[412,532,1361,1378]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="53" value="-13.514722">13°30’53’’S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[539,655,1361,1378]" degrees="53" direction="west" minutes="5" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="40" value="-53.094444">53°5’40’’W</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD5C2FAE6D9E7FABE" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Gaucha do Norte" pageId="7" pageNumber="522">Gaúcha do Norte</location>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[241,342,1390,1407]" pageId="7" pageNumber="522" value="2007-05">
|
|||
|
|
<collectingDate box="[241,342,1390,1407]" pageId="7" pageNumber="522" value="2007-05">May 2007</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[351,492,1390,1407]" pageId="7" pageNumber="522">J. L. Birindelli</collectorName>
|
|||
|
|
<emphasis box="[497,538,1390,1407]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[550,695,1390,1407]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 94308</specimenCode>
|
|||
|
|
(1 c&s, 1, 114.0-116.0 mm SL), marginal lake,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD552FA3CDBDDFA5D" box="[522,641,1419,1436]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="rio Culuene" pageId="7" pageNumber="522">rio Culuene</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[652,771,1419,1436]" degrees="13" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="51" value="-13.514167">13°30’51’’S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[129,244,1448,1465]" degrees="53" direction="west" minutes="5" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="49" value="-53.096943">53°5’49’’W</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:03FE1801FFC4FF82D48BF962DA89FB85:8E88FFCCFFC6FF8FD7A5FA1FD8FEFA78" box="[253,418,1448,1465]" country="Brazil" county="rio Jatoba" latitude="-12.696944" longLatPrecision="21" longitude="-54.152225" municipality="rio Culuene" name="Gaucha do Norte" pageId="7" pageNumber="522">Gaúcha do Norte</location>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[472,572,1448,1465]" pageId="7" pageNumber="522" value="2007-05">
|
|||
|
|
<collectingDate box="[472,572,1448,1465]" pageId="7" pageNumber="522" value="2007-05">May 2007</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[580,719,1448,1465]" pageId="7" pageNumber="522">J. L. Birindelli</collectorName>
|
|||
|
|
<emphasis box="[724,765,1448,1465]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[129,274,1477,1494]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 94374</specimenCode>
|
|||
|
|
(4,
|
|||
|
|
<quantity box="[309,455,1477,1494]" metricMagnitude="-2" metricUnit="m" metricValue="9.965" metricValueMax="11.74" metricValueMin="8.19" pageId="7" pageNumber="522" unit="mm" value="99.65" valueMax="117.4" valueMin="81.9">81.9-117.4 mm</quantity>
|
|||
|
|
SL), lago
|
|||
|
|
<collectorName box="[554,645,1477,1494]" pageId="7" pageNumber="522">da Mirim</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality box="[653,768,1477,1494]" pageId="7" pageNumber="522">rio Culuene</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[129,247,1505,1522]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="48" value="-13.43">13°25’48’’S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[252,367,1505,1522]" degrees="53" direction="west" minutes="2" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="24" value="-53.04">53°2’24’’W</geoCoordinate>
|
|||
|
|
, Canarana, MT,
|
|||
|
|
<date box="[524,624,1505,1522]" pageId="7" pageNumber="522" value="2007-05">
|
|||
|
|
<collectingDate box="[524,624,1505,1522]" pageId="7" pageNumber="522" value="2007-05">May 2007</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[632,771,1505,1522]" pageId="7" pageNumber="522">J. L. Birindelli</collectorName>
