treatments-xml/data/53/08/87/530887A4E066FFD39DEA5F78FA5AFBC7.xml

<document id="EFF47F587923D953F6BE447795820EDA" ID-DOI="10.5281/zenodo.183769" ID-GBIF-Dataset="2c4f3956-2fad-4239-8494-fbf20984d1a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="183769" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459935462660" checkinUser="plazi" docAuthor="Kaiser, Stefanie" docDate="2008" docId="530887A4E066FFD39DEA5F78FA5AFBC7" docLanguage="en" docName="zt01866p481.pdf" docOrigin="Zootaxa 1866" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hebefustis clareolithis Kaiser, 2008, sp. nov." docType="treatment" docVersion="10" lastPageNumber="479" masterDocId="AF31FFDCE065FFDF9D7D5D37FFC7FFE5" masterDocTitle="First record and new species of the genus Hebefustis Siebenaller &amp; Hessler, 1977 (Isopoda: Asellota: Nannoniscidae) from the deep Weddell Sea Basin *" masterLastPageNumber="481" masterPageNumber="467" pageNumber="470" updateTime="1698230051120" updateUser="plazi">
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<mods:title id="87FE321CF80EC034F37F508C4674D5D3">First record and new species of the genus Hebefustis Siebenaller &amp; Hessler, 1977 (Isopoda: Asellota: Nannoniscidae) from the deep Weddell Sea Basin *</mods:title>
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<mods:name id="541F6DE373CA85166FD0E40D7F82CABD" type="personal">
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<mods:namePart id="BAE6C00B019C66325F445533A362095A">Kaiser, Stefanie</mods:namePart>
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<mods:date id="A3A8E76E754EA28805A5F2EA3D594079">2008</mods:date>
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<treatment id="530887A4E066FFD39DEA5F78FA5AFBC7" ID-DOI="http://doi.org/10.5281/zenodo.5670990" ID-GBIF-Taxon="119375102" ID-Zenodo-Dep="5670990" LSID="urn:lsid:plazi:treatment:530887A4E066FFD39DEA5F78FA5AFBC7" httpUri="http://treatment.plazi.org/id/530887A4E066FFD39DEA5F78FA5AFBC7" lastPageId="12" lastPageNumber="479" pageId="3" pageNumber="470">
<subSubSection id="93BB6539E066FFDC9DEA5F78FED6FD69" pageId="3" pageNumber="470" type="nomenclature">
<paragraph id="DB1E36B2E066FFDC9DEA5F78FDCCFD8C" blockId="3.[151,523,591,652]" box="[151,523,591,617]" pageId="3" pageNumber="470">
<heading id="805681DEE066FFDC9DEA5F78FDCCFD8C" bold="true" box="[151,523,591,617]" fontSize="11" level="1" pageId="3" pageNumber="470" reason="1">
<emphasis id="E9D5EAA0E066FFDC9DEA5F78FDCCFD8C" bold="true" box="[151,523,591,617]" pageId="3" pageNumber="470">
<taxonomicName id="1CA14D31E066FFDC9DEA5F78FE61FD8C" ID-CoL="3JXH2" box="[151,422,591,617]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="470" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E066FFDC9DEA5F78FE61FD8C" bold="true" box="[151,422,591,617]" italics="true" pageId="3" pageNumber="470">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE066FFDC9CD05F78FDCCFD8C" box="[429,523,591,617]" pageId="3" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DEA5F45FED6FD69" blockId="3.[151,523,591,652]" box="[151,273,626,652]" pageId="3" pageNumber="470">
(
<figureCitation id="439A2A37E066FFDC9DE25F45FEC0FD69" box="[159,263,626,652]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,255,1977,2001]" captionStartId-1="5.[151,255,1771,1795]" captionStartId-2="7.[151,255,1897,1921]" captionStartId-3="9.[151,255,1934,1958]" captionStartId-4="11.[151,255,1910,1934]" captionTargetBox-0="[281,1301,465,1964]" captionTargetBox-1="[227,1291,157,1759]" captionTargetBox-2="[181,1393,156,1896]" captionTargetBox-3="[187,1426,181,1933]" captionTargetBox-4="[235,1345,436,1895]" captionTargetId-0="figure@4.[268,1319,459,1969]" captionTargetId-1="figure@5.[209,1378,149,1765]" captionTargetId-2="figure@7.[151,1436,71,1961]" captionTargetId-3="figure@9.[151,1436,108,1970]" captionTargetId-4="figure@11.[222,1365,416,1905]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="11" captionText-0="FIGURE 1. Hebefustis clareolithis sp. nov.; A – C, holotype female (ZMH K – 41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm." captionText-1="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." captionText-2="FIGURE 3. