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<document id="C21D425FB41CE1F62ADAC82DE16E3E4E" ID-CLB-Dataset="26345" ID-DOI="10.11646/zootaxa.4624.3.11" ID-GBIF-Dataset="29510c70-d277-45f5-aba5-2a779b460b37" ID-ISSN="1175-5326" ID-Zenodo-Dep="3264492" ID-ZooBank="2F12876C-2DA7-4452-9200-65F4BE1EC243" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561958693893" checkinUser="plazi" docAuthor="Michelsen, Verner &amp; Grootaert, Patrick" docDate="2019" docId="7B5E8D08AF07FFF2FF084ECC4CFD4582" docLanguage="en" docName="zootaxa.4624.3.11.pdf" docOrigin="Zootaxa 4624 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drapetis abrollensis Michelsen &amp; Grootaert 2019, sp. nov." docType="treatment" docUuid="BA913310-D251-4F8F-8024-92A3E506415B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="433" masterDocId="8767F570AF06FFF0FF9F4D354D55466E" masterDocTitle="Co-existing species of Drapetis Meigen in Skåne (S Sweden) with description of a new species and a key to males of NW European species (Diptera: Hybotidae)" masterLastPageNumber="441" masterPageNumber="431" pageNumber="432" updateTime="1698737728541" updateUser="ExternalLinkService">
<mods:mods id="5F61B51AAD9B42CC01063C288B583AAD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3467D5B46587DC3B70D73AC2D89B6778">
<mods:title id="2816811481810E65352D321ECB312BF3">Co-existing species of Drapetis Meigen in Skåne (S Sweden) with description of a new species and a key to males of NW European species (Diptera: Hybotidae)</mods:title>
</mods:titleInfo>
<mods:name id="8D748E711F7E1CC7ECAC4D1F2B9D2247" type="personal">
<mods:role id="47668D31FAAB80E6CD70C4CC09487688">
<mods:roleTerm id="7B95C81F828DE9BE68077846817E0650">Author</mods:roleTerm>
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<mods:namePart id="8A96D9FD3F73DB952885B35361E87DB6">Michelsen, Verner</mods:namePart>
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<mods:role id="268D051E875AC3DF35C3F4326D0F707A">
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<mods:namePart id="3E24AE2535E08942D0FB81B51ECE5A79">Grootaert, Patrick</mods:namePart>
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<mods:typeOfResource id="172CD359A12D892E54271E8DE4E6B91F">text</mods:typeOfResource>
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<mods:title id="9DB79D9FA64B26B88B46C05466609D02">Zootaxa</mods:title>
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<mods:part id="1377080F261DA50AC009230390FA949B">
<mods:date id="2B217A5299C0D88C99891AC7EACD6351">2019</mods:date>
<mods:detail id="6AE3EB2F863F490F36AD2B8866A3DF23" type="pubDate">
<mods:number id="9A4C5239092C76695937EC60ED18CBA7">2019-07-01</mods:number>
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<mods:detail id="F6E526624DA8B404528C70464E988553" type="volume">
<mods:number id="A9A0507AF346AEDE56E2CDA60FFFD7E9">4624</mods:number>
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<mods:detail id="7405EF32D0B3596372581AB3E17AC08E" type="issue">
<mods:number id="BF4B9C98016A34D982C66A6928889C65">3</mods:number>
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<mods:start id="7A37D27DBAFC412C1606D929B3B32A22">431</mods:start>
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<mods:classification id="B3F8CA6194D9C095841DDCA25578E9F1">journal article</mods:classification>
<mods:identifier id="EB111D6D2FB1DE3924B62FB0EF2576F6" type="CLB-Dataset">26345</mods:identifier>