|
|||
|
|
<emphasis box="[129,168,1534,1551]" italics="true" pageId="7" pageNumber="522">et al</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[179,321,1534,1551]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 97219</specimenCode>
|
|||
|
|
(3, 124.7-
|
|||
|
|
<quantity box="[416,511,1534,1551]" metricMagnitude="-1" metricUnit="m" metricValue="1.338" pageId="7" pageNumber="522" unit="mm" value="133.8">133.8 mm</quantity>
|
|||
|
|
SL), same data as
|
|||
|
|
<typeStatus box="[682,768,1534,1551]" pageId="7" pageNumber="522">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
<specimenCode box="[129,291,1563,1580]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="522">MZUSP 100328</specimenCode>
|
|||
|
|
(3,
|
|||
|
|
<quantity box="[335,476,1563,1580]" metricMagnitude="-2" metricUnit="m" metricValue="6.315" metricValueMax="6.59" metricValueMin="6.04" pageId="7" pageNumber="522" unit="mm" value="63.15" valueMax="65.9" valueMin="60.4">60.4-65.9 mm</quantity>
|
|||
|
|
SL), lagoas
|
|||
|
|
<collectorName box="[607,732,1563,1580]" pageId="7" pageNumber="522">Tarik Yegun</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectingCounty pageId="7" pageNumber="522">rio Jatobá</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate box="[204,324,1592,1609]" degrees="12" direction="south" minutes="41" orientation="latitude" pageId="7" pageNumber="522" precision="15" seconds="49" value="-12.696944">12°41’49’’S</geoCoordinate>
|
|||
|
|
<geoCoordinate box="[330,434,1592,1609]" degrees="54" direction="west" minutes="9" orientation="longitude" pageId="7" pageNumber="522" precision="15" seconds="8" value="-54.152225">54°9’8’’W</geoCoordinate>
|
|||
|
|
, MT,
|
|||
|
|
<date box="[494,580,1592,1609]" pageId="7" pageNumber="522" value="2008-07">
|
|||
|
|
<collectingDate box="[494,580,1592,1609]" pageId="7" pageNumber="522" value="2008-07">Jul 2008</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName box="[591,718,1592,1609]" pageId="7" pageNumber="522">M. C. Lopes</collectorName>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="7" pageNumber="522" type="diagnosis">
|
|||
|
|
<paragraph blockId="7.[129,772,1653,1960]" lastBlockId="7.[809,1452,178,584]" pageId="7" pageNumber="522">
|
|||
|
|
<emphasis bold="true" box="[129,247,1653,1672]" pageId="7" pageNumber="522">Diagnosis.</emphasis>
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[255,414,1653,1672]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[255,414,1653,1672]" italics="true" pageId="7" pageNumber="522">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is distinguished from
|
|||
|
|
<taxonomicName box="[664,771,1653,1672]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[664,771,1653,1672]" italics="true" pageId="7" pageNumber="522">H. orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName authorityName="Kindle" authorityYear="1895" box="[176,286,1685,1704]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="wilderi">
|
|||
|
|
<emphasis box="[176,286,1685,1704]" italics="true" pageId="7" pageNumber="522">H. wilderi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having two weakly-branched diverticula restricted to each side of anterior chamber of gas bladder (rarely one extra pair on posterior chambers) (
|
|||
|
|
<emphasis box="[615,645,1749,1768]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
gas bladder with many well-branched diverticula along margins of entire bladder); and gas bladder rounded posteriorly (
|
|||
|
|
<emphasis box="[618,647,1813,1832]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
gas bladder triangular posteriorly, each posterior chamber extended posteriorly into a short terminal diverticulum sharing medial septum with its pair).