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 2 – 4. D, P 1. Scale bar = 100 µm." captionText-3="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," captionText-4="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri-0="https://zenodo.org/record/183770/files/figure.png" httpUri-1="https://zenodo.org/record/183771/files/figure.png" httpUri-2="https://zenodo.org/record/183772/files/figure.png" httpUri-3="https://zenodo.org/record/183773/files/figure.png" httpUri-4="https://zenodo.org/record/183774/files/figure.png" pageId="3" pageNumber="470">Figs 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E066FFDC9DEA5FF5FBA5FC99" pageId="3" pageNumber="470" type="materials_examined">
<paragraph id="DB1E36B2E066FFDC9DEA5FF5FEB2FD3E" blockId="3.[151,1436,706,2012]" box="[151,373,706,731]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DEA5FF5FEB2FD3E" box="[151,373,706,731]" italics="true" pageId="3" pageNumber="470">Material examined</emphasis>
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DBB5FDDFB90FCC9" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5FDDFEFFFCE1" bold="true" box="[198,312,746,772]" pageId="3" pageNumber="470">
<typeStatus id="041A8810E066FFDC9DBB5FDDFEFFFCE1" box="[198,312,746,772]" pageId="3" pageNumber="470" type="holotype">Holotype</typeStatus>
</emphasis>
: female, (preparatory, 
<quantity id="1C599B57E066FFDC9F295FDDFD70FCE1" box="[596,695,746,772]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="470" unit="mm" value="1.9">1.9 mm</quantity>
), eastern Weddell Sea, 
<geoCoordinate id="BE955075E066FFDC9EA05FDDFB98FCE1" box="[989,1119,746,772]" direction="south" orientation="latitude" pageId="3" pageNumber="470" precision="9" value="-70.518">70°31.08'S</geoCoordinate>
– 
<geoCoordinate id="BE955075E066FFDC99125FDDFB37FCE1" box="[1135,1264,746,772]" direction="south" orientation="latitude" pageId="3" pageNumber="470" precision="9" value="-70.53717">70°32.23'S</geoCoordinate>
, 
<geoCoordinate id="BE955075E066FFDC987C5FDDFA4BFCE1" box="[1281,1420,746,772]" direction="west" orientation="longitude" pageId="3" pageNumber="470" precision="9" value="-14.580334">14°34.82'W</geoCoordinate>
– 
<geoCoordinate id="BE955075E066FFDC9DEA5E25FEE4FCC9" box="[151,291,786,812]" direction="west" orientation="longitude" pageId="3" pageNumber="470" precision="9" value="-14.581667">14°34.90'W</geoCoordinate>
, PS ANT XXII/3, Station 81–8, 4382 m, 
<date id="AF1F1072E066FFDC9E7A5E25FC49FCC9" box="[775,910,786,812]" pageId="3" pageNumber="470" value="2005-02-24">24.02.2005</date>
, 
<collectionCode id="BDB0AE77E066FFDC9EE75E25FC1CFCC9" box="[922,987,786,812]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="470">ZMH</collectionCode>
K–41261.
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DBB5E0DFBA5FC99" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5E0DFE84FCB1" bold="true" box="[198,323,826,852]" pageId="3" pageNumber="470">
<typeStatus id="041A8810E066FFDC9DBB5E0DFE84FCB1" box="[198,323,826,852]" pageId="3" pageNumber="470" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="CDA7FD3BE066FFDC9C2E5E0DFE02FCB1" box="[339,453,826,852]" pageId="3" pageNumber="470" type="female">9 females</specimenCount>
(7 preparatory, 
<specimenCount id="CDA7FD3BE066FFDC9FFD5E0DFD3EFCB1" box="[640,761,826,852]" pageId="3" pageNumber="470" type="juvenile">1 juvenile</specimenCount>
, 1 damaged), 
<specimenCount id="CDA7FD3BE066FFDC9EE05E0DFC2BFCB1" box="[925,1004,826,852]" pageId="3" pageNumber="470" type="male">1 male</specimenCount>
(juvenile), same station as 
<typeStatus id="041A8810E066FFDC98535E0DFA50FCB1" box="[1326,1431,826,852]" pageId="3" pageNumber="470" type="holotype">holotype</typeStatus>
, 
<collectionCode id="BDB0AE77E066FFDC9DEA5E55FF1FFC99" box="[151,216,866,892]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="470">ZMH</collectionCode>
K–41262; 
<specimenCount id="CDA7FD3BE066FFDC9C215E55FE6CFC99" box="[348,427,866,892]" pageId="3" pageNumber="470" type="male">1 male</specimenCount>
(adult, 
<quantity id="1C599B57E066FFDC9F785E55FDA4FC99" box="[517,611,866,892]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="470" unit="mm" value="1.4">1.4 mm</quantity>