<mods:identifier id="18AD719B09E65751ED88E07E5E222144" type="DOI">10.11646/zootaxa.4624.3.11</mods:identifier>
<mods:identifier id="77086830550F1B95967B75394E3718E6" type="GBIF-Dataset">29510c70-d277-45f5-aba5-2a779b460b37</mods:identifier>
<mods:identifier id="B9D8B55529ACA324C89DC22492F999C9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="584458613F851987C419789E2A4A310E" type="Zenodo-Dep">3264492</mods:identifier>
<mods:identifier id="4122C0214A0A64FAB59BB6B7B741A4C8" type="ZooBank">2F12876C-2DA7-4452-9200-65F4BE1EC243</mods:identifier>
</mods:mods>
<treatment id="7B5E8D08AF07FFF2FF084ECC4CFD4582" ID-GBIF-Taxon="158485174" LSID="urn:lsid:zoobank.org:act:BA913310-D251-4F8F-8024-92A3E506415B" httpUri="http://treatment.plazi.org/id/7B5E8D08AF07FFF2FF084ECC4CFD4582" lastPageId="2" lastPageNumber="433" pageId="1" pageNumber="432">
<subSubSection id="BBED6F95AF07FFF1FF084ECC4C8A427A" box="[151,479,1017,1044]" pageId="1" pageNumber="432" type="nomenclature">
<paragraph id="F3483C1EAF07FFF1FF084ECC4C8A427A" blockId="1.[151,479,1017,1079]" box="[151,479,1017,1044]" pageId="1" pageNumber="432">
<heading id="A8008B72AF07FFF1FF084ECC4C8A427A" bold="true" box="[151,479,1017,1044]" fontSize="11" level="1" pageId="1" pageNumber="432" reason="1">
<emphasis id="C183E00CAF07FFF1FF084ECC4C8A427A" bold="true" box="[151,479,1017,1044]" pageId="1" pageNumber="432">
<taxonomicName id="34F7479DAF07FFF1FF084ECC4C2B427A" authority="Michelsen &amp; Grootaert, 2019" authorityName="Michelsen &amp; Grootaert" authorityYear="2019" box="[151,382,1017,1044]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="abrollensis" status="sp. nov.">
<emphasis id="C183E00CAF07FFF1FF084ECC4C2B427A" bold="true" box="[151,382,1017,1044]" italics="true" pageId="1" pageNumber="432">Drapetis abrollensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAB05D77AF07FFF1FE1A4ECF4C8A427A" box="[389,479,1018,1044]" pageId="1" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF07FFF1FF0849284C834259" box="[151,470,1053,1079]" pageId="1" pageNumber="432" type="description">
<paragraph id="F3483C1EAF07FFF1FF0849284C834259" blockId="1.[151,479,1017,1079]" box="[151,470,1053,1079]" pageId="1" pageNumber="432">
(
<figureCitation id="6BCC209BAF07FFF1FF0049284DB44259" box="[159,225,1053,1079]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="1" pageNumber="432">Figs 1</figureCitation>
A–D, 2B, 3C, 4A–D)
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF07FFF1FF0849504EDC4282" pageId="1" pageNumber="432" type="materials_examined">
<paragraph id="F3483C1EAF07FFF1FF0849504EDC4282" blockId="1.[151,1437,1125,2016]" pageId="1" pageNumber="432">
<emphasis id="C183E00CAF07FFF1FF0849504CD542EE" bold="true" box="[151,384,1125,1152]" pageId="1" pageNumber="432">Material examined.</emphasis>
<materialsCitation id="439F3643AF07FFF1FE1A49504C9E42CD" ID-GBIF-Occurrence="2273188155" collectingDate="2016-06-11" collectingDateMax="2016-06-13" collectingDateMin="2016-06-11" collectionCode="NHMD" collectorName="V. Michelsen" country="Sweden" elevation="142" latitude="56.433334" location="Osby" longLatPrecision="1057" longitude="14.133333" municipality="Abrolla" pageId="1" pageNumber="432" specimenCount="1" specimenCount-male="1" stateProvince="Skane" typeStatus="holotype">
<typeStatus id="2C4C82BCAF07FFF1FE1A49504F7142EE" box="[389,548,1125,1152]" pageId="1" pageNumber="432" type="holotype">
<emphasis id="C183E00CAF07FFF1FE1A49504F7142EE" bold="true" box="[389,548,1125,1152]" pageId="1" pageNumber="432">HOLOTYPE</emphasis>
