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<document ID-DOI="10.5252/a2012n1a8" ID-ISSN="1639-4798" ID-Zenodo-Dep="4599314" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1615490889614" checkinUser="carolina" docAuthor="McPHERSON, Gordon" docDate="2012" docId="E2678A6A1E33175BFEACE5FEFB83FA29" docLanguage="en" docName="Adansonia.34.1.71-75.pdf" docOrigin="Adansonia (3) 34 (1)" docSource="http://dx.doi.org/10.5252/a2012n1a8" docStyle="DocumentStyle:B98EC8A323C0F63E1C55336AD58D9273.1:Adansonia.2011-2014.journal_article" docStyleId="B98EC8A323C0F63E1C55336AD58D9273" docStyleName="Adansonia.2011-2014.journal_article" docStyleVersion="1" docTitle="Hancea inhospita McPherson 2012, sp. nov." docType="treatment" docVersion="2" lastPageId="3" lastPageNumber="74" masterDocId="1E5EF2121E321758FE06E072FFEDFF9C" masterDocTitle="A new Madagascan species of Hancea Seem. (Euphorbiaceae)" masterLastPageNumber="75" masterPageNumber="71" pageId="1" pageNumber="72" updateTime="1615492920194" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new Madagascan species of Hancea Seem. (Euphorbiaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>McPHERSON, Gordon</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Adansonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="series">
<mods:title>3</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2012-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>71</mods:start>
<mods:end>75</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.5252/a2012n1a8</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/a2012n1a8</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
<mods:identifier type="Zenodo-Dep">4599314</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29" httpUri="http://treatment.plazi.org/id/E2678A6A1E33175BFEACE5FEFB83FA29" lastPageId="3" lastPageNumber="74" pageId="1" pageNumber="72">
<subSubSection box="[170,590,1419,1446]" pageId="1" pageNumber="72" type="nomenclature">
<paragraph blockId="1.[170,590,1419,1477]" box="[170,590,1419,1446]" pageId="1" pageNumber="72">
<heading box="[170,590,1419,1446]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="72" reason="3">
<taxonomicName authority="McPherson" authorityName="McPherson" authorityYear="2012" box="[170,494,1419,1446]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="72" phylum="Tracheophyta" rank="species" species="inhospita" status="sp. nov.">
<emphasis bold="true" box="[170,360,1419,1446]" italics="true" pageId="1" pageNumber="72">Hancea inhospita</emphasis>
McPherson
</taxonomicName>
,
<taxonomicNameLabel box="[505,590,1420,1446]" pageId="1" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="74" pageId="1" pageNumber="72" type="description">
<paragraph blockId="1.[170,590,1419,1477]" box="[340,420,1451,1477]" pageId="1" pageNumber="72">
(
<figureCitation box="[347,409,1451,1477]" captionStart="FIG" captionStartId="2.[255,266,1680,1697]" captionTargetBox="[168,1155,215,1629]" captionTargetId="figure-38@2.[168,1155,215,1629]" captionTargetPageId="2" captionText="FIG. 1.— Hancea inhospita McPherson, sp. nov.: fruiting branchlets, McPherson 17461, holo-, MO." figureDoi="http://doi.org/10.5281/zenodo.4599316" httpUri="https://zenodo.org/record/4599316/files/figure.png" pageId="1" pageNumber="72">Fig. 1</figureCitation>
)
</paragraph>
<paragraph blockId="1.[111,648,1516,1699]" pageId="1" pageNumber="72">
<emphasis italics="true" pageId="1" pageNumber="72">Ab ceteris speciebus madagascariensibus cognitis foliis ovatis vel ellipticis domatiis nullis differt.Ab</emphasis>
<taxonomicName box="[453,573,1543,1567]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="72" phylum="Tracheophyta" rank="species" species="spinulosa">H. spinulosa</taxonomicName>
<emphasis italics="true" pageId="1" pageNumber="72">
laminis foliorum apice late acuminatis vel aliquando obtusis (nec anguste acuminatis), nervis in axillis domatiis non instructis, stipulis brevibus (
<quantity box="[262,335,1622,1646]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="72" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
contra
<quantity box="[398,472,1622,1646]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="72" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
), floribus staminatis bracteis brevibus (
<quantity box="[282,337,1649,1673]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="72" unit="mm" value="1.0">1 mm</quantity>
contra
<quantity box="[409,507,1649,1673]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="72" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5-3 mm</quantity>
) et staminibus paucioribus (45-50 contra 80-90) distinguenda.
</emphasis>
</paragraph>
<paragraph blockId="1.[674,1212,215,1028]" pageId="1" pageNumber="72">
TYPUS. —
<materialsCitation collectingDate="1998-02-26" collectionCode="NE, MO, K, P, TAN" country="Madagascar" elevation="1100" latitude="-18.853334" location="Moramanga" longLatPrecision="21" longitude="48.31333" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" stateProvince="Toamasina">
<emphasis bold="true" box="[788,909,215,239]" pageId="1" pageNumber="72">
<collectingCountry box="[788,909,215,239]" name="Madagascar" pageId="1" pageNumber="72">Madagascar</collectingCountry>
</emphasis>
. Prov.
