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156 lines
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<document id="E7D71D4FA060972A455E4900648813F8" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA030AFF88FF2FFE7DFDA113AE" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus dacatraianus Deuve 1996" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="37" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9E28578674FF9E430D2C633B84C3A91B">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="6B494C3EB2BFCEDE36E5F8689FDEA013">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="6EC262413067456AA4BEC1BDDCF5E3FE">text</mods:typeOfResource>
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<mods:title id="F6887501806C6E149F6CB3CA5F5F7B21">Zootaxa</mods:title>
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<mods:date id="420573456A6CCBF3883B60E67AE29EC1">2009</mods:date>
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<mods:number id="809007FF18F6389E572798747340FA9A">2009-08-06</mods:number>
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<mods:number id="4C7BAF559460D201FB4F47E8F817D188">2178</mods:number>
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<mods:number id="096EC2EB7F97FD960E794E9383F81BF6">1</mods:number>
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<mods:url id="A5C30A00A06F4D83587A430F64814752">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="CCA7405D072137BD272B9EEB95B68177">journal article</mods:classification>
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<mods:identifier id="4C8DF502ABF43C05363AFFA591ED55CC" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="4EFE405C281D6B64BF5B29B1207BA302" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="783405605E19832CF5D131C44E237CCF" type="Zenodo-Dep">5312276</mods:identifier>
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<treatment id="733A87FA030AFF88FF2FFE7DFDA113AE" ID-DOI="http://doi.org/10.5281/zenodo.5320034" ID-Zenodo-Dep="5320034" LSID="urn:lsid:plazi:treatment:733A87FA030AFF88FF2FFE7DFDA113AE" httpUri="http://treatment.plazi.org/id/733A87FA030AFF88FF2FFE7DFDA113AE" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection id="B3896567030AFF88FF2FFE7DFDE414DB" box="[151,552,461,487]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="FB2C36EC030AFF88FF2FFE7DFDE414DB" blockId="36.[151,552,461,487]" box="[151,552,461,487]" pageId="36" pageNumber="37">
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<heading id="A0648180030AFF88FF2FFE7DFDE414DB" bold="true" box="[151,552,461,487]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
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<emphasis id="C9E7EAFE030AFF88FF2FFE7DFDE414DB" bold="true" box="[151,552,461,487]" pageId="36" pageNumber="37">
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The
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<taxonomicName id="3C934D6F030AFF88FF77FE7DFE1A14DB" ID-CoL="57ZJ3" authorityName="Deuve" authorityYear="1996" box="[207,470,461,487]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dacatraianus">
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<emphasis id="C9E7EAFE030AFF88FF77FE7DFE1A14DB" bold="true" box="[207,470,461,487]" italics="true" pageId="36" pageNumber="37">Trechus dacatraianus</emphasis>
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</taxonomicName>
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group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567030AFF88FF2FFDA6FECB1033" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="FB2C36EC030AFF88FF2FFDA6FECB1033" blockId="36.[151,1436,534,1682]" pageId="36" pageNumber="37">
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<emphasis id="C9E7EAFE030AFF88FF2FFDA6FED7170C" bold="true" box="[151,283,534,560]" pageId="36" pageNumber="37">Diagnosis:</emphasis>
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Head with frontal furrows deep, flattened and indistinct at level of hind suborbital seta in some species, +/- strongly curved at middle. Frons and supraorbital areas strongly convex. Temples distinctly pubescent. Mandibles normal. Pronotum subcordate, with hind angles well produced. Pronotal base rectilinear or the outer fifth slightly curved anteriorly. Basal transverse depression of pronotum diffuse, limited towards disc; laterobasal foveae broadly developed. Pronotal median line distinct, but not deepened before base. Hind wings reduced to small stubs. Humerus broadly rounded. Each elytron with parascutellar seta, preapical seta and two discal setae on third interval, with anterior discal seta located on stria III at the end of the anterior elytral quarter, and with middle dorsal seta located on stria III somewhat behind elytral middle. Stria VIII slightly (
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<emphasis id="C9E7EAFE030AFF88FEB8FCFCFD541659" box="[256,664,844,869]" italics="true" pageId="36" pageNumber="37">
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<taxonomicName id="3C934D6F030AFF88FEB8FCFCFE761659" authorityName="Deuve" authorityYear="1996" box="[256,442,844,869]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dacatraianus">T. dacatraianus</taxonomicName>
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,
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<taxonomicName id="3C934D6F030AFF88FE7EFCFCFD541659" authorityName="Deuve" authorityYear="1997" box="[454,664,844,869]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="damchungensis">T. damchungensis</taxonomicName>