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[362,518,1909,1928]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[362,518,1909,1928]" italics="true" pageId="7" pageNumber="522">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is further distinguished from
|
|||
|
|
<taxonomicName box="[190,298,1941,1960]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[190,298,1941,1960]" italics="true" pageId="7" pageNumber="522">H. orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having 11
|
|||
|
|
<superScript attach="left" box="[456,468,1938,1949]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
through 15
|
|||
|
|
<superScript attach="none" box="[598,610,1938,1949]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
, modally 13
|
|||
|
|
<superScript attach="none" box="[752,764,1938,1949]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
, midlateral scute as the anteriormost with median thorn (
|
|||
|
|
<emphasis box="[1394,1423,181,200]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
1
|
|||
|
|
<superScript attach="none" box="[1441,1451,178,189]" fontSize="6" pageId="7" pageNumber="522">st</superScript>
|
|||
|
|
through 8
|
|||
|
|
<superScript attach="right" box="[917,929,210,221]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
, modally 3
|
|||
|
|
<superScript attach="right" box="[1058,1071,210,221]" fontSize="6" pageId="7" pageNumber="522">rd</superScript>
|
|||
|
|
), tip of upper caudal-fin lobe not darkened (
|
|||
|
|
<emphasis box="[927,957,245,264]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
usually darkened), body depth at dorsal-fin origin 24.3-33.1%, mean 25.8% SL (
|
|||
|
|
<emphasis box="[1169,1197,277,296]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
16.8-22.2%, mean 21.1% SL), body depth at anal-fin origin 15.9-20.7%, mean 17.3% SL (
|
|||
|
|
<emphasis box="[818,847,341,360]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
10.0-14.7%, mean 13.1% SL), and caudal peduncle depth 6.6-8.6%, mean 7.1% SL (
|
|||
|
|
<emphasis box="[1074,1102,373,392]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
4.3-6.4%, mean 5.5% SL).
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis italics="true" pageId="7" pageNumber="522">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is distinguished from
|
|||
|
|
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1881" box="[1126,1229,405,424]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="affinis">
|
|||
|
|
<emphasis box="[1126,1229,405,424]" italics="true" pageId="7" pageNumber="522">H. affinis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having the distal tip of the first branched dorsal-fin rays and membranes pale (
|
|||
|
|
<emphasis box="[818,847,469,488]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
first branched dorsal-fin rays and membranes distally darkened); and lateral diverticula on the gas bladder fingerlike, weakly-branched in large specimens (
|
|||
|
|
<emphasis box="[1257,1286,533,552]" italics="true" pageId="7" pageNumber="522">vs.</emphasis>
|
|||
|
|
rounded, rarely branched in large specimens).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection lastPageId="10" lastPageNumber="525" pageId="7" pageNumber="522" type="description">
|
|||
|
|
<paragraph blockId="7.[809,1453,629,1960]" pageId="7" pageNumber="522">
|
|||
|
|
<emphasis bold="true" box="[809,949,629,648]" pageId="7" pageNumber="522">Description.</emphasis>
|
|||
|
|
Morphometric data are summarized in
|
|||
|
|
<tableCitation box="[1363,1451,629,648]" captionStart="Table 1" captionStartId="21.[129,192,182,201]" captionText="Table 1. Morphometric data for non-type specimens of the Hassar affinis and H. gabiru. SD = standard deviation." pageId="7" pageNumber="522">Tables 1</tableCitation>
|
|||
|
|
and
|
|||
|
|
<tableCitation box="[854,870,661,680]" captionStart="Table 2" captionStartId="22.[136,199,182,201]" captionText="Table 2. Morphometric data for type specimens of the Hassar affinis and H. gabiru. SD = standard deviation." pageId="7" pageNumber="522">2</tableCitation>
|
|||
|
|
;
|
|||
|
|
<typeStatus box="[880,974,661,680]" pageId="7" pageNumber="522">holotype</typeStatus>