<collectingRegion box="[981,1091,215,239]" country="Madagascar" name="Toamasina" pageId="1" pageNumber="72">Toamasina</collectingRegion>
,
<collectingMunicipality box="[1102,1205,215,239]" pageId="1" pageNumber="72">Ambatovy</collectingMunicipality>
,
<collectionCode box="[678,713,242,265]" country="Australia" lsid="urn:lsid:biocol.org:col:14575" name="University of New England" pageId="1" pageNumber="72" type="Herbarium">NE</collectionCode>
of
<location LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29:6F116DA71E331759FCEDE080FC89FE96" box="[747,868,242,266]" country="Madagascar" latitude="-18.853334" longLatPrecision="21" longitude="48.31333" municipality="Ambatovy" name="Moramanga" pageId="1" pageNumber="72" stateProvince="Toamasina">Moramanga</location>
, low, dense forest on lateritic soils,
<geoCoordinate box="[678,793,268,292]" degrees="18" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="12" value="-18.853334">18°51’12”S</geoCoordinate>
,
<geoCoordinate box="[805,923,268,292]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="48" value="48.31333">48°18’48”E</geoCoordinate>
,
<quantity box="[935,1016,268,292]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="72" unit="m" value="1100.0">
<elevation box="[935,1016,268,292]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="72" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date box="[1028,1137,268,292]" pageId="1" pageNumber="72" value="1998-02-26">
<collectingDate box="[1028,1137,268,292]" pageId="1" pageNumber="72" value="1998-02-26">26.II.1998</collectingDate>
</date>
, fl., fr.,
<emphasis box="[678,858,294,318]" italics="true" pageId="1" pageNumber="72">McPherson 17461</emphasis>
(holo-,
<collectionCode box="[942,988,295,319]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
! [#5888427]; iso-,
<collectionCode box="[1187,1207,295,318]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="72" type="Herbarium">K</collectionCode>
,
<collectionCode box="[678,726,321,345]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
!,
<collectionCode box="[740,752,321,344]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[761,816,321,345]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[674,1212,215,1028]" pageId="1" pageNumber="72">
<materialsCitation collectingDate="2000-10-24" collectionCode="MO, P, TAN" collectorName="de Zahamena, NW &amp; Moango &amp; Randriamanarivo" country="Madagascar" elevation="740" latitude="-17.565832" location="Prov." longLatPrecision="21" longitude="48.899445" pageId="1" pageNumber="72" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus box="[677,762,369,392]" pageId="1" pageNumber="72">PARATYPI</typeStatus>
. —
<emphasis bold="true" box="[802,916,369,393]" pageId="1" pageNumber="72">
<collectingCountry box="[802,916,369,393]" name="Madagascar" pageId="1" pageNumber="72">Madagascar</collectingCountry>
</emphasis>
.
<location LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29:6F116DA71E331759FD98E103FC22FE14" box="[926,975,369,392]" country="Madagascar" latitude="-17.565832" longLatPrecision="21" longitude="48.899445" name="Prov." pageId="1" pageNumber="72" stateProvince="Toamasina">Prov.</location>
<collectingRegion box="[976,1080,368,392]" country="Madagascar" name="Toamasina" pageId="1" pageNumber="72">Toamasina</collectingRegion>
, parc national
<collectorName box="[675,865,395,419]" pageId="1" pageNumber="72">de Zahamena, NW</collectorName>
de la rivière
<collectorName box="[994,1079,395,419]" pageId="1" pageNumber="72">Moango</collectorName>
,
<geoCoordinate box="[1089,1203,395,419]" degrees="17" direction="south" minutes="33" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="57" value="-17.565832">17°33’57”S</geoCoordinate>
,
<geoCoordinate box="[677,794,421,445]" degrees="48" direction="east" minutes="53" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="58" value="48.899445">48°53’58”E</geoCoordinate>
,
<quantity box="[805,872,422,446]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="72" unit="m" value="740.0">
<elevation box="[805,872,422,446]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="72" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<date box="[884,991,422,446]" pageId="1" pageNumber="72" value="2000-10-24">
<collectingDate box="[884,991,422,446]" pageId="1" pageNumber="72" value="2000-10-24">24.X.2000</collectingDate>
</date>
, fl., fr.,
<emphasis italics="true" pageId="1" pageNumber="72">
<collectorName pageId="1" pageNumber="72">Randriamanarivo</collectorName>
17
</emphasis>
(
<collectionCode box="[757,803,448,472]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[812,824,448,471]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[832,887,448,472]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
. —
<materialsCitation collectingDate="2004-12-18" collectionCode="MO, P, TAN" collectorName="Andriamahefarivo" country="Madagascar" elevation="1220" latitude="-18.804998" location="d'Analamay Sud" longLatPrecision="21" longitude="48.329998" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<collectingMunicipality box="[933,1034,448,472]" pageId="1" pageNumber="72">Ambatovy</collectingMunicipality>
, forêt
<location LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29:6F116DA71E331759FA40E1B2FD21FE6E" country="Madagascar" latitude="-18.804998" longLatPrecision="21" longitude="48.329998" municipality="Ambatovy" name="d'Analamay Sud" pageId="1" pageNumber="72">d’Analamay Sud</location>
,
<geoCoordinate box="[723,835,474,498]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="18" value="-18.804998">18°48’18”S</geoCoordinate>
,
<geoCoordinate box="[844,959,474,498]" degrees="48" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="48" value="48.329998">48°19’48”E</geoCoordinate>
,
<quantity box="[967,1044,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="72" unit="m" value="1220.0">
<elevation box="[967,1044,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="72" unit="m" value="1220.0">1220 m</elevation>
</quantity>
,