), same station as 
<typeStatus id="041A8810E066FFDC9E4D5E55FC5EFC99" box="[816,921,866,892]" pageId="3" pageNumber="470" type="holotype">holotype</typeStatus>
, 
<collectionCode id="BDB0AE77E066FFDC9ED85E55FC21FC99" box="[933,998,866,892]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="470">ZMH</collectionCode>
K–41263.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E066FFDC9DBB5EBDFF16FBF9" pageId="3" pageNumber="470" type="diagnosis">
<paragraph id="DB1E36B2E066FFDC9DBB5EBDFF16FBF9" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5EBDFE15FC46" box="[198,466,906,931]" italics="true" pageId="3" pageNumber="470">Diagnosis. Pereonite 2</emphasis>
with a robust spine on anterolateral margins. Pereonites 6 and 7 with acute, distally directed posterolateral margins. Second article of antennula with 4 broom setae distally. Antenna with 12 flagellar articles. Posterolateral spines well developed and acute, halfway along margin. Pleopod 1 widest distally.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E066FFD59DBB591DFAFEF917" lastPageId="10" lastPageNumber="477" pageId="3" pageNumber="470" type="description">
<paragraph id="DB1E36B2E066FFDC9DBB591DFBCDFA19" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB591DFE79FBA6" box="[198,446,1066,1091]" italics="true" pageId="3" pageNumber="470">Description. Habitus</emphasis>
of female 
<typeStatus id="041A8810E066FFDC9F3D591DFD60FBA1" box="[576,679,1066,1092]" pageId="3" pageNumber="470" type="holotype">holotype</typeStatus>
(
<figureCitation id="439A2A37E066FFDC9FCA591DFD39FBA1" box="[695,766,1066,1092]" captionStart="FIGURE 1" captionStartId="4.[151,255,1977,2001]" captionTargetBox="[281,1301,465,1964]" captionTargetId="figure@4.[268,1319,459,1969]" captionTargetPageId="4" captionText="FIGURE 1. Hebefustis clareolithis sp. nov.; A – C, holotype female (ZMH K – 41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm." httpUri="https://zenodo.org/record/183770/files/figure.png" pageId="3" pageNumber="470">Figs 1</figureCitation>
A–C): 
<emphasis id="E9D5EAA0E066FFDC9E28591DFC56FBA6" box="[853,913,1066,1091]" italics="true" pageId="3" pageNumber="470">Body</emphasis>
dorsoventrally slightly flattened and broadened, 3.7 times longer than wide. Surface of dorsal cuticle with numerous setules (
<figureCitation id="439A2A37E066FFDC99095965FB7BFB89" box="[1140,1212,1106,1132]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="3" pageNumber="470">Fig. 2</figureCitation>
H). Ventral surface alveolarly structured (
<figureCitation id="439A2A37E066FFDC9CE7594DFE1EFB71" box="[410,473,1146,1172]" captionStart="FIGURE 1" captionStartId="4.[151,255,1977,2001]" captionTargetBox="[281,1301,465,1964]" captionTargetId="figure@4.[268,1319,459,1969]" captionTargetPageId="4" captionText="FIGURE 1. Hebefustis clareolithis sp. nov.; A – C, holotype female (ZMH K – 41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm." httpUri="https://zenodo.org/record/183770/files/figure.png" pageId="3" pageNumber="470">Fig.1</figureCitation>
B). 
<emphasis id="E9D5EAA0E066FFDC9F7E594CFD89FB71" box="[515,590,1147,1172]" italics="true" pageId="3" pageNumber="470">Coxae</emphasis>
not visible in dorsal view. 
<emphasis id="E9D5EAA0E066FFDC9EF6594CFBFDFB71" box="[907,1082,1147,1172]" italics="true" pageId="3" pageNumber="470">Pereonites 1–3</emphasis>
with frontally directed laterofrontal margins. 
<emphasis id="E9D5EAA0E066FFDC9C215994FE22FB59" box="[348,485,1187,1212]" italics="true" pageId="3" pageNumber="470">Pereonite 1</emphasis>
and 2 nearly of same length and width, p 
<emphasis id="E9D5EAA0E066FFDC9EA45994FB88FB59" box="[985,1103,1187,1212]" italics="true" pageId="3" pageNumber="470">ereonite 2</emphasis>
widest, 4 times longer than wide, with a robust spine anterolaterally. Pereonite 4 longest, nearly twice as wide as long. Pereonites 1–4 decreasing, pereonites 5–7 increasing in width distally. Body gradually flattening from pereonites 1–4 and slightly increasing in height from pereonites 5–7. Pereonites 6 and 7 medially fused, with acute, distally directed posterolateral margins and 2 ventral depressions. 