</typeStatus>
<specimenCount id="E5F1F797AF07FFF1FDB549534F6A42EE" box="[554,575,1126,1152]" pageId="1" pageNumber="432" type="male">♂</specimenCount>
,
<collectingCountry id="8BE07C8EAF07FFF1FDD349504F904211" box="[588,709,1125,1151]" name="Sweden" pageId="1" pageNumber="432">SWEDEN</collectingCountry>
:
<collectingRegion id="3133F2FCAF07FFF1FD5149504E4242EE" box="[718,791,1125,1152]" country="Sweden" name="Skane" pageId="1" pageNumber="432">Skåne</collectingRegion>
:
<collectingMunicipality id="132CA664AF07FFF1FCBE49504E2C42EE" box="[801,889,1125,1152]" pageId="1" pageNumber="432">Åbrolla</collectingMunicipality>
,
<location id="F6286AC5AF07FFF1FC1C49504EEB4211" LSID="urn:lsid:plazi:treatment:7B5E8D08AF07FFF2FF084ECC4CFD4582:F6286AC5AF07FFF1FC1C49504EEB4211" box="[899,958,1125,1151]" country="Sweden" latitude="56.433334" longLatPrecision="1057" longitude="14.133333" municipality="Abrolla" name="Osby" pageId="1" pageNumber="432" stateProvince="Skane">Osby</location>
(
<geoCoordinate id="96C35AD9AF07FFF1FC534950497E42EE" box="[972,1067,1125,1152]" degrees="56" direction="north" minutes="26" orientation="latitude" pageId="1" pageNumber="432" precision="925" value="56.433334">56°26′N</geoCoordinate>
<geoCoordinate id="96C35AD9AF07FFF1FBAE495049DA42EE" box="[1073,1167,1125,1152]" degrees="14" direction="east" minutes="08" orientation="longitude" pageId="1" pageNumber="432" precision="925" value="14.133333">14°08′E</geoCoordinate>
),
<elevation id="78DADB2DAF07FFF1FB3D495049BF42EE" box="[1186,1258,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="432" unit="m" value="142.0">
<quantity id="340F91FBAF07FFF1FB3D495049BF42EE" box="[1186,1258,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="432" unit="m" value="142.0">142 m</quantity>
</elevation>
,
<collectingDate id="970DE336AF07FFF1FB6B495048C24211" box="[1268,1431,1125,1151]" pageId="1" pageNumber="432" value="2016-06-11" valueMax="2016-06-13" valueMin="2016-06-11">
<date id="87491ADEAF07FFF1FB6B495048C24211" box="[1268,1431,1125,1151]" pageId="1" pageNumber="432" value="2016-06-11" valueMax="2016-06-13" valueMin="2016-06-11">11–13.vi.2016</date>
</collectingDate>
, leg.
<collectorName id="5E0259C8AF07FFF1FF5949BF4C0C42CD" box="[198,345,1161,1188]" pageId="1" pageNumber="432">V. Michelsen</collectorName>
(
<collectionCode id="95E6A4DBAF07FFF1FEF849BC4C9642CD" box="[359,451,1161,1187]" pageId="1" pageNumber="432">NHMD</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2C4C82BCAF07FFF1FE4B49BC4F6942CD" box="[468,572,1161,1187]" pageId="1" pageNumber="432" type="holotype">Holotype</typeStatus>
condition: micro-pinned from the left and staged; abdomen dissected (middle segments missing); dissected parts stored in glycerin in a microvial with the specimen; right postpedicel, right wing and right midleg missing; left midleg glued separately on the stage.
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF07FFF1FF5849C349D8437E" box="[199,1165,1269,1296]" pageId="1" pageNumber="432" type="etymology">
<paragraph id="F3483C1EAF07FFF1FF5849C349D8437E" blockId="1.[151,1437,1125,2016]" box="[199,1165,1269,1296]" pageId="1" pageNumber="432">
<emphasis id="C183E00CAF07FFF1FF5849C34C1B437E" bold="true" box="[199,334,1270,1296]" pageId="1" pageNumber="432">Etymology.</emphasis>
The species epithet is derived from the name of the
<typeStatus id="2C4C82BCAF07FFF1FC0749C34E9D437E" box="[920,968,1270,1296]" pageId="1" pageNumber="432">type</typeStatus>
locality, Åbrolla.