<date box="[1052,1175,474,498]" pageId="1" pageNumber="72" value="2004-12-18">
<collectingDate box="[1052,1175,474,498]" pageId="1" pageNumber="72" value="2004-12-18">18.XII.2004</collectingDate>
</date>
, fl.,
<emphasis box="[675,876,501,525]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[675,846,501,525]" pageId="1" pageNumber="72">Andriamahefarivo</collectorName>
47
</emphasis>
(
<collectionCode box="[888,934,501,525]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[943,955,501,524]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[962,1016,501,525]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2007-10-09" collectionCode="MO, P, TAN" collectorName="Bernard" country="Madagascar" elevation="1095" latitude="-18.806944" location="d'Analamay Sud" longLatPrecision="21" longitude="48.345276" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[1032,1078,501,525]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[1090,1204,501,525]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="25" value="-18.806944">18°48’25”S</geoCoordinate>
,
<geoCoordinate box="[675,795,527,551]" degrees="48" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="43" value="48.345276">48°20’43”E</geoCoordinate>
,
<quantity box="[807,890,527,551]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="1" pageNumber="72" unit="m" value="1095.0">
<elevation box="[807,890,527,551]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="1" pageNumber="72" unit="m" value="1095.0">1095 m</elevation>
</quantity>
,
<date box="[902,999,527,551]" pageId="1" pageNumber="72" value="2007-10-09">
<collectingDate box="[902,999,527,551]" pageId="1" pageNumber="72" value="2007-10-09">9.X.2007</collectingDate>
</date>
, fl., fr.,
<emphasis box="[1083,1208,527,551]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[1083,1161,527,551]" pageId="1" pageNumber="72">Bernard</collectorName>
602
</emphasis>
(
<collectionCode box="[684,730,554,578]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[738,750,554,577]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[757,811,554,578]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-03-31" collectionCode="MO, TAN" collectorName="Rakotovao" country="Madagascar" elevation="1068" latitude="-18.823612" location="d'Analamay Sud" longLatPrecision="21" longitude="48.334442" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[826,872,554,578]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[883,996,554,578]" degrees="18" direction="south" minutes="49" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="25" value="-18.823612">18°49’25”S</geoCoordinate>
,
<geoCoordinate box="[1004,1120,554,578]" degrees="48" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="04" value="48.334442">48°20’04”E</geoCoordinate>
,
<quantity box="[1128,1205,554,578]" metricMagnitude="3" metricUnit="m" metricValue="1.068" pageId="1" pageNumber="72" unit="m" value="1068.0">
<elevation box="[1128,1205,554,578]" metricMagnitude="3" metricUnit="m" metricValue="1.068" pageId="1" pageNumber="72" unit="m" value="1068.0">1068 m</elevation>
</quantity>
,
<date box="[675,795,580,604]" pageId="1" pageNumber="72" value="2005-03-31">
<collectingDate box="[675,795,580,604]" pageId="1" pageNumber="72" value="2005-03-31">31.III.2005</collectingDate>
</date>
, fr.,
<emphasis box="[844,1004,580,604]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[844,944,580,604]" pageId="1" pageNumber="72">Rakotovao</collectorName>
1701
</emphasis>
(
<collectionCode box="[1020,1067,580,604]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1078,1134,580,604]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-03-11" collectionCode="MO, P, TAN" collectorName="Rakotovao" country="Madagascar" elevation="1061" latitude="-18.832779" location="d'Analamay Sud" longLatPrecision="21" longitude="48.329445" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[1155,1202,580,604]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[677,788,606,630]" degrees="18" direction="south" minutes="49" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="58" value="-18.832779">18°49’58”S</geoCoordinate>
,
<geoCoordinate box="[796,910,607,631]" degrees="48" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="46" value="48.329445">48°19’46”E</geoCoordinate>
,
<quantity box="[919,995,607,631]" metricMagnitude="3" metricUnit="m" metricValue="1.061" pageId="1" pageNumber="72" unit="m" value="1061.0">
<elevation box="[919,995,607,631]" metricMagnitude="3" metricUnit="m" metricValue="1.061" pageId="1" pageNumber="72" unit="m" value="1061.0">1061 m</elevation>
</quantity>
,
<date box="[1003,1116,606,630]" pageId="1" pageNumber="72" value="2005-03-11">
<collectingDate box="[1003,1116,606,630]" pageId="1" pageNumber="72" value="2005-03-11">11.III.2005</collectingDate>
</date>
, fr.,
<emphasis italics="true" pageId="1" pageNumber="72">
<collectorName pageId="1" pageNumber="72">Rakotovao</collectorName>
1614
</emphasis>
(
<collectionCode box="[787,831,633,657]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[839,851,634,657]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[857,909,633,657]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-10-12" collectionCode="MO, P, TAN" collectorName="Razanatsoa" country="Madagascar" elevation="1124" latitude="-18.848057" location="d'Analamay Sud" longLatPrecision="21" longitude="48.308056" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[924,968,633,657]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[978,1087,633,657]" degrees="18" direction="south" minutes="50" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="53" value="-18.848057">18°50’53”S</geoCoordinate>
,
<geoCoordinate box="[1094,1206,633,657]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="29" value="48.308056">48°18’29”E</geoCoordinate>
,
<quantity box="[675,753,660,684]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="1" pageNumber="72" unit="m" value="1124.0">
<elevation box="[675,753,660,684]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="1" pageNumber="72" unit="m" value="1124.0">1124 m</elevation>