|
|||
|
|
and additional specimens illustrated in
|
|||
|
|
<figureCitation captionStart="Fig" captionStartId="9.[129,168,1873,1892]" captionTargetBox="[129,1452,280,1834]" captionTargetId="figure-56@9.[129,1452,280,1834]" captionTargetPageId="9" captionText="Fig. 7. Hassar gabiru: (a) MZUSP 108440, 103.4 mm SL, holotype, rio Curuá, Altamira, PA; (b) INPA 26726, 156.6 mm SL, rio Xingu at Ilha do Babaquara, Altamira, PA; (c) MZUSP 87026, 65.3 mm SL, paratype, rio Curisevo, Gaúcha do Norte, MT." figureDoi="http://doi.org/10.5281/zenodo.5423010" httpUri="https://zenodo.org/record/5423010/files/figure.png" pageId="7" pageNumber="522">Figs. 7</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation box="[866,881,693,712]" captionStart="Fig" captionStartId="10.[136,175,757,776]" captionTargetBox="[156,1415,217,715]" captionTargetId="figure-515@10.[136,1459,193,715]" captionTargetPageId="10" captionText="Fig. 8. Hassar gabiru, paratype, MZUSP 97219, c.130 mm SL, rio Curuá, Iriri-Xingu drainage, Altamira, PA. Photo by Mark Sabaj Pérez." figureDoi="http://doi.org/10.5281/zenodo.5423012" httpUri="https://zenodo.org/record/5423012/files/figure.png" pageId="7" pageNumber="522">8</figureCitation>
|
|||
|
|
. Largest specimen examined
|
|||
|
|
<quantity box="[1168,1270,693,712]" metricMagnitude="-1" metricUnit="m" metricValue="1.566" pageId="7" pageNumber="522" unit="mm" value="156.6">156.6 mm</quantity>
|
|||
|
|
SL (INPA 26726). Dorsal profile of head rising moderately, evenly (usually in smaller specimens) or slightly convex (especially in larger specimens) from snout tip to anterior margin of orbit, and relatively straight form latter point to dorsal-fin spine. Dorsal profile of body descending gradually, approximately straight from dorsal-fin spine to caudal peduncle. Ventral contour shallowly concave from snout tip to pectoral girdle, and slightly convex from latter point to caudal peduncle. Caudal peduncle short with shallow hourglass shape in lateral view.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="7.[809,1453,629,1960]" pageId="7" pageNumber="522">Body elongate with prominent conical snout. Mouth subterminal, each premaxilla bearing small patch of approximately 5 to 10 acicular teeth, and each dentary with approximately 10 to 20 acicular teeth. Oval orbit with weakly developed adipose eyelid in juveniles and adults, extended slightly beyond anterior and posterior limits of eye. Eyes positioned about half way between tip of snout and dorsalfin origin.</paragraph>
|
|||
|
|
<paragraph blockId="7.[809,1453,629,1960]" pageId="7" pageNumber="522">Three pairs of barbels (maxillary, inner and outer mental), all fimbriate. Maxillary barbel usually reaching base of first pectoral-fin ray; with 10 to 14 (mode 12, n=13) fimbriae along ventrolateral face. Inner and outer mentonian barbels of approximately equal size, covered with many rounded papillae, and falling short of ventralmost opening of gill slit.</paragraph>
|
|||
|
|
<paragraph blockId="7.[809,1453,629,1960]" pageId="7" pageNumber="522">Gill rakers small on first gill arch, absent on remaining arches. Accessory branchial lamellae on inner face of first gill arch well developed in approximately ten rows from insertion of rakers to origin of branchial filaments (but not contacting the latter); accessory lamellae gradually reduced on remaining (second to fifth) gill arches.</paragraph>
|
|||
|
|
<paragraph blockId="7.[809,1453,629,1960]" lastBlockId="8.[136,779,1362,1960]" lastPageId="8" lastPageNumber="523" pageId="7" pageNumber="522">
|
|||
|
|
Lateral-line tubules ossified, forming a row of 30 to 33 (mode 31, n=13) midlateral scutes beginning with infranuchal. Three tympanal scutes, inconspicuous, usually without emergent thorn. Infranuchal scute with dorsal wing extremely thin and ventral wing dilated, expanded anteriorly, connected to posterior cleithral process; scute usually without medial thorn. Postinfranuchal scutes reduced anteriorly, nonoverlapping; each with posterior margin bicuspid (without medial thorn) or tricuspid (including medial thorn), latter condition usually starting at 13
|
|||
|
|