<emphasis id="E9D5EAA0E066FFDC9E3C5874FC7AFAB9" box="[833,957,1347,1372]" italics="true" pageId="3" pageNumber="470">Pleotelson</emphasis>
0.23 times body length, as long as wide, width 0.9 times pereonite 2 width, with well-developed posterolateral spines, halfway along margin. Posterior margin strongly convex, anterior dorsal margin straight. 
<emphasis id="E9D5EAA0E066FFDC9E5158A4FCA0FA49" box="[812,871,1427,1452]" italics="true" pageId="3" pageNumber="470">Anus</emphasis>
(
<figureCitation id="439A2A37E066FFDC9E0B58A5FC7FFA49" box="[886,952,1426,1452]" captionStart="FIGURE 1" captionStartId="4.[151,255,1977,2001]" captionTargetBox="[281,1301,465,1964]" captionTargetId="figure@4.[268,1319,459,1969]" captionTargetPageId="4" captionText="FIGURE 1. Hebefustis clareolithis sp. nov.; A – C, holotype female (ZMH K – 41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm." httpUri="https://zenodo.org/record/183770/files/figure.png" pageId="3" pageNumber="470">Fig. 1</figureCitation>
B) covered by anus valves laterally. 
<emphasis id="E9D5EAA0E066FFDC981958A4FF08FA31" italics="true" pageId="3" pageNumber="470">Uropods</emphasis>
inserting closely to the anus valves, short in relation to pleotelson, hardly projecting above posterior margin. Operculum about 0.6 times pleotelson length (ventrally measured).
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DBB5B3CFECBF991" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5B3CFEB1F9C1" box="[198,374,1547,1572]" italics="true" pageId="3" pageNumber="470">Cephalothorax</emphasis>
(
<figureCitation id="439A2A37E066FFDC9CF75B3DFE08F9C1" box="[394,463,1546,1572]" captionStart="FIGURE 1" captionStartId="4.[151,255,1977,2001]" captionTargetBox="[281,1301,465,1964]" captionTargetId="figure@4.[268,1319,459,1969]" captionTargetPageId="4" captionText="FIGURE 1. Hebefustis clareolithis sp. nov.; A – C, holotype female (ZMH K – 41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm." httpUri="https://zenodo.org/record/183770/files/figure.png" pageId="3" pageNumber="470">Fig. 1</figureCitation>
A, B) as wide as long, 
<emphasis id="E9D5EAA0E066FFDC9F895B3CFC42F9C1" box="[756,901,1547,1572]" italics="true" pageId="3" pageNumber="470">rostral crest</emphasis>
slightly developed, anterior margin of head strongly, posterior margin slightly rounded, lateral margins straight. 
<emphasis id="E9D5EAA0E066FFDC9EAC5B04FB87F9A9" box="[977,1088,1587,1612]" italics="true" pageId="3" pageNumber="470">Antennae</emphasis>
inserting frontolaterally in a deep fold.
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DBB5BB4FCCAF881" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5BB4FEF8F979" box="[198,319,1667,1692]" italics="true" pageId="3" pageNumber="470">Antennula</emphasis>
(
<figureCitation id="439A2A37E066FFDC9C325BB5FE52F979" box="[335,405,1666,1692]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="3" pageNumber="470">Fig. 2</figureCitation>
B) of female 
<typeStatus id="041A8810E066FFDC9F4C5BB5FD51F979" box="[561,662,1666,1692]" pageId="3" pageNumber="470" type="paratype">paratype</typeStatus>
0.16 times body length, with 5 articles. Basal article short, nearly as long as wide, with 2 broom setae of varying size and 2 slender seta distally. Article 2 slender, 2.1 times longer than wide, with 2 long and 2 shorter broom setae distally, with 1 simple seta distally. Articles 3 and 4 short, almost of same length and width, each about 0.3 times article 2 length. Article 4 with 2 simple setae distally (1 broken off). Article 5 long and slender, nearly 3 times longer than article 4, 3.9 times longer than wide, with 2 long, slender setae and 1 aesthetasc terminally.
</paragraph>
<paragraph id="DB1E36B2E066FFDC9DBB5A44FA5BF839" blockId="3.[151,1436,706,2012]" pageId="3" pageNumber="470">
<emphasis id="E9D5EAA0E066FFDC9DBB5A44FEEFF869" box="[198,296,1907,1932]" italics="true" pageId="3" pageNumber="470">Antenna</emphasis>
(
<figureCitation id="439A2A37E066FFDC9C445A45FE46F869" box="[313,385,1906,1932]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="3" pageNumber="470">Fig. 2</figureCitation>
A) length 0.6 times body length, with 6 peduncular and 12 flagellar articles. Peduncular articles 1–4 short; article 3 with 1 robust spine tipped with a small seta distally, with 2 slender setae of same size and 1 small spine distally and a fringe of short setules laterally. Article 4 with a single small seta distally.