</quantity>
,
<date box="[764,871,659,683]" pageId="1" pageNumber="72" value="2005-10-12">
<collectingDate box="[764,871,659,683]" pageId="1" pageNumber="72" value="2005-10-12">12.X.2005</collectingDate>
</date>
, fl.,
<emphasis box="[913,1051,659,683]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[913,1020,660,683]" pageId="1" pageNumber="72">Razanatsoa</collectorName>
67
</emphasis>
(
<collectionCode box="[1065,1111,660,684]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1122,1134,660,683]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[1142,1196,660,684]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-02-16" collectionCode="MO, P, TAN" collectorName="Rakotovao" country="Madagascar" elevation="1109" latitude="-18.851944" location="d'Analamay Sud" longLatPrecision="21" longitude="48.296944" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[676,722,686,710]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[734,847,686,710]" degrees="18" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="07" value="-18.851944">18°51’07”S</geoCoordinate>
,
<geoCoordinate box="[856,973,686,710]" degrees="48" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="49" value="48.296944">48°17’49”E</geoCoordinate>
,
<quantity box="[981,1059,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="1" pageNumber="72" unit="m" value="1109.0">
<elevation box="[981,1059,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="1" pageNumber="72" unit="m" value="1109.0">1109 m</elevation>
</quantity>
,
<date box="[1067,1175,686,710]" pageId="1" pageNumber="72" value="2005-02-16">
<collectingDate box="[1067,1175,686,710]" pageId="1" pageNumber="72" value="2005-02-16">16.II.2005</collectingDate>
</date>
, fl., fr.,
<emphasis box="[711,866,712,737]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[711,808,712,736]" pageId="1" pageNumber="72">Rakotovao</collectorName>
1183
</emphasis>
(
<collectionCode box="[880,926,713,737]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[937,949,713,736]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[959,1013,713,737]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-03-02" collectionCode="MO, TAN" collectorName="Rakotovao" country="Madagascar" elevation="1141" latitude="-18.856112" location="d'Analamay Sud" longLatPrecision="21" longitude="48.309444" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[1031,1077,712,736]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[1091,1205,712,736]" degrees="18" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="22" value="-18.856112">18°51’22”S</geoCoordinate>
,
<geoCoordinate box="[677,795,739,763]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="34" value="48.309444">48°18’34”E</geoCoordinate>
,
<quantity box="[808,889,739,763]" metricMagnitude="3" metricUnit="m" metricValue="1.141" pageId="1" pageNumber="72" unit="m" value="1141.0">
<elevation box="[808,889,739,763]" metricMagnitude="3" metricUnit="m" metricValue="1.141" pageId="1" pageNumber="72" unit="m" value="1141.0">1141 m</elevation>
</quantity>
,
<date box="[901,1006,739,763]" pageId="1" pageNumber="72" value="2005-03-02">
<collectingDate box="[901,1006,739,763]" pageId="1" pageNumber="72" value="2005-03-02">2.III.2005</collectingDate>
</date>
, fl.,
<emphasis box="[1053,1210,739,763]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[1053,1151,739,763]" pageId="1" pageNumber="72">Rakotovao</collectorName>
1461
</emphasis>
(
<collectionCode box="[684,730,765,789]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[740,794,765,789]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1997-03-06" collectionCode="MO, P, TEF" collectorName="Rakotomalaza" country="Madagascar" elevation="1000" latitude="-18.856667" location="d'Analamay Sud" longLatPrecision="21" longitude="48.294724" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[812,858,765,789]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[872,986,765,789]" degrees="18" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="24" value="-18.856667">18°51’24”S</geoCoordinate>
,
<geoCoordinate box="[997,1114,765,789]" degrees="48" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="41" value="48.294724">48°17’41”E</geoCoordinate>
,
<quantity box="[1125,1205,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="72" unit="m" value="1000.0">
<elevation box="[1125,1205,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="72" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date box="[677,780,792,816]" pageId="1" pageNumber="72" value="1997-03-06">
<collectingDate box="[677,780,792,816]" pageId="1" pageNumber="72" value="1997-03-06">6.III.1997</collectingDate>
</date>
, fl.,
<emphasis box="[820,1007,792,816]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[820,953,792,816]" pageId="1" pageNumber="72">Rakotomalaza</collectorName>
1258
</emphasis>
(
<collectionCode box="[1020,1066,792,816]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1074,1086,792,815]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[1094,1142,792,816]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="1" pageNumber="72" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1997-02-27" collectionCode="MO, P, TEF" collectorName="Rakotomalaza" country="Madagascar" elevation="1050" latitude="-18.859446" location="d'Analamay Sud" longLatPrecision="21" longitude="48.306942" municipality="Ambatovy" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[1157,1203,792,816]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[677,791,818,842]" degrees="18" direction="south" minutes="51" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="34" value="-18.859446">18°51’34”S</geoCoordinate>
,
<geoCoordinate box="[802,919,818,842]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="25" value="48.306942">48°18’25”E</geoCoordinate>
,
<quantity box="[931,1011,818,842]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="72" unit="m" value="1050.0">