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</emphasis>
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) or faintly impressed behind the level of the fifth umbilicate pore and deeply impressed at levels of seventh and eighth umbilicate pores. Recurrent elytral preapical sulcus deeply impressed and directed to the end of the fifth stria. Ventral surface smooth. Legs average, with moderately slender (
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<emphasis id="C9E7EAFE030AFF88FE2BFC70FD1116E5" box="[403,733,960,985]" italics="true" pageId="36" pageNumber="37">
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<taxonomicName id="3C934D6F030AFF88FE2BFC70FDE416E5" authorityName="Deuve" authorityYear="1997" box="[403,552,960,985]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hodeberti">T. hodeberti</taxonomicName>
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,
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<taxonomicName id="3C934D6F030AFF88FD8EFC70FD1116E5" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[566,733,960,985]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mieheorum" status="sp. nov.">T. mieheorum</taxonomicName>
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</emphasis>
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<emphasis id="C9E7EAFE030AFF88FD5EFC0FFCE716E5" bold="true" box="[742,811,959,985]" pageId="36" pageNumber="37">
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<taxonomicNameLabel id="D2D45785030AFF88FD5EFC0FFCE716E5" box="[742,811,959,985]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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) or thick femora and relatively thin tibia and tarsi; protibiae distinctly dilated towards apices, hardly bowed, each without a longitudinal groove on external surface. Two basal protarsi of male dilated, dentoid at the inner apical border. Aedeagal median lobe average to large, with basal bulb spherically enlarged and strongly bent downwards, with basal velum small or completely reduced, and with terminal lamella strongly hooked at tip. Internal sac with sclerotized portion quadripartite, extending almost half of the length of median lobe or even more; folding lobes in dorsal view bilaterally symmetrical, whereas the more basal pair of lobes forms a sheath for the more distal pair. Parameres average to relatively stout, broad at tip, both with four setae; left paramere slightly longer than right one.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567030AFF88FF7EFAABFDA113AE" pageId="36" pageNumber="37" type="discussion">
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<paragraph id="FB2C36EC030AFF88FF7EFAABFDF01322" blockId="36.[151,1436,534,1682]" pageId="36" pageNumber="37">
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<emphasis id="C9E7EAFE030AFF88FF7EFAABFE8D1009" bold="true" box="[198,321,1307,1333]" pageId="36" pageNumber="37">Remarks:</emphasis>
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Although this group includes species with relatively different external morphology, it is very likely a natural group because all the species share a more complicated structure of the internal sac of the aedeagus which is unique within the
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<taxonomicName id="3C934D6F030AFF88FDD4FAD9FD0110BE" authorityName="Clairville" authorityYear="1806" box="[620,717,1385,1410]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="C9E7EAFE030AFF88FDD4FAD9FD0110BE" box="[620,717,1385,1410]" italics="true" pageId="36" pageNumber="37">Trechus</emphasis>
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</taxonomicName>
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fauna of
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<collectingRegion id="3957F80E030AFF88FCF0FAD9FC4410BF" box="[840,904,1385,1411]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
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and adjacent mountainous regions, and is certainly a synapomorphic feature. However, the taxonomic position of the
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<taxonomicName id="3C934D6F030AFF88FBAFFA20FB1C1095" authorityName="Deuve" authorityYear="1996" box="[1047,1232,1424,1449]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dacatraianus">
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<emphasis id="C9E7EAFE030AFF88FBAFFA20FB1C1095" box="[1047,1232,1424,1449]" italics="true" pageId="36" pageNumber="37">T. dacatraianus</emphasis>
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</taxonomicName>
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group within the genus is quite difficult to trace without taking into account a much more comprehensive data set of characters. Therefore, a more detailed phylogenetic study which includes the highly diverse
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<taxonomicName id="3C934D6F030AFF88FBF0FA6DFB6810CA" authorityName="Clairville" authorityYear="1806" box="[1096,1188,1501,1526]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="C9E7EAFE030AFF88FBF0FA6DFB6810CA" box="[1096,1188,1501,1526]" italics="true" pageId="36" pageNumber="37">Trechus</emphasis>
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</taxonomicName>
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fauna of the Western Chinese mountain ranges is needed.