<superScript attach="left" box="[1154,1166,1938,1949]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
scute (range 11
|
|||
|
|
<superScript attach="left" box="[1346,1358,1938,1949]" fontSize="6" pageId="7" pageNumber="522">th</superScript>
|
|||
|
|
through 15
|
|||
|
|
<superScript attach="none" box="[164,176,1362,1373]" fontSize="6" pageId="8" pageNumber="523">th</superScript>
|
|||
|
|
, n=13); medial thorn and dorsal and ventral wings gradually increasing in size posteriorly; scutes with serrated posterior margins and overlapping on the posterior third of body.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.5423008" ID-Zenodo-Dep="5423008" httpUri="https://zenodo.org/record/5423008/files/figure.png" pageId="8" pageNumber="523" startId="8.[136,175,1184,1203]" targetBox="[143,1453,179,1149]" targetPageId="8">
|
|||
|
|
<paragraph blockId="8.[136,1459,1184,1267]" pageId="8" pageNumber="523">
|
|||
|
|
<emphasis bold="true" box="[136,207,1184,1203]" pageId="8" pageNumber="523">Fig. 6.</emphasis>
|
|||
|
|
Distribution map of
|
|||
|
|
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1881" box="[428,580,1184,1203]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="affinis">
|
|||
|
|
<emphasis box="[428,580,1184,1203]" italics="true" pageId="8" pageNumber="523">Hassar affinis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(yellow triangles),
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[789,893,1184,1203]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[789,893,1184,1203]" italics="true" pageId="8" pageNumber="523">H. gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(white diamonds),
|
|||
|
|
<taxonomicName box="[1099,1203,1184,1203]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[1099,1203,1184,1203]" italics="true" pageId="8" pageNumber="523">H. orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(red circles),
|
|||
|
|
<taxonomicName authorityName="Kindle" authorityYear="1895" box="[1351,1459,1184,1203]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="wilderi">
|
|||
|
|
<emphasis box="[1351,1459,1184,1203]" italics="true" pageId="8" pageNumber="523">H. wilderi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(orange pentamer). Stars represent type localities. Brown star indicates the type locality of
|
|||
|
|
<taxonomicName box="[1101,1310,1216,1235]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="ucayalensis">
|
|||
|
|
<emphasis box="[1101,1310,1216,1235]" italics="true" pageId="8" pageNumber="523">Hassar ucayalensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and pink star indicates that of
|
|||
|
|
<taxonomicName authorityName="Fowler" authorityYear="1941" box="[317,468,1248,1267]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="woodi">
|
|||
|
|
<emphasis box="[317,468,1248,1267]" italics="true" pageId="8" pageNumber="523">Hassar woodi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph blockId="8.[136,779,1362,1960]" lastBlockId="8.[816,1459,1365,1672]" pageId="8" pageNumber="523">Dorsal-fin II,6 (n=13), triangular with distal margin approximately straight, vertical when erected. Dorsal-fin spine slightly compressed and curved, with relatively small antrorse serrations along anterior margin (serrations reduced or absent on distal third); slightly larger retrorse serrations along posterior margin (serrations absent on proximal portion). Pectoral fin modally I,9, range I,8-9 (n=13); distal margin straight, oblique relative to body axis. Pectoral-fin spine slightly depressed and curved, with antrorse serrations along anterior margin (serrations absent on distal third); slightly larger retrorse serrations on posterior margin (serrations larger distally). Pelvic fin i,6 (n=13); distal margin rounded. Anal fin modally iv,9, range iii-iv,8-9 (n=13); subtriangular with scarcely rounded distal margin. Adipose fin relatively small, teardrop-shaped, with rounded free posterior margin. Caudal fin i,7/8,i (n=13, rarely i,7/7,i), distinctly forked, with lobes approximately equal in size.</paragraph>
|
|||
|
|
<paragraph blockId="8.[816,1459,1365,1672]" pageId="8" pageNumber="523">
|
|||
|
|
Gas bladder (
|
|||
|
|