<elevation box="[931,1011,818,842]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="72" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date box="[1023,1130,818,842]" pageId="1" pageNumber="72" value="1997-02-27">
<collectingDate box="[1023,1130,818,842]" pageId="1" pageNumber="72" value="1997-02-27">27.II.1997</collectingDate>
</date>
, fl.,
<emphasis italics="true" pageId="1" pageNumber="72">
<collectorName pageId="1" pageNumber="72">Rakotomalaza</collectorName>
1196
</emphasis>
(
<collectionCode box="[854,900,845,869]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[909,921,845,868]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[929,978,845,869]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="1" pageNumber="72" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
. —
<materialsCitation collectingDate="2004-03-24" collectionCode="MO, P, TAN" collectorName="de Manombo &amp; Ludovic" country="Madagascar" elevation="21" latitude="-23.17" location="Farafangana" longLatPrecision="21" longitude="47.71389" municipality="Prov." pageId="1" pageNumber="72" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingMunicipality box="[1025,1075,845,868]" pageId="1" pageNumber="72">Prov.</collectingMunicipality>
<collectingRegion box="[1080,1206,845,869]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="72">Fianarantsoa</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29:6F116DA71E331759FCA2E315FCFBFCE3" box="[676,790,871,895]" country="Madagascar" latitude="-23.17" longLatPrecision="21" longitude="47.71389" municipality="Prov." name="Farafangana" pageId="1" pageNumber="72" stateProvince="Fianarantsoa">Farafangana</location>
, forêt
<collectorName box="[843,971,871,895]" pageId="1" pageNumber="72">de Manombo</collectorName>
,
<geoCoordinate box="[978,1087,871,895]" degrees="23" direction="south" minutes="10" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="12" value="-23.17">23°10’12”S</geoCoordinate>
,
<geoCoordinate box="[1094,1206,871,895]" degrees="47" direction="east" minutes="42" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="50" value="47.71389">47°42’50”E</geoCoordinate>
,
<quantity box="[674,728,898,922]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="72" unit="m" value="21.0">
<elevation box="[674,728,898,922]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="72" unit="m" value="21.0">21 m</elevation>
</quantity>
,
<date box="[739,856,898,922]" pageId="1" pageNumber="72" value="2004-03-24">
<collectingDate box="[739,856,898,922]" pageId="1" pageNumber="72" value="2004-03-24">24.III.2004</collectingDate>
</date>
, fl.,
<emphasis box="[899,1018,897,922]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[899,973,898,922]" pageId="1" pageNumber="72">Ludovic</collectorName>
672
</emphasis>
(
<collectionCode box="[1032,1078,898,922]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1089,1101,898,921]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[1110,1165,898,922]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
. —
<materialsCitation collectingDate="2005-11-28" collectionCode="MO, P, TAN" collectorName="Razakamalala" country="Madagascar" county="Prov. Tulear" elevation="265" latitude="-24.570557" location="d'Ivohibe" longLatPrecision="21" longitude="47.20111" municipality="Fort-Dauphin" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<collectingCounty box="[677,790,924,948]" pageId="1" pageNumber="72">Prov.Tulear</collectingCounty>
,
<collectingMunicipality box="[797,935,924,948]" pageId="1" pageNumber="72">Fort-Dauphin</collectingMunicipality>
, forêt
<location LSID="urn:lsid:plazi:treatment:E2678A6A1E33175BFEACE5FEFB83FA29:6F116DA71E331759FDE6E3EEFBD0FC28" box="[992,1085,924,948]" country="Madagascar" county="Prov. Tulear" latitude="-24.570557" longLatPrecision="21" longitude="47.20111" municipality="Fort-Dauphin" name="d'Ivohibe" pageId="1" pageNumber="72">d’Ivohibe</location>
,
<geoCoordinate box="[1093,1206,924,948]" degrees="24" direction="south" minutes="34" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="14" value="-24.570557">24°34’14”S</geoCoordinate>
,
<geoCoordinate box="[675,808,951,975]" degrees="047" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="04" value="47.20111">047°12’04”E</geoCoordinate>
,
<quantity box="[820,890,950,974]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="1" pageNumber="72" unit="m" value="265.0">
<elevation box="[820,890,950,974]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="1" pageNumber="72" unit="m" value="265.0">265 m</elevation>
</quantity>
,
<date box="[902,1021,950,974]" pageId="1" pageNumber="72" value="2005-11-28">
<collectingDate box="[902,1021,950,974]" pageId="1" pageNumber="72" value="2005-11-28">28.XI.2005</collectingDate>
</date>
, fr.,
<emphasis italics="true" pageId="1" pageNumber="72">
<collectorName box="[1070,1208,950,974]" pageId="1" pageNumber="72">Razakamalala</collectorName>
2438
</emphasis>
(
<collectionCode box="[743,789,977,1001]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[802,814,977,1000]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[825,880,977,1001]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2005-11" collectionCode="MO, P, TAN" collectorName="Razakamalala" country="Madagascar" county="Prov. Tulear" elevation="386" latitude="-24.564444" location="d'Ivohibe" longLatPrecision="21" longitude="47.19528" municipality="Fort-Dauphin" pageId="1" pageNumber="72" specimenCount="1" typeStatus="paratype">
<emphasis box="[898,945,977,1001]" italics="true" pageId="1" pageNumber="72">idem</emphasis>
,
<geoCoordinate box="[959,1075,977,1001]" degrees="24" direction="south" minutes="33" orientation="latitude" pageId="1" pageNumber="72" precision="15" seconds="52" value="-24.564444">24°33’52”S</geoCoordinate>
,
<geoCoordinate box="[1087,1205,977,1001]" degrees="47" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="72" precision="15" seconds="43" value="47.19528">47°11’43”E</geoCoordinate>
,
<quantity box="[678,742,1003,1027]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="1" pageNumber="72" unit="m" value="386.0">