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</paragraph>
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<paragraph id="FB2C36EC030AFF88FF7EF99AFDA113AE" blockId="36.[151,1436,534,1682]" pageId="36" pageNumber="37">
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<emphasis id="C9E7EAFE030AFF88FF7EF99AFE531378" bold="true" box="[198,415,1578,1604]" pageId="36" pageNumber="37">Species included:</emphasis>
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<taxonomicName id="3C934D6F030AFF88FE11F99BFDEB1378" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[425,551,1579,1604]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bastropi" status="sp. nov.">
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<emphasis id="C9E7EAFE030AFF88FE11F99BFDEB1378" box="[425,551,1579,1604]" italics="true" pageId="36" pageNumber="37">T. bastropi</emphasis>
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</taxonomicName>
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<emphasis id="C9E7EAFE030AFF88FD89F99AFDB81378" bold="true" box="[561,628,1578,1604]" pageId="36" pageNumber="37">
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<taxonomicNameLabel id="D2D45785030AFF88FD89F99AFDB81378" box="[561,628,1578,1604]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(South Central
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<collectingRegion id="3957F80E030AFF88FC8BF99BFCB91379" box="[819,885,1579,1605]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
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),
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<materialsCitation id="4BFB3CB1030AFF88FC33F99BFF2F1357" location="North Eastern" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Xizang">
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<taxonomicName id="3C934D6F030AFF88FC33F99BFB2F1379" authority="Deuve, 1996" authorityName="Deuve" authorityYear="1996" box="[907,1251,1579,1605]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dacatraianus">
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<emphasis id="C9E7EAFE030AFF88FC33F99BFB8F1378" box="[907,1091,1579,1604]" italics="true" pageId="36" pageNumber="37">T. dacatraianus</emphasis>
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<bibRefCitation id="9F024B1D030AFF88FBF4F99BFB2F1379" author="Deuve, T." box="[1100,1251,1579,1605]" pageId="36" pageNumber="37" pagination="65 - 70" refId="ref48181" refString="Deuve, T. (1996) Nouveaux Trechus du Tibet et des regions limitrophes (Coleoptera, Trechidae). Bulletin de la Societe entomologique de France, 101 (1), 65 - 70." type="journal article" year="1996">Deuve, 1996</bibRefCitation>
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</taxonomicName>
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(
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<location id="FE4C6037030AFF88FB4DF99BFA571379" LSID="urn:lsid:plazi:treatment:733A87FA030AFF88FF2FFE7DFDA113AE:FE4C6037030AFF88FB4DF99BFA571379" box="[1269,1435,1579,1605]" name="North Eastern" pageId="36" pageNumber="37" stateProvince="Xizang">North Eastern</location>
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<collectingRegion id="3957F80E030AFF88FF2FF9E1FF171357" box="[151,219,1617,1643]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
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)
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</materialsCitation>
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,
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<materialsCitation id="4BFB3CB1030AFF88FF57F9E1FCE61357" box="[239,810,1617,1643]" location="Eastern" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Xizang">
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<taxonomicName id="3C934D6F030AFF88FF57F9E1FDA61357" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[239,618,1617,1643]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="damchungensis">
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<emphasis id="C9E7EAFE030AFF88FF57F9E1FE0A1356" box="[239,454,1617,1642]" italics="true" pageId="36" pageNumber="37">T. damchungensis</emphasis>