</paragraph>
<paragraph id="DB1E36B2E061FFDB9DEA5DAFFC68FE47" blockId="4.[151,1437,152,418]" pageId="4" pageNumber="471">Article 5 nearly as long as articles 1–4 combined, about 3.4 times longer than wide, with 4 simple setae laterally, with 1 small broom seta and 4 simple setae distally. Article 6 length 1.3 times article 5 length, about 4.6 times longer than wide; with 5 short simple setae laterally, with 4 broom setae and 5 slender setae distally. Flagellar article 1 longest, about 0.5 times peduncular article 6 length, 3.6 times longer than wide, with 3 simple setae laterally and 3 simple setae distally. Flagellar articles 2–10 of similar length; width decreasing distally, each article with 1–4 short simple setae distally. Articles 11–12 somewhat shorter; article 11 with 3 simple setae distally, article 12 with 6 long slender setae terminally.</paragraph>
<caption id="8FDE663AE061FFDB9DEA5A8EFE39F816" httpUri="https://zenodo.org/record/183770/files/figure.png" pageId="4" pageNumber="471" targetBox="[281,1301,465,1964]" targetPageId="4">
<paragraph id="DB1E36B2E061FFDB9DEA5A8EFE39F816" blockId="4.[151,1436,1977,2035]" pageId="4" pageNumber="471">
<emphasis id="E9D5EAA0E061FFDB9DEA5A8EFEDEF834" bold="true" box="[151,281,1977,2001]" pageId="4" pageNumber="471">FIGURE 1.</emphasis>
<taxonomicName id="1CA14D31E061FFDB9C5D5A8DFDD7F834" box="[288,528,1978,2001]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="471" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E061FFDB9C5D5A8DFDD7F834" box="[288,528,1978,2001]" italics="true" pageId="4" pageNumber="471">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE061FFDB9F6A5A8DFDAAF834" box="[535,621,1978,2001]" pageId="4" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
; A–C, holotype female (ZMH K–41261); A, dorsal view. B, lateral view. C, Plt ventrally. Scale bar = 500 µm.
</paragraph>
</caption>
<caption id="8FDE663AE060FFDA9DEA5BDCFF05F8AD" httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="5" pageNumber="472" targetBox="[227,1291,157,1759]" targetPageId="5">
<paragraph id="DB1E36B2E060FFDA9DEA5BDCFF05F8AD" blockId="5.[151,1436,1771,1864]" pageId="5" pageNumber="472">
<emphasis id="E9D5EAA0E060FFDA9DEA5BDCFEDDF8E6" bold="true" box="[151,282,1771,1795]" pageId="5" pageNumber="472">FIGURE 2.</emphasis>
<taxonomicName id="1CA14D31E060FFDA9C5C5BDBFDD6F8E6" box="[289,529,1772,1795]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="472" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E060FFDA9C5C5BDBFDD6F8E6" box="[289,529,1772,1795]" italics="true" pageId="5" pageNumber="472">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE060FFDA9F645BDBFDA8F8E6" box="[537,623,1772,1795]" pageId="5" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
; A–G paratype female (ZMH K–41262); A, A2. B, A1. C, rMd. D, lMd. E, Mxp. F, Mx1. G, Mx2. H, holotype female (ZMH K–41261); habitus, dorsal view. Scale bar: A–G = 100 µm, H = 500 µm.
</paragraph>
</caption>
<paragraph id="DB1E36B2E060FFD99DBB5A93FC57FE2F" blockId="5.[151,1436,1955,2021]" lastBlockId="6.[151,1437,152,1978]" lastPageId="6" lastPageNumber="473" pageId="5" pageNumber="472">
<emphasis id="E9D5EAA0E060FFDA9DBB5A93FE55F858" box="[198,402,1956,1981]" italics="true" pageId="5" pageNumber="472">Mandibular palp</emphasis>
of left and right 
<emphasis id="E9D5EAA0E060FFDA9F115A93FD1EF858" box="[620,729,1956,1981]" italics="true" pageId="5" pageNumber="472">mandible</emphasis>
(
<figureCitation id="439A2A37E060FFDA9F935A94FCFFF858" box="[750,824,1955,1981]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="5" pageNumber="472">Fig. 2</figureCitation>
C, D) well developed, consisting of 3 articles and almost reaching incisor. Palpal article 1 of right mandible 1.6 times longer than article 2. Terminal article tapering distally, with a fringe of small setules ventrally and 2 robust setae terminally. Article 1 and 2 of left mandible of similar length, article 2 with fringe of fine setules and 2 simple setae distally; article 3 tapering in width distally, with a fringe of fine setules and 2 longer simple setae ventrally. Incisor of right mandible with 3 teeth, of left mandible with 6 teeth. Lacinia mobilis of left mandible with 4 teeth (
<figureCitation id="439A2A37E063FFD999ED5C27FB1DFECF" box="[1168,1242,272,298]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="6" pageNumber="473">Fig. 2</figureCitation>
D). Setal row of right mandible with 9 robust setae and several slender setae in between; distal robust setae dentate, dentation decreasing proximally. Setal row of left mandible with 8 setae; distal setae 1–3 most robust, proximal setae more slender. Molar of right and left mandible triangular, on right mandible with 15 long, slender setae distally, on left mandible with 11 long, slender serrate setae distally.