<elevation box="[678,742,1003,1027]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="1" pageNumber="72" unit="m" value="386.0">386 m</elevation>
</quantity>
,
<date box="[751,834,1003,1027]" pageId="1" pageNumber="72" value="2005-11">
<collectingDate box="[751,834,1003,1027]" pageId="1" pageNumber="72" value="2005-11">XI.2005</collectingDate>
</date>
, fl.,
<emphasis box="[873,1061,1003,1027]" italics="true" pageId="1" pageNumber="72">
<collectorName box="[873,1006,1003,1027]" pageId="1" pageNumber="72">Razakamalala</collectorName>
2446
</emphasis>
(
<collectionCode box="[1073,1119,1004,1028]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="72" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1128,1140,1004,1027]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="72" type="Herbarium">P</collectionCode>
,
<collectionCode box="[1147,1202,1004,1028]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="72" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[674,1212,1072,1700]" box="[677,821,1072,1098]" pageId="1" pageNumber="72">DESCRIPTION</paragraph>
<paragraph blockId="1.[674,1212,1072,1700]" pageId="1" pageNumber="72">
Small tree
<quantity box="[792,861,1103,1129]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="1" pageNumber="72" unit="m" value="5.0" valueMax="8.0" valueMin="2.0">2-8 m</quantity>
; diameter at breast height up to
<quantity box="[678,744,1135,1161]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="72" unit="cm" value="12.0">12 cm</quantity>
; young twigs glabrous,often resinous near tips; stipules subulate,
<quantity box="[858,964,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="72" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
long, caducous. Leaves opposite to subopposite and unequal, or alternate. Blades elliptic-oblong to occasionally ovate,the smaller 1.4-6.5 ×
<quantity box="[772,889,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.8499999999999999" metricValueMax="2.8" metricValueMin="0.9" pageId="1" pageNumber="72" unit="cm" value="1.8499999999999999" valueMax="2.8" valueMin="0.9">0.9-2.8 cm</quantity>
, the larger 2.3-16×
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, somewhat coriaceous, obtuse to broadly cuneate at the base, typically rather broadly acuminate at the apex but sometimes merely acute or even obtuse (most often in smallest leaves); midrib raised on both surfaces, usually pubescent abaxially with erect hairs more than
<quantity box="[791,875,1451,1478]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="72" unit="mm" value="0.5">0.5 mm</quantity>
long, but sometimes glabrate or glabrous;secondary veins8-12on each side of the midrib, domatia absent; both surfaces without glandular granules or embedded circular glands, glabrous except for the midrib abaxially.Shorter petioles
<quantity box="[1070,1167,1578,1605]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.5" metricValueMin="0.2" pageId="1" pageNumber="72" unit="mm" value="13.5" valueMax="25.0" valueMin="2.0">2-25 mm</quantity>
, the longer
<quantity box="[745,846,1610,1636]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="4.3" metricValueMin="0.4" pageId="1" pageNumber="72" unit="mm" value="23.5" valueMax="43.0" valueMin="4.0">4-43 mm</quantity>
, narrowly canaliculate, pulvinate at both ends, often pubescent abaxially like the midrib but sometimes glabrate or glabrous.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4599316" ID-Zenodo-Dep="4599316" box="[255,1069,1680,1697]" httpUri="https://zenodo.org/record/4599316/files/figure.png" pageId="2" pageNumber="73" startId="2.[255,266,1680,1697]" targetBox="[168,1155,215,1629]" targetPageId="2">
<paragraph blockId="2.[255,1069,1680,1697]" box="[255,1069,1680,1697]" pageId="2" pageNumber="73">
FIG. 1.—
<taxonomicName authority="McPherson" authorityName="McPherson" authorityYear="2012" box="[329,575,1680,1697]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="inhospita" status="sp. nov.">
<emphasis box="[329,472,1680,1697]" italics="true" pageId="2" pageNumber="73">Hancea inhospita</emphasis>
McPherson
</taxonomicName>
,
<taxonomicNameLabel box="[583,652,1680,1697]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
: fruiting branchlets,
<emphasis box="[817,969,1680,1697]" italics="true" pageId="2" pageNumber="73">McPherson 17461</emphasis>
, holo-, MO.
</paragraph>
</caption>
<paragraph blockId="3.[111,649,216,654]" pageId="3" pageNumber="74">
Plants monoecious or (apparently) dioecious. Staminate inflorescences racemiform, axillary,
<quantity box="[113,188,279,306]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="3" pageNumber="74" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">2-5 cm</quantity>
long, resinous-scurfy, flowers solitary along the axis or in groups of 2-3, bracts up to
<quantity box="[577,646,311,337]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="74" unit="mm" value="1.0">1 mm</quantity>
long, quickly caducous, pedicels
<quantity box="[472,585,342,369]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="74" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
; buds
<quantity box="[113,227,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="74" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
in diameter; sepals 3-4,
<quantity box="[511,645,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="74" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5-2.5 mm</quantity>
long; stamens
<quantity box="[277,374,406,433]" metricMagnitude="0" metricUnit="m" metricValue="1.2065" metricValueMax="1.27" metricValueMin="1.143" pageId="3" pageNumber="74" unit="in" value="47.5" valueMax="50.0" valueMin="45.0">45-50 in</quantity>
number. Pistillate inflorescences axillary; peduncle
<quantity box="[418,495,438,464]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="3" pageNumber="74" unit="cm" value="4.5" valueMax="7.0" valueMin="2.0">2-7 cm</quantity>
long, bearing a single terminal flower; pedicel
<emphasis box="[462,471,470,495]" italics="true" pageId="3" pageNumber="74">c</emphasis>
.