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<bibRefCitation id="9F024B1D030AFF88FE77F9E1FDA61357" author="Deuve, T." box="[463,618,1617,1643]" pageId="36" pageNumber="37" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
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</taxonomicName>
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(
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<location id="FE4C6037030AFF88FDC2F9E1FD1A1357" LSID="urn:lsid:plazi:treatment:733A87FA030AFF88FF2FFE7DFDA113AE:FE4C6037030AFF88FDC2F9E1FD1A1357" box="[634,726,1617,1643]" name="Eastern" pageId="36" pageNumber="37" stateProvince="Xizang">Eastern</location>
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<collectingRegion id="3957F80E030AFF88FD66F9E1FCED1357" box="[734,801,1617,1643]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
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)
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</materialsCitation>
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,
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<materialsCitation id="4BFB3CB1030AFF88FC8FF9E1FAB51357" box="[823,1401,1617,1643]" location="South Central" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Xizang">
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<taxonomicName id="3C934D6F030AFF88FC8FF9E1FBA71357" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[823,1131,1617,1643]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hodeberti">
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<emphasis id="C9E7EAFE030AFF88FC8FF9E1FC051356" box="[823,969,1617,1642]" italics="true" pageId="36" pageNumber="37">T. hodeberti</emphasis>
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|
|
<bibRefCitation id="9F024B1D030AFF88FC69F9E1FBA71357" author="Deuve, T." box="[977,1131,1617,1643]" pageId="36" pageNumber="37" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
|
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|
|
</taxonomicName>
|
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|
|
(
|
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|
|
<location id="FE4C6037030AFF88FBC4F9E1FAE81357" LSID="urn:lsid:plazi:treatment:733A87FA030AFF88FF2FFE7DFDA113AE:FE4C6037030AFF88FBC4F9E1FAE81357" box="[1148,1316,1617,1643]" name="South Central" pageId="36" pageNumber="37" stateProvince="Xizang">South Central</location>
|
|||
|
|
<collectingRegion id="3957F80E030AFF88FA95F9E1FABC1357" box="[1325,1392,1617,1643]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
|
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|
|
)
|
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|
|
</materialsCitation>
|
|||
|
|
,
|
|||
|
|
<materialsCitation id="4BFB3CB1030AFF88FA3EF9E1FDA413AE" location="South Central" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Xizang">
|
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|
|
<taxonomicName id="3C934D6F030AFF88FA3EF9E1FED713AD" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mieheorum" status="sp. nov.">
|
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|
|
<emphasis id="C9E7EAFE030AFF88FA3EF9E1FED713AD" italics="true" pageId="36" pageNumber="37">T. mieheorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="C9E7EAFE030AFF88FE9AF9C7FEA813AD" bold="true" box="[290,356,1655,1681]" pageId="36" pageNumber="37">
|
|||
|
|
<taxonomicNameLabel id="D2D45785030AFF88FE9AF9C7FEA813AD" box="[290,356,1655,1681]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<location id="FE4C6037030AFF88FECBF9C8FDDB13AE" LSID="urn:lsid:plazi:treatment:733A87FA030AFF88FF2FFE7DFDA113AE:FE4C6037030AFF88FECBF9C8FDDB13AE" box="[371,535,1656,1682]" name="South Central" pageId="36" pageNumber="37" stateProvince="Xizang">South Central</location>
|
|||
|
|
<collectingRegion id="3957F80E030AFF88FDA6F9C8FDAC13AE" box="[542,608,1656,1682]" country="China" name="Xizang" pageId="36" pageNumber="37">Tibet</collectingRegion>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|