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF07FFF1FF58482C4F614065" pageId="1" pageNumber="432" type="diagnosis">
<paragraph id="F3483C1EAF07FFF1FF58482C4F614065" blockId="1.[151,1437,1125,2016]" pageId="1" pageNumber="432">
<emphasis id="C183E00CAF07FFF1FF58482C4C6F435A" bold="true" box="[199,314,1305,1332]" pageId="1" pageNumber="432">Diagnosis</emphasis>
(male): Radial medial crossvein (r-m) lying distinctly beyond middle of basal medial cell (bm) (
<figureCitation id="6BCC209BAF07FFF1FAF0482C4DEE4339" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1985]" captionTargetBox="[334,1249,595,1923]" captionTargetId="figure@3.[324,1264,587,1937]" captionTargetPageId="3" captionText="FIGURE 2. Drapetis males. Right wing. A. D. assimilis (Fallén); B. D. abrollensis sp. nov.; C. D. incompleta Collin. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264498/files/figure.png" pageId="1" pageNumber="432">Fig. 2B</figureCitation>
). Antenna brownish black; pedicel beneath with short distal setulae; postpedicel subtriangular, slightly longer than wide. Prothorax except on epimeron greyish pruinose; proepisternum without upturned seta. Halter blackish on knob. Legs light brownish. Mid femur simple except for subapical field of scaly cuticle (
<figureCitation id="6BCC209BAF07FFF1FB3E48B049BC43F1" box="[1185,1257,1413,1439]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1571]" captionTargetBox="[153,1436,181,1520]" captionTargetId="figure@5.[151,1436,181,1520]" captionTargetPageId="5" captionText="FIGURE 4. Drapetis abrollensis sp. nov. Right midleg, SEM images showing modified cuticle. A. Femur and tibia; B. Subapical cuticle of femur; C. Ventral edge of tibia; D. Detail of C showing accumulations of brochosomes. Scales: A = 0.1 mm; C = 1 μm." httpUri="https://zenodo.org/record/3264504/files/figure.png" pageId="1" pageNumber="432">Figs 4</figureCitation>
A–B). Mid tibia ventrally expanded and compressed (
<figureCitation id="6BCC209BAF07FFF1FDA3489C4FC043AD" box="[572,661,1449,1475]" captionStart="FIGURE 3" captionStartId="4.[151,250,1431,1456]" captionTargetBox="[430,1255,239,1406]" captionTargetId="figure@4.[332,1257,200,1409]" captionTargetPageId="4" captionText="FIGURE 3. Drapetis males. Legs. A. D. assimilis (Fallén), right hind femur; B. D. pusilla Loew, left fore femur; C. D. abrollensis sp. nov., right mid femur and tibia. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264500/files/figure.png" pageId="1" pageNumber="432">Fig. 3C</figureCitation>
), at ventral edge bearing numerous cuticular denticles (
<figureCitation id="6BCC209BAF07FFF1FA92489C48DB43AD" box="[1293,1422,1449,1475]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1571]" captionTargetBox="[153,1436,181,1520]" captionTargetId="figure@5.[151,1436,181,1520]" captionTargetPageId="5" captionText="FIGURE 4. Drapetis abrollensis sp. nov. Right midleg, SEM images showing modified cuticle. A. Femur and tibia; B. Subapical cuticle of femur; C. Ventral edge of tibia; D. Detail of C showing accumulations of brochosomes. Scales: A = 0.1 mm; C = 1 μm." httpUri="https://zenodo.org/record/3264504/files/figure.png" pageId="1" pageNumber="432">Figs 4A, C</figureCitation>
); posterior side of tibia with a longitudinal impression (
<figureCitation id="6BCC209BAF07FFF1FD6548F84E014389" box="[762,852,1485,1511]" captionStart="FIGURE 3" captionStartId="4.[151,250,1431,1456]" captionTargetBox="[430,1255,239,1406]" captionTargetId="figure@4.[332,1257,200,1409]" captionTargetPageId="4" captionText="FIGURE 3. Drapetis males. Legs. A. D. assimilis (Fallén), right hind femur; B. D. pusilla Loew, left fore femur; C. D. abrollensis sp. nov., right mid femur and tibia. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264500/files/figure.png" pageId="1" pageNumber="432">Fig. 3C</figureCitation>
). Apical margin of right epandrial lamella with two narrow projections (
<figureCitation id="6BCC209BAF07FFF1FEE648C44CAE4065" box="[377,507,1521,1547]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="1" pageNumber="432">Fig. 1A, p1</figureCitation>
, p
<figureCitation id="6BCC209BAF07FFF1FD8748C44F724065" box="[536,551,1521,1547]" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1985]" captionTargetBox="[334,1249,595,1923]" captionTargetId="figure@3.[324,1264,587,1937]" captionTargetPageId="3" captionText="FIGURE 2. Drapetis males. Right wing. A. D. assimilis (Fallén); B. D. abrollensis sp. nov.; C. D. incompleta Collin. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264498/files/figure.png" pageId="1" pageNumber="432">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF07FFF2FF584B204D9145CB" lastPageId="2" lastPageNumber="433" pageId="1" pageNumber="432" type="description">
<paragraph id="F3483C1EAF07FFF2FF584B204C7E44A1" blockId="1.[151,1437,1125,2016]" lastBlockId="2.[151,1437,152,1004]" lastPageId="2" lastPageNumber="433" pageId="1" pageNumber="432">
<emphasis id="C183E00CAF07FFF1FF584B204CF7405E" bold="true" box="[199,418,1557,1584]" pageId="1" pageNumber="432">Description. Male.</emphasis>
Small sized: wing length
<quantity id="340F91FBAF07FFF1FD5E4B204E43405E" box="[705,790,1557,1584]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="432" unit="mm" value="1.3">1.3 mm</quantity>
(n = 1).