</paragraph>
<paragraph id="DB1E36B2E063FFD99DBB5CEFFE75FDA7" blockId="6.[151,1437,152,1978]" pageId="6" pageNumber="473">
<emphasis id="E9D5EAA0E063FFD99DBB5CEFFE9BFE14" box="[198,348,472,497]" italics="true" pageId="6" pageNumber="473">Outer endite</emphasis>
of 
<emphasis id="E9D5EAA0E063FFD99CFB5CEFFE33FE14" box="[390,500,472,497]" italics="true" pageId="6" pageNumber="473">maxillula</emphasis>
(
<figureCitation id="439A2A37E063FFD99F7B5CEFFD89FE17" box="[518,590,472,498]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="6" pageNumber="473">Fig. 2</figureCitation>
F) with 11 robust spine-like setae distally. Outer margin with 7 slender setae, inner margin with several (=15) slender setae. Inner endite 0.6 times outer endite length, with numerous (=20) fine setae distally.
</paragraph>
<paragraph id="DB1E36B2E063FFD99DBB5F67FDD2FD5F" blockId="6.[151,1437,152,1978]" pageId="6" pageNumber="473">
<emphasis id="E9D5EAA0E063FFD99DBB5F67FEE6FD8C" box="[198,289,592,617]" italics="true" pageId="6" pageNumber="473">Maxilla</emphasis>
(
<figureCitation id="439A2A37E063FFD99C4E5F67FEBCFD8F" box="[307,379,592,618]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="6" pageNumber="473">Fig. 2</figureCitation>
G) outer margins of lateral and inner endites with several setae. 3 strong setae on lateral and medial endites. Distal margin of inner endite with numerous long setae of varying size. Surface of inner endite with several short setules.
</paragraph>
<paragraph id="DB1E36B2E063FFD99DBB5FFFFCD2FC1F" blockId="6.[151,1437,152,1978]" pageId="6" pageNumber="473">
Left and right 
<emphasis id="E9D5EAA0E063FFD99C095FFFFE37FD04" box="[372,496,712,737]" italics="true" pageId="6" pageNumber="473">maxilliped</emphasis>
(
<figureCitation id="439A2A37E063FFD99F7F5FFFFD8DFD07" box="[514,586,712,738]" captionStart="FIGURE 2" captionStartId="5.[151,255,1771,1795]" captionTargetBox="[227,1291,157,1759]" captionTargetId="figure@5.[209,1378,149,1765]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis clareolithis sp. nov.; A – G paratype female (ZMH K – 41262); A, A 2. B, A 1. C, rMd. D, lMd. E, Mxp. F, Mx 1. G, Mx 2. H, holotype female (ZMH K – 41261); habitus, dorsal view. Scale bar: A – G = 100 µm, H = 500 µm." httpUri="https://zenodo.org/record/183771/files/figure.png" pageId="6" pageNumber="473">Fig. 2</figureCitation>
E) connected by 3 retinacula. Epipodite smooth, triangular, slender, 3.5 times longer than wide, reaching to mid of palpal article 3. Palpal article 1 short, about 2.5 times wider than long. Article 2 longest, 3 times longer than article 1, almost quadrangular. Palpal article 3 length 2.3 times article 1 length, 1.4 times wider than long, with 3 robust sensory setae and 2 more slender setae distally. Article 4 about 1.4 times article 1 length, 1.6 times longer than wide, with a projection reaching to mid of article 5, with 3 slender setae distally. Article 5 about 0.7 times article 4 length, twice as long as wide, with 3 long, slen- der setae terminally. Endite distal margin with some robust, dentate setae and several fine setae laterally. Protopod nearly triangular, about 3 times wider than long.