<figureCitation box="[1005,1072,1429,1448]" captionStart="Fig" captionStartId="6.[136,174,1781,1800]" captionTargetBox="[136,1459,174,1761]" captionTargetId="figure-174@6.[136,1459,174,1761]" captionTargetPageId="6" captionText="Fig. 4. Gas bladders in ventral view of Hassar affinis (a-d): (a) MZUSP 90583, 54.4 mm SL; (b) MZUSP 74890, 85.9 mm SL; (c) MZUSP 43604, 228.0 mm SL; (d) ANSP 69392, 201.6 mm SL (holotype of Hassar woodi, photo by Mark Sabaj Pérez); Hassar gabiru (e-h): (e) MZUSP 87026, 69.0 mm SL; (f) MZUSP 94142, 137.6 mm SL; (g) INPA 31043, 122.5 mm SL; (h) MZUSP 94142, 128.6 mm SL; Hassar wilderi (i-l): (i) MZUSP 46016, 67.0 mm SL; (j) MZUSP 4857, 103.6 mm SL; (k) MZUSP 63148, 148.8 mm SL; (l) MZUSP 62998, 161.6 mm SL; Hassar orestis (m-p): (m) MZUSP 6991, 71.0 mm SL; (n) MZUSP 74680, 123.5 mm SL; (o) MZUSP 15512, 132.0 mm SL; (p) MZUSP 32542, 220.0 mm SL. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5423004" httpUri="https://zenodo.org/record/5423004/files/figure.png" pageId="8" pageNumber="523">Fig. 4</figureCitation>
|
|||
|
|
) moderately large, cordiform. Gas bladder walls entirely smooth in small specimens (up to
|
|||
|
|
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="523" unit="mm" value="50.0">50 mm</quantity>
|
|||
|
|
SL). Large specimens have only two short weaklybranched diverticula on each side of anterior chamber; in the largest specimens examined a third pair of diverticula are present at middle of lateral margin of posterior chambers. Gas bladder rounded posteriorly, not extended into terminal diverticula.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="8.[816,1459,1717,1960]" pageId="8" pageNumber="523">
|
|||
|
|
<emphasis bold="true" box="[816,934,1717,1736]" pageId="8" pageNumber="523">Osteology.</emphasis>
|
|||
|
|
Osteology generally similar to that described for
|
|||
|
|
<taxonomicName box="[816,962,1749,1768]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="523" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[816,962,1749,1768]" italics="true" pageId="8" pageNumber="523">Hassar orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, excepting differences as follows. Total vertebrae 35 (n=1), vertebrae 6-13 bearing ribs. Eight (n=1) dorsal-fin pterygiophores, 10 (n=1) pelvic-fin pterygiophores; 12 (n=2) dorsal and 12 (n=1) ventral caudal-fin procurrent rays. All examined specimens with hypural fusion pattern
|
|||
|
|
<collectingCountry box="[1322,1359,1877,1896]" name="Philippines" pageId="8" pageNumber="523">PH</collectingCountry>
|
|||
|
|
; HY 1+2; HY 3+4; HY 5, and with complete ossification of sutural joint at junction of sphenotic, parieto-supraoccipital and frontal.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.5423010" ID-Zenodo-Dep="5423010" httpUri="https://zenodo.org/record/5423010/files/figure.png" pageId="9" pageNumber="524" startId="9.[129,168,1873,1892]" targetBox="[129,1452,280,1834]" targetPageId="9">
|
|||
|
|
<paragraph blockId="9.[129,1451,1873,1924]" pageId="9" pageNumber="524">
|
|||
|
|
<emphasis bold="true" box="[129,196,1873,1892]" pageId="9" pageNumber="524">Fig. 7.</emphasis>
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[201,352,1873,1892]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="524" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[201,352,1873,1892]" italics="true" pageId="9" pageNumber="524">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: (
|
|||
|
|
<emphasis bold="true" box="[373,386,1873,1892]" pageId="9" pageNumber="524">a</emphasis>
|
|||
|
|
) MZUSP 108440, 103.4 mm SL, holotype, rio Curuá, Altamira, PA; (
|
|||
|
|
<emphasis bold="true" box="[1103,1118,1873,1892]" pageId="9" pageNumber="524">b</emphasis>
|
|||
|
|
) INPA 26726, 156.6 mm SL, rio Xingu at Ilha do Babaquara, Altamira, PA; (
|
|||
|
|
<emphasis bold="true" box="[587,599,1905,1924]" pageId="9" pageNumber="524">c</emphasis>
|
|||
|
|
) MZUSP 87026, 65.3 mm SL, paratype, rio Curisevo, Gaúcha do Norte, MT.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.5423012" ID-Zenodo-Dep="5423012" httpUri="https://zenodo.org/record/5423012/files/figure.png" pageId="10" pageNumber="525" startId="10.[136,175,757,776]" targetBox="[156,1415,217,715]" targetPageId="10">