<emphasis id="C183E00CAF07FFF1FCEC4B234EE9405E" bold="true" box="[883,956,1558,1584]" pageId="1" pageNumber="432">Head:</emphasis>
Black in ground-colour, shiny through thin grey pruinosity. Frons narrowed downwards; face linear, narrower than frons, widening immediately above clypeus. All setulae and setae on head and antennae dark, on palpi and labella pale yellowish. Anterior ocellars shorter than postpedicel, crossing; median ocellars same size, diverging. Pair of long, convergent (inner) vertical setae flanked by two convergent pairs of postocular setae. Occiput densely set with setulae, on postgena with pair of fine setae. Antenna brownish black, ventro-apical setulae on pedicel shorter than length of pedicel; postpedicel slightly longer than wide, subtriangular, with apical stylus about 2.2 times as long as remaining antenna.
<emphasis id="C183E00CAF07FFF1FBF14BDB49854166" bold="true" box="[1134,1232,1774,1800]" pageId="1" pageNumber="432">Thorax:</emphasis>
Extensively shiny black, except grey pruinose on prothorax other than on proepimeron, scutellum and upper margins of anepisternum. No upturned seta on proepisternum. Mesonotum densely set with setulae; acrostichals not distinct from surrounding setulae. Prominent setae: a pair of prescutellar dorsocentrals, 4 notopleurals, 2 pairs of scutellars, apical pair longer and crossing.
<emphasis id="C183E00CAF07FFF1FEAF4A484C2F41F6" bold="true" box="[304,378,1917,1944]" pageId="1" pageNumber="432">Wing:</emphasis>
Hyaline, with brown veins (
<figureCitation id="6BCC209BAF07FFF1FD2C4A484E5F41F9" box="[691,778,1917,1943]" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1985]" captionTargetBox="[334,1249,595,1923]" captionTargetId="figure@3.[324,1264,587,1937]" captionTargetPageId="3" captionText="FIGURE 2. Drapetis males. Right wing. A. D. assimilis (Fallén); B. D. abrollensis sp. nov.; C. D. incompleta Collin. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264498/files/figure.png" pageId="1" pageNumber="432">Fig. 2B</figureCitation>
). Vein R
<subScript id="6F733E5BAF07FFF1FCF74ABA4ED741F3" attach="left" box="[872,898,1935,1949]" fontSize="6" pageId="1" pageNumber="432">2+3</subScript>
almost parallel with vein R
<subScript id="6F733E5BAF07FFF1FB2D4ABA499941F3" attach="left" box="[1202,1228,1935,1949]" fontSize="6" pageId="1" pageNumber="432">4+5</subScript>
on basal half; vein R
<subScript id="6F733E5BAF07FFF1FF354A864D9141AF" attach="left" box="[170,196,1971,1985]" fontSize="6" pageId="1" pageNumber="432">4+5</subScript>
slightly sigmoid before joining costa well before wing apex; M
<subScript id="6F733E5BAF07FFF1FC164A864EC441AF" attach="left" box="[905,913,1971,1985]" fontSize="6" pageId="1" pageNumber="432">1</subScript>
curved on basal part but straight on distal half, meeting costa only slightly posterior of wing apex. Distance between cross-veins r–m and bm–m 0.6 times length of basal portion of vein M
<subScript id="6F733E5BAF04FFF2FE044D9F4CF646D6" attach="left" box="[411,419,170,184]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
and 0.35 times length of apical section of vein Cu. Calypter brownish black with long, dark marginal setae. Halter knob blackish, pale yellowish on stalk.