<quantity box="[485,553,469,496]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="74" unit="mm" value="1.0">1 mm</quantity>
; calyx of five subulate sepals
<quantity box="[326,414,501,527]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="74" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mm</quantity>
long; styles not seen. Fruit
<emphasis box="[171,180,533,558]" italics="true" pageId="3" pageNumber="74">c</emphasis>
.
<quantity box="[192,254,532,559]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="74" unit="mm" value="6.0">6 mm</quantity>
high,
<quantity box="[318,416,533,559]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="74" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9-10 mm</quantity>
in diameter, minutely papillate-granular and puberulent-glabrate, spinulose, each sixth of the capsule bearing one spinule up to
<quantity box="[179,244,628,654]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="74" unit="mm" value="2.0">2 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="74" type="distribution">
<paragraph blockId="3.[112,646,692,844]" box="[113,428,692,718]" pageId="3" pageNumber="74">DISTRIBUTION AND ECOLOGY</paragraph>
<paragraph blockId="3.[112,646,692,844]" pageId="3" pageNumber="74">
<taxonomicName authority="McPHERSON, 2012" authorityName="McPherson" authorityYear="2012" box="[113,293,723,749]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="inhospita" status="sp. nov.">
<emphasis box="[113,293,723,749]" italics="true" pageId="3" pageNumber="74">Hancea inhospita</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[299,380,723,749]" pageId="3" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to the humid evergreen forests of eastern
<collectingCountry box="[399,525,754,780]" name="Madagascar" pageId="3" pageNumber="74">Madagascar</collectingCountry>
on lateritic soils or sand at recorded elevations from
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.205" metricValueMax="12.2" metricValueMin="0.21" pageId="3" pageNumber="74" unit="m" value="620.5" valueMax="1220.0" valueMin="21.0">21 to 1220m</quantity>
(
<figureCitation box="[208,273,818,844]" captionStart="FIG" captionStartId="4.[677,688,1138,1155]" captionTargetBox="[676,1210,215,1082]" captionTargetId="graphics-364@4.[674,1210,216,1125]" captionTargetPageId="4" captionText="FIG. 2.— Distribution of Hancea inhospita McPherson,sp.nov.(l) mapped on the bioclimatic zones of Madagascar (after Cornet 1974; see Schatz 2000)." figureDoi="http://doi.org/10.5281/zenodo.4599318" httpUri="https://zenodo.org/record/4599318/files/figure.png" pageId="3" pageNumber="74">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="74" type="discussion">
<paragraph blockId="3.[675,1210,216,623]" box="[677,777,216,242]" pageId="3" pageNumber="74">REMARKS</paragraph>
<paragraph blockId="3.[675,1210,216,623]" pageId="3" pageNumber="74">
The new species differs in leaf shape and apex from
<taxonomicName authority="(Leandri) S. E. C. Sierra, Kulju &amp; Welzen" authorityName="S. E. C. Sierra, Kulju &amp; Welzen" baseAuthorityName="Leandri" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis box="[675,867,280,305]" italics="true" pageId="3" pageNumber="74">Hancea capuronii</emphasis>
(Leandri) S.E.C.Sierra, Kulju &amp; Welzen
</taxonomicName>
, which is the only Madagascan species having obovate to oblanceolate leaves with obtuse apices. From the four other Madagascan species known,
<taxonomicName authorityName="McPherson" authorityYear="2012" box="[767,899,406,432]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="inhospita">
<emphasis box="[767,899,406,432]" italics="true" pageId="3" pageNumber="74">H. inhospita</emphasis>
</taxonomicName>
is easily distinguished by its lack of leaf domatia. It most closely resembles
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="spinulosa">
<emphasis italics="true" pageId="3" pageNumber="74">H. spinulosa</emphasis>
</taxonomicName>
but differs in having leaves with shorter and broader tips, in lacking leaf domatia entirely, and in having shorter stipules (
<quantity box="[995,1078,533,559]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="74" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
vs
<quantity box="[1109,1198,532,559]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="74" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
), shorter staminate bracts (
<quantity box="[957,1021,564,591]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="74" unit="mm" value="1.0">1 mm</quantity>
vs
<quantity box="[1060,1151,564,591]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="74" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
) and fewer stamens per staminate flower (45-50 vs
<emphasis box="[1134,1142,597,622]" italics="true" pageId="3" pageNumber="74">c</emphasis>
. 100).
</paragraph>
<paragraph blockId="3.[674,1210,660,844]" box="[674,811,660,686]" pageId="3" pageNumber="74">E TYMOLOGY</paragraph>
<paragraph blockId="3.[674,1210,660,844]" pageId="3" pageNumber="74">
The specific epithet, meaning “not providing shelter”, was suggested by the fact that
<taxonomicName authorityName="McPherson" authorityYear="2012" box="[987,1165,723,749]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="inhospita">
<emphasis box="[987,1165,723,749]" italics="true" pageId="3" pageNumber="74">Hancea inhospita</emphasis>
</taxonomicName>
can be most readily separated from its congeners with similar leaves by its lack of domatia in the axils of the leaf veins.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="74" type="key">
<paragraph blockId="3.[189,1135,892,1461]" box="[446,877,892,919]" pageId="3" pageNumber="74">
KEY TO
<taxonomicName authority="SEEM. IN" authorityName="SEEM. IN" box="[533,722,892,919]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="genus">
<emphasis box="[533,620,892,917]" italics="true" pageId="3" pageNumber="74">HANCEA</emphasis>
SEEM. IN
</taxonomicName>
<collectingCountry box="[729,877,892,918]" name="Madagascar" pageId="3" pageNumber="74">MADAGASCAR</collectingCountry>
</paragraph>
<key pageId="3" pageNumber="74">
<keyStep pageId="3" pageNumber="74">
<paragraph blockId="3.[189,1135,892,1461]" pageId="3" pageNumber="74">
<keyLead pageId="3" pageNumber="74">
1. Leaves obovate to oblanceolate, the apex obtuse ............................................................ ....................................................