|
|||
|
|
<paragraph blockId="10.[136,1406,757,808]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,206,757,776]" pageId="10" pageNumber="525">Fig. 8.</emphasis>
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[211,367,757,776]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[211,367,757,776]" italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, paratype, MZUSP 97219, c.130 mm SL, rio Curuá, Iriri-Xingu drainage, Altamira, PA. Photo by Mark Sabaj Pérez.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph blockId="10.[136,779,880,1155]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,267,880,899]" pageId="10" pageNumber="525">Coloration.</emphasis>
|
|||
|
|
In alcohol, head and body tan to brown, or grey, countershaded. Faint dark stripe from anterior margin of upper lip to anterior margin of eye. A conspicuous dark blotch on the first three branched dorsal-fin rays and membranes, blotch starting from midlength of rays and membranes and almost reaching their distal tips, which are pale. Tip of upper caudalfin lobe not conspicuously darkened.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="10.[136,779,880,1155]" pageId="10" pageNumber="525">
|
|||
|
|
In life, ground color yellowish or greenish laterally and white ventrally; lower lip pinkish; eye silvery (
|
|||
|
|
<figureCitation box="[628,692,1136,1155]" captionStart="Fig" captionStartId="10.[136,175,757,776]" captionTargetBox="[156,1415,217,715]" captionTargetId="figure-515@10.[136,1459,193,715]" captionTargetPageId="10" captionText="Fig. 8. Hassar gabiru, paratype, MZUSP 97219, c.130 mm SL, rio Curuá, Iriri-Xingu drainage, Altamira, PA. Photo by Mark Sabaj Pérez." figureDoi="http://doi.org/10.5281/zenodo.5423012" httpUri="https://zenodo.org/record/5423012/files/figure.png" pageId="10" pageNumber="525">Fig. 8</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="10" pageNumber="525" type="distribution">
|
|||
|
|
<paragraph blockId="10.[136,779,1200,1283]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,284,1200,1219]" pageId="10" pageNumber="525">Distribution.</emphasis>
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[292,449,1200,1219]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[292,449,1200,1219]" italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
occurs in the middle to upper Xingu river basin (
|
|||
|
|
<figureCitation box="[340,406,1232,1251]" captionStart="Fig" captionStartId="8.[136,175,1184,1203]" captionTargetBox="[143,1453,179,1149]" captionTargetId="figure-349@8.[136,1459,174,1156]" captionTargetPageId="8" captionText="Fig. 6. Distribution map of Hassar affinis (yellow triangles), H. gabiru (white diamonds), H. orestis (red circles), H. wilderi (orange pentamer). Stars represent type localities. Brown star indicates the type locality of Hassar ucayalensis, and pink star indicates that of Hassar woodi." figureDoi="http://doi.org/10.5281/zenodo.5423008" httpUri="https://zenodo.org/record/5423008/files/figure.png" pageId="10" pageNumber="525">Fig. 6</figureCitation>
|
|||
|
|
), and is apparently endemic to the Xingu basin above the rapids of Volta Grande, near Altamira.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="10" pageNumber="525" type="biology_ecology">
|
|||
|
|
<paragraph blockId="10.[136,779,1328,1475]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,233,1328,1347]" pageId="10" pageNumber="525">Ecology.</emphasis>
|
|||
|
|
Like its congeners,
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[461,620,1328,1347]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[461,620,1328,1347]" italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was collected usually in swift water over sand beaches at night. Camargo (2009) considered that specimens of
|
|||
|
|
<taxonomicName box="[520,673,1392,1411]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[520,673,1392,1411]" italics="true" pageId="10" pageNumber="525">Hassar orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(=
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) collected at the Volta Grande rapids have preference for benthic invertebrates and detritus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="10" pageNumber="525" type="etymology">