<emphasis id="C183E00CAF04FFF2FCD24D884EDA46B9" bold="true" box="[845,911,189,215]" pageId="2" pageNumber="433">Legs:</emphasis>
Light brownish. Chaetotaxy mainly consisting of short pale setulae; even few setae pale yellowish, except for some relatively strong, dark anterior setae on mid coxa. Fore femur stout, decidedly thicker than mid femur, slightly thicker than hind femur. Anterior surface of mid femur simple except for subapical field of scaly cuticle (
<figureCitation id="6BCC209BAF04FFF2FC8B4C1C4EC6472D" box="[788,915,297,323]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1571]" captionTargetBox="[153,1436,181,1520]" captionTargetId="figure@5.[151,1436,181,1520]" captionTargetPageId="5" captionText="FIGURE 4. Drapetis abrollensis sp. nov. Right midleg, SEM images showing modified cuticle. A. Femur and tibia; B. Subapical cuticle of femur; C. Ventral edge of tibia; D. Detail of C showing accumulations of brochosomes. Scales: A = 0.1 mm; C = 1 μm." httpUri="https://zenodo.org/record/3264504/files/figure.png" pageId="2" pageNumber="433">Figs 4A, B</figureCitation>
). Mid tibia ventrally crest-like expanded (
<figureCitation id="6BCC209BAF04FFF2FAF04C1C4DEF4709" captionStart="FIGURE 3" captionStartId="4.[151,250,1431,1456]" captionTargetBox="[430,1255,239,1406]" captionTargetId="figure@4.[332,1257,200,1409]" captionTargetPageId="4" captionText="FIGURE 3. Drapetis males. Legs. A. D. assimilis (Fallén), right hind femur; B. D. pusilla Loew, left fore femur; C. D. abrollensis sp. nov., right mid femur and tibia. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264500/files/figure.png" pageId="2" pageNumber="433">Fig. 3C</figureCitation>
), with numerous cuticular denticles along ventral edge spreading onto anterior surface (
<figureCitation id="6BCC209BAF04FFF2FB154C78485D4709" box="[1162,1288,333,359]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1571]" captionTargetBox="[153,1436,181,1520]" captionTargetId="figure@5.[151,1436,181,1520]" captionTargetPageId="5" captionText="FIGURE 4. Drapetis abrollensis sp. nov. Right midleg, SEM images showing modified cuticle. A. Femur and tibia; B. Subapical cuticle of femur; C. Ventral edge of tibia; D. Detail of C showing accumulations of brochosomes. Scales: A = 0.1 mm; C = 1 μm." httpUri="https://zenodo.org/record/3264504/files/figure.png" pageId="2" pageNumber="433">Figs 4A, C</figureCitation>
). Hind femur distinctly curved outwards. Hind tarsomere 1 slightly swollen, as long as combined length of hind tarsomeres 2–4.
<emphasis id="C183E00CAF04FFF2FF084CA14C4047C1" bold="true" box="[151,277,404,431]" pageId="2" pageNumber="433">Abdomen:</emphasis>
Basal sclerites brownish black, rather shiny through thin pruinosity. Tergite I not sclerotized, sternite I very short. Middle part of preabdomen destroyed during dissection.