</paragraph>
<paragraph id="DB1E36B2E063FFD99DBB593FFE3CFA3F" blockId="6.[151,1437,152,1978]" pageId="6" pageNumber="473">
<emphasis id="E9D5EAA0E063FFD99DBB593FFE8AFBC4" box="[198,333,1032,1057]" italics="true" pageId="6" pageNumber="473">Pereopod 1</emphasis>
(
<figureCitation id="439A2A37E063FFD99C21593FFE58FBC7" box="[348,415,1032,1058]" captionStart="FIGURE 3" captionStartId="7.[151,255,1897,1921]" captionTargetBox="[181,1393,156,1896]" captionTargetId="figure@7.[151,1436,71,1961]" captionTargetPageId="7" captionText="FIGURE 3. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 2 – 4. D, P 1. Scale bar = 100 µm." httpUri="https://zenodo.org/record/183772/files/figure.png" pageId="6" pageNumber="473">Fig. 3</figureCitation>
D) more robust than pereopods 2–7. Basis 3.6 times longer than wide, with 5 short simple setae dorsally and 4 simple setae ventrally. Ischium length 0.7 times basis length, 2.3 times longer than wide; with 2 simple setae dorsally (one broken off) and 2 simple setae ventrally. Merus length 0.6 times ischium length, 1.3 times longer than wide; with 2 long robust setae distally and a smaller seta on a small projection distodorsally, with 4 setae ventrally. Carpus length 1.5 times merus length, 2.4 times longer than wide; with 2 slender setae dorsally, ventral margin slightly concave, with numerous small setae, membranously embedded, and 3 robust unequally bifid setae in between ventrally and 1 very long simple seta distally. Propodus length 0.9 times carpus length, 3 times longer than wide; with 3 slender simple setae dorsally, with numerous small setae, membranously embedded, 1 robust unequally bifid seta in between ventrally and 2 more slender setae distally. Dactylus nearly half of propodus length, 2.8 times longer than wide; with 3 slender setae medially and with 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and with 2 slender, longer setae ventrally.
</paragraph>
<paragraph id="DB1E36B2E063FFD99DBB58DFFDEEF85F" blockId="6.[151,1437,152,1978]" pageId="6" pageNumber="473">
Basis of 
<emphasis id="E9D5EAA0E063FFD99C4D58DFFE73F9E4" box="[304,436,1512,1537]" italics="true" pageId="6" pageNumber="473">pereopod 2</emphasis>
(
<figureCitation id="439A2A37E063FFD99CBA58DFFDCBF9E7" box="[455,524,1512,1538]" captionStart="FIGURE 3" captionStartId="7.[151,255,1897,1921]" captionTargetBox="[181,1393,156,1896]" captionTargetId="figure@7.[151,1436,71,1961]" captionTargetPageId="7" captionText="FIGURE 3. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 2 – 4. D, P 1. Scale bar = 100 µm." httpUri="https://zenodo.org/record/183772/files/figure.png" pageId="6" pageNumber="473">Fig. 3</figureCitation>
A) proximally damaged during dissection, about 4.1 times longer than wide; with 3 simple setae (one broken off) dorsally, with 4 long setae ventrally and 2 setae distally. Ischium about half of basis length, 2.5 times longer than wide; with 1 simple seta distodorsally and 3 simple setae ventrally. Merus length 0.6 times ischium length, 1.7 times longer than wide, with 2 robust simple setae distodorsally and with 3 more slender setae ventrally. Carpus length 2.6 times merus length, 4.6 times longer than wide; with 1 slender seta dorsally and 2 simple setae distally, with numerous small setae, membranously embedded, and with 4 stout unequally bifid setae in between ventrally. Propodus length 0.8 times carpus length, 6.6 times longer than wide, with 4 simple setae and 1 small broom seta dorsally, with numerous small setae, membranously embedded, and with 3 stout unequally bifid setae and 1 slender seta ventrally. Dactylus length nearly half of propodus length, 4.3 times longer than wide; with 3 thin, simple setae medially, with numerous small setae, membranously embedded, and 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae ventrally.