<taxonomicName authority="(Leandri) S. E. C. Sierra, Kulju &amp; Welzen" authorityName="S. E. C. Sierra, Kulju &amp; Welzen" baseAuthorityName="Leandri" box="[590,1134,973,997]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis box="[590,717,974,997]" italics="true" pageId="3" pageNumber="74">H. capuronii</emphasis>
(Leandri) S.E.C.Sierra, Kulju &amp; Welzen
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[189,1135,892,1461]" box="[189,1134,1002,1026]" pageId="3" pageNumber="74">
<keyLead box="[189,1134,1002,1026]" pageId="3" pageNumber="74">— Leaves ovate or elliptic, the apex acuminate or acute, rarely obtuse ............................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="74">
<paragraph blockId="3.[189,1135,892,1461]" box="[189,1134,1053,1077]" pageId="3" pageNumber="74">
<keyLead box="[189,1134,1053,1077]" pageId="3" pageNumber="74">
2. Twigs densely pubescent ...............
<taxonomicName authority="(Baill.) S. E. C. Sierra, Kulju &amp; Welzen" authorityName="S. E. C. Sierra, Kulju &amp; Welzen" baseAuthorityName="Baill." box="[604,1134,1053,1077]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="acuminata">
<emphasis box="[604,744,1054,1077]" italics="true" pageId="3" pageNumber="74">H. acuminata</emphasis>
(Baill.) S.E.C.Sierra, Kulju &amp; Welzen
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[189,1135,892,1461]" box="[189,1134,1082,1106]" pageId="3" pageNumber="74">
<keyLead box="[189,1134,1082,1106]" pageId="3" pageNumber="74">— Twigs glabrous or obscurely puberulent ...................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="74">
<paragraph blockId="3.[189,1135,892,1461]" box="[189,1134,1133,1157]" pageId="3" pageNumber="74">
<keyLead box="[189,1134,1133,1157]" pageId="3" pageNumber="74">3. Domatia present in axils of leaf veins .......................................................................... 4</keyLead>
</paragraph>
<paragraph blockId="3.[189,1135,892,1461]" box="[189,1135,1162,1186]" pageId="3" pageNumber="74">
<keyLead box="[189,1135,1162,1186]" pageId="3" pageNumber="74">
— Domatia absent .........................................................
<taxonomicName authority="McPherson" authorityName="McPherson" authorityYear="2012" box="[795,1045,1162,1186]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="inhospita" status="sp. nov.">
<emphasis box="[795,919,1162,1186]" italics="true" pageId="3" pageNumber="74">H. inhospita</emphasis>
McPherson
</taxonomicName>
,
<taxonomicNameLabel box="[1056,1135,1163,1186]" pageId="3" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="74">
<paragraph blockId="3.[189,1135,892,1461]" pageId="3" pageNumber="74">
<keyLead pageId="3" pageNumber="74">
4. Fruit less than
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in diameter (excluding spinules); spinules up to
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long, 6-
<emphasis box="[1087,1103,1214,1237]" italics="true" pageId="3" pageNumber="74">c.</emphasis>
<quantity metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="3" pageNumber="74" unit="in" value="24.0">24 in</quantity>
number .............................
<taxonomicName authority="(McPherson) S. E. C. Sierra, Kulju &amp; Welzen" authorityName="S. E. C. Sierra, Kulju &amp; Welzen" baseAuthorityName="McPherson" box="[556,1134,1242,1266]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hancea" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="spinulosa">
<emphasis box="[556,679,1242,1266]" italics="true" pageId="3" pageNumber="74">H. spinulosa</emphasis>
(McPherson) S.E.C.Sierra, Kulju &amp; Welzen
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[189,1135,892,1461]" pageId="3" pageNumber="74">
<keyLead pageId="3" pageNumber="74">
— Fruit more than
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in diameter (excluding spines); spines
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long, more than
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number ............................................................................................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="74">
<paragraph blockId="3.[189,1135,892,1461]" pageId="3" pageNumber="74">
<keyLead pageId="3" pageNumber="74">
5. Leaf apex long-acuminate; petiole not canaliculate; known only from Ampasindava Peninsula ..............................................................................................................
<emphasis box="[1052,1079,1381,1404]" italics="true" pageId="3" pageNumber="74">H.</emphasis>
sp. 1
</keyLead>
</paragraph>
<paragraph blockId="3.[189,1135,892,1461]" pageId="3" pageNumber="74">
<keyLead pageId="3" pageNumber="74">
— Leaf apex shortly and broadly acuminate; petiole canaliculate, at least distally; known only from the summit of Mt. Vohimarangitra .................................................
<emphasis box="[1052,1079,1438,1461]" italics="true" pageId="3" pageNumber="74">H.</emphasis>
sp. 2
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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