<emphasis id="C183E00CAF04FFF2FC004C8C493547BD" bold="true" box="[927,1120,440,467]" pageId="2" pageNumber="433">Male terminalia</emphasis>
(
<figureCitation id="6BCC209BAF04FFF2FBEE4C8C49E047BD" box="[1137,1205,441,467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Figs 1</figureCitation>
A–D): Right sursty- lus fused with right epandrial lamella; right epandrial lamella with 2 projections (p1, p2) from apical border (
<figureCitation id="6BCC209BAF04FFF2FAF54CE84D884475" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Figs 1A, B</figureCitation>
). Cerci (
<figureCitation id="6BCC209BAF04FFF2FEDF4F344CC34475" box="[320,406,513,539]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Fig. 1B</figureCitation>
) fused mediodistally, tips of smaller right cercus (rc) and larger left cercus (lc) still clearly indicated. Left epandrial lamella (lel) fused with hypandrium (hy), with 5 setae on apical border (
<figureCitation id="6BCC209BAF04FFF2FB4E4F1048724451" box="[1233,1319,549,575]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Fig. 1B</figureCitation>
). Left surstylus 1 (ls1) pale brown, only with a few small ventral setae (
<figureCitation id="6BCC209BAF04FFF2FCCA4F7C4E81440D" box="[853,980,585,611]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Figs 1B, C</figureCitation>
). Left surstylus 2 (ls2) large, enveloping apical part of left epandrial lamella (lel), with median apical projection bearing distinct apical seta (
<figureCitation id="6BCC209BAF04FFF2FB724F58481144E9" box="[1261,1348,621,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Fig. 1C</figureCitation>
). Tip of hypandrium bearing stout black seta at tip (
<figureCitation id="6BCC209BAF04FFF2FD1D4FA44F8944C5" box="[642,732,657,683]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Fig. 1D</figureCitation>
). Ejaculatory apodeme (ej) arising from long ventral apodeme (
<figureCitation id="6BCC209BAF04FFF2FF004F804C4B44A1" box="[159,286,693,719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1800,1825]" captionTargetBox="[184,1391,1142,1738]" captionTargetId="figure@2.[183,1404,1024,1773]" captionTargetPageId="2" captionText="FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm." httpUri="https://zenodo.org/record/3264496/files/figure.png" pageId="2" pageNumber="433">Figs 1B, D</figureCitation>
).
</paragraph>
<paragraph id="F3483C1EAF04FFF2FF584FED4D9145CB" blockId="2.[151,1437,152,1004]" pageId="2" pageNumber="433">
<emphasis id="C183E00CAF04FFF2FF584FED4CC2449D" bold="true" box="[199,407,728,755]" pageId="2" pageNumber="433">Taxonomic notes.</emphasis>
The close-set cross-veins r–m and bm–m assign
<taxonomicName id="34F7479DAF04FFF2FC214FEF49F7449D" authority="Michelsen &amp; Grootaert, 2019" authorityName="Michelsen &amp; Grootaert" authorityYear="2019" box="[958,1186,729,755]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="abrollensis" status="sp. nov.">
<emphasis id="C183E00CAF04FFF2FC214FEF49F7449D" box="[958,1186,729,755]" italics="true" pageId="2" pageNumber="433">Drapetis abrollensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAB05D77AF04FFF2FB364FEC4856449D" box="[1193,1283,729,755]" pageId="2" pageNumber="433" rank="species">
<emphasis id="C183E00CAF04FFF2FB364FEC4856449D" bold="true" box="[1193,1283,729,755]" pageId="2" pageNumber="433">sp. nov.</emphasis>
</taxonomicNameLabel>
to the
<taxonomicName id="34F7479DAF04FFF2FACF4FEF4DE44579" authorityName="Meigen" authorityYear="1822" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="C183E00CAF04FFF2FACF4FEF4831449D" box="[1360,1380,730,755]" italics="true" pageId="2" pageNumber="433">D</emphasis>
.
<emphasis id="C183E00CAF04FFF2FAED4FEF4DE44579" italics="true" pageId="2" pageNumber="433">exilis</emphasis>
</taxonomicName>
-group as defined by
<bibRefCitation id="976641EFAF04FFF2FE004FC84F114579" author="Chvala, M." box="[415,580,764,791]" pageId="2" pageNumber="433" pagination="1 - 336" refId="ref6525" refString="Chvala, M. (1975) The Tachydromiinae (Dipt. Empididae) of Fennoscandia and Denmark. Fauna entomologica scandinavica, 3, 1 - 336. http: // dx. doi. org / 10.1163 / 187631272 X 00012" type="journal article" year="1975">Chvála (1975)</bibRefCitation>
. The anterior surface of the male mid femur is lacking a longitudinal field of vertical ribs or furrows and has a normal cover of fine setulae. Within the
<taxonomicName id="34F7479DAF04FFF2FC7E4E1449494555" authorityName="Meigen" authorityYear="1822" box="[993,1052,801,827]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="C183E00CAF04FFF2FC7E4E1449494555" box="[993,1052,801,827]" italics="true" pageId="2" pageNumber="433">exilis</emphasis>
</taxonomicName>
-group a simple mid femur is also seen in males of
<taxonomicName id="34F7479DAF04FFF2FECB4E714F114533" authority="Kovalev" authorityName="Kovalev" authorityYear="1972" box="[340,580,835,862]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="completa">
<emphasis id="C183E00CAF04FFF2FECB4E714C884530" box="[340,477,836,862]" italics="true" pageId="2" pageNumber="433">D. completa</emphasis>
Kovalev
</taxonomicName>
,
<taxonomicName id="34F7479DAF04FFF2FDCF4E714E694533" authority="Collin" authorityName="Collin. Scale" authorityYear="1926" box="[592,828,835,862]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="incompleta">
<emphasis id="C183E00CAF04FFF2FDCF4E714FBB4530" box="[592,750,836,862]" italics="true" pageId="2" pageNumber="433">D. incompleta</emphasis>
Collin
</taxonomicName>
and
<taxonomicName id="34F7479DAF04FFF2FCED4E7149DD4533" authority="Kovalev." authorityName="Kovalev." box="[882,1160,835,862]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="stackelbergi">
<emphasis id="C183E00CAF04FFF2FCED4E7149484530" box="[882,1053,836,862]" italics="true" pageId="2" pageNumber="433">D. stackelbergi</emphasis>
Kovalev.