|
|||
|
|
<paragraph blockId="10.[136,779,1520,1699]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,260,1520,1539]" pageId="10" pageNumber="525">Etymology.</emphasis>
|
|||
|
|
Named in honor of Leandro Melo de Sousa, known to his friends as “Gabiru”, for his many contributions to the understanding of the
|
|||
|
|
<taxonomicName authorityName="Bleeker" authorityYear="1858" box="[361,476,1584,1603]" class="Actinopterygii" family="Doradidae" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="family">Doradidae</taxonomicName>
|
|||
|
|
, including his MSc and PhD dissertations (Sousa & Rapp Py-Daniel, 2005;
|
|||
|
|
<bibRefCitation author="Sousa, L" box="[632,768,1616,1635]" pageId="10" pageNumber="525" refId="ref23371" refString="Sousa, L. M. 2010. Revisao taxonomica e filogenia de Astrodoradinae (Siluriformes, Doradidae). Unpublished Ph. D. Dissertation, Universidade de Sao Paulo, Sao Paulo, 276 p." type="book" year="2010">Sousa, 2010</bibRefCitation>
|
|||
|
|
). Leandro also helped to collect part of the
|
|||
|
|
<typeStatus box="[593,639,1648,1667]" pageId="10" pageNumber="525">type</typeStatus>
|
|||
|
|
series of the new species. Treated as a noun in apposition.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="10" pageNumber="525" type="discussion">
|
|||
|
|
<paragraph blockId="10.[136,779,1744,1923]" lastBlockId="10.[816,1459,881,1092]" pageId="10" pageNumber="525">
|
|||
|
|
<emphasis bold="true" box="[136,246,1744,1763]" pageId="10" pageNumber="525">Remarks.</emphasis>
|
|||
|
|
Specimens of
|
|||
|
|
<taxonomicName authorityName="Eigenmann & Eigenmann" authorityYear="1888" box="[397,474,1744,1763]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis box="[397,474,1744,1763]" italics="true" pageId="10" pageNumber="525">Hassar</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the rio Xingu above the rapids of Volta Grande are distinct from the specimens below the rapids (including the area near Belo Monte) in overall body shape and gas bladder morphology, and are similar to the specimens from the upper Xingu river basin, herein referred to as the new species
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[459,615,1904,1923]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[459,615,1904,1923]" italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. This indicates that the Volta Grande rapids impose a distributional limit between
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[922,1037,913,932]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[922,1037,913,932]" italics="true" pageId="10" pageNumber="525">H. gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName box="[1100,1215,913,932]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="orestis">
|
|||
|
|
<emphasis box="[1100,1215,913,932]" italics="true" pageId="10" pageNumber="525">H. orestis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The identity of the specimens cited by
|
|||
|
|
<bibRefCitation author="Camargo, M" box="[1029,1270,945,964]" pageId="10" pageNumber="525" pagination="123 - 147" refId="ref21541" refString="Camargo, M, T. Giarrizzo & V. Isaac. 2004. Review of the geographic distribution of fish fauna of the Xingu river basin, Brazil. Ecotropica, 10: 123 - 147." type="journal article" year="2004">
|
|||
|
|
Camargo
|
|||
|
|
<emphasis box="[1135,1182,945,964]" italics="true" pageId="10" pageNumber="525">et al</emphasis>
|
|||
|
|
. (2004)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, Camargo (2009) and Camargo & Giarrizzo (2009) were not verified by us. However, specimens collected at the rapids of Volta Grande or upstream (putatively including rio Bacajá) are likely to be
|
|||
|
|
<taxonomicName authorityName="Birindelli & Fayal & Wosiacki" authorityYear="2011" box="[816,974,1073,1092]" class="Actinopterygii" family="Doradidae" genus="Hassar" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="gabiru">
|
|||
|
|
<emphasis box="[816,974,1073,1092]" italics="true" pageId="10" pageNumber="525">Hassar gabiru</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|