</taxonomicName>
<taxonomicName id="34F7479DAF04FFF2FB104E7148264530" authority="Michelsen &amp; Grootaert, 2019" authorityName="Michelsen &amp; Grootaert" authorityYear="2019" box="[1167,1395,836,862]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="abrollensis" status="sp. nov.">
<emphasis id="C183E00CAF04FFF2FB104E7148264530" box="[1167,1395,836,862]" italics="true" pageId="2" pageNumber="433">Drapetis abrollensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAB05D77AF04FFF2FAE44E714D9F45EE" pageId="2" pageNumber="433" rank="species">
<emphasis id="C183E00CAF04FFF2FAE44E714D9645EE" bold="true" pageId="2" pageNumber="433">sp. nov</emphasis>
.
</taxonomicNameLabel>
is to our knowledge the only species of the
<taxonomicName id="34F7479DAF04FFF2FD2A4E524E4745EE" authorityName="Meigen" authorityYear="1822" box="[693,786,870,897]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="C183E00CAF04FFF2FD2A4E524F9C45EE" box="[693,713,871,896]" italics="true" pageId="2" pageNumber="433">D</emphasis>
.
<emphasis id="C183E00CAF04FFF2FD484E534E4745EE" box="[727,786,870,896]" italics="true" pageId="2" pageNumber="433">exilis</emphasis>
</taxonomicName>
-group reported to have the mid tibia modified in the male sex.
</paragraph>
</subSubSection>
<subSubSection id="BBED6F95AF04FFF2FF584E984CFD4582" pageId="2" pageNumber="433" type="distribution">
<paragraph id="F3483C1EAF04FFF2FF584E984CFD4582" blockId="2.[151,1437,152,1004]" pageId="2" pageNumber="433">
<emphasis id="C183E00CAF04FFF2FF584E984C3445A6" bold="true" box="[199,353,941,968]" pageId="2" pageNumber="433">Distribution.</emphasis>
<taxonomicName id="34F7479DAF04FFF2FEF54E9A4F0545A6" authority="Michelsen &amp; Grootaert, 2019" authorityName="Michelsen &amp; Grootaert" authorityYear="2019" box="[362,592,942,968]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="abrollensis" status="sp. nov.">
<emphasis id="C183E00CAF04FFF2FEF54E9A4F0545A6" box="[362,592,942,968]" italics="true" pageId="2" pageNumber="433">Drapetis abrollensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAB05D77AF04FFF2FDC64E9B4FE045A6" box="[601,693,942,968]" pageId="2" pageNumber="433" rank="species">
<emphasis id="C183E00CAF04FFF2FDC64E9B4FE045A6" bold="true" box="[601,693,942,968]" pageId="2" pageNumber="433">sp. nov.</emphasis>
</taxonomicNameLabel>
is known so far only by the male
<typeStatus id="2C4C82BCAF04FFF2FBDB4E9B49F345A6" box="[1092,1190,942,968]" pageId="2" pageNumber="433" type="holotype">holotype</typeStatus>
from the province of
<collectingRegion id="3133F2FCAF04FFF2FF084EE44D8E4582" box="[151,219,977,1004]" country="Sweden" name="Skane" pageId="2" pageNumber="433">Skåne</collectingRegion>
in South
<collectingCountry id="8BE07C8EAF04FFF2FED74EE74CF14582" box="[328,420,978,1004]" name="Sweden" pageId="2" pageNumber="433">Sweden</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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