</paragraph>
<caption id="8FDE663AE062FFD89DEA5A5EFA55F864" box="[151,1426,1897,1921]" httpUri="https://zenodo.org/record/183772/files/figure.png" pageId="7" pageNumber="474" targetBox="[181,1393,156,1896]" targetPageId="7">
<paragraph id="DB1E36B2E062FFD89DEA5A5EFA55F864" blockId="7.[151,1426,1897,1921]" box="[151,1426,1897,1921]" pageId="7" pageNumber="474">
<emphasis id="E9D5EAA0E062FFD89DEA5A5EFEDEF864" bold="true" box="[151,281,1897,1921]" pageId="7" pageNumber="474">FIGURE 3.</emphasis>
<taxonomicName id="1CA14D31E062FFD89C5D5A5DFDC8F864" box="[288,527,1898,1921]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="474" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E062FFD89C5D5A5DFDC8F864" box="[288,527,1898,1921]" italics="true" pageId="7" pageNumber="474">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE062FFD89F6B5A5DFDACF864" box="[534,619,1898,1921]" pageId="7" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female (ZMH K–41262); A–C, P2–4. D, P1. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph id="DB1E36B2E06DFFD79DBB5DAFFD97FDA7" blockId="8.[151,1437,152,1938]" pageId="8" pageNumber="475">
Basis of 
<emphasis id="E9D5EAA0E06DFFD79C515DAFFE76FF54" box="[300,433,152,177]" italics="true" pageId="8" pageNumber="475">pereopod 3</emphasis>
(
<figureCitation id="439A2A37E06DFFD79CBD5DAFFDC2FF57" box="[448,517,152,178]" captionStart="FIGURE 3" captionStartId="7.[151,255,1897,1921]" captionTargetBox="[181,1393,156,1896]" captionTargetId="figure@7.[151,1436,71,1961]" captionTargetPageId="7" captionText="FIGURE 3. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 2 – 4. D, P 1. Scale bar = 100 µm." httpUri="https://zenodo.org/record/183772/files/figure.png" pageId="8" pageNumber="475">Fig. 3</figureCitation>
B) 4.3 times longer than wide; with 1 broom seta and 4 long slender setae dorsally, with 4 long simple setae (2 broken off) setae ventrally, with 2 long, slender setae distally. Ischium length about half of basis length, 2.3 times longer than wide; with 3 simple setae dorsally, and 3 simple setae ventrally. Merus length 0.6 times ischium length, 1.4 times longer than wide; with 2 long setae distodorsally and 2 simple setae of varying size ventrally. Carpus length 2.6 times merus length, 4.6 times longer than wide; with 4 slender setae dorsally, with numerous small setae, membranously embedded, and 4 stout unequally bifid setae in between ventrally, with a small, stout setae distally. Propodus length 0.9 times carpus length, 6.5 times longer than wide, with 3 simple setae dorsally, with numerous small setae, membranously embedded, and 3 stout unequally bifid setae ventrally, with 1 slender longer seta distally. Dactylus length half of propodus length, 4.5 times longer than wide; with 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae ventrally.
</paragraph>
<paragraph id="DB1E36B2E06DFFD79DBB5F67FE2DFBC7" blockId="8.[151,1437,152,1938]" pageId="8" pageNumber="475">
Basis of 
<emphasis id="E9D5EAA0E06DFFD79C525F67FE73FD8C" box="[303,436,592,617]" italics="true" pageId="8" pageNumber="475">pereopod 4</emphasis>
(
<figureCitation id="439A2A37E06DFFD79CB85F67FDCCFD8F" box="[453,523,592,618]" captionStart="FIGURE 3" captionStartId="7.[151,255,1897,1921]" captionTargetBox="[181,1393,156,1896]" captionTargetId="figure@7.[151,1436,71,1961]" captionTargetPageId="7" captionText="FIGURE 3. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 2 – 4. D, P 1. Scale bar = 100 µm." httpUri="https://zenodo.org/record/183772/files/figure.png" pageId="8" pageNumber="475">Fig. 3</figureCitation>
C) 4.1 times longer than wide; with 2 long broom setae and 3 smaller simple setae dorsally, with 4 simple setae ventrally and 2 robust setae distally. Ischium length 0.6 times basis length, 2.5 times longer than wide; with 3 simple setae and 1 more robust seta dorsally, with 4 simple setae ventrally. Merus length 0.6 times ischium length, 1.6 times longer than wide; with 2 long setae distodorsally and 2 long setae ventrally. Carpus length 2.6 times merus length, 5.9 times longer than wide, with 2 long simple setae dorsally, 1 simple seta medially and 1 broom seta and 1 small simple seta distally, with numerous small setae, membranously embedded, and 4 stout unequally bifid setae ventrally. Propodus length about carpus length, 7.5 times longer than wide; with 2 long simple setae and 1 broom seta dorsally, with numerous small setae, membranously embedded, and 3 stout unequally bifid setae and 1 slender, simple seta in between ventrally, with 1 long simple seta distally. Dactylus length 0.4 times propodus length, 3.4 times longer than wide; with 3 thin, simple setae medially and 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae ventrally.
</paragraph>
<paragraph id="DB1E36B2E06DFFD79DBB5907FD02F9E7" blockId="8.[151,1437,152,1938]" pageId="8" pageNumber="475">
Pereopods 5–7 more slender and longer than pereopods 1–4, due to comparably increased length of basis, carpus and propodus. Basis of 
<emphasis id="E9D5EAA0E06DFFD79F7C596FFD42FB94" box="[513,645,1112,1137]" italics="true" pageId="8" pageNumber="475">pereopod 5</emphasis>
(
<figureCitation id="439A2A37E06DFFD79FEB596FFD1BFB97" box="[662,732,1112,1138]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="8" pageNumber="475">Fig. 4</figureCitation>
A) 4.6 times longer than wide; with 2 long broom setae and 5 simple setae dorsally and with 4 long simple setae ventrally. Ischium length 0.7 times basis length, 3.6 times longer than wide; with 3 simple setae (1 broken off) dorsally and with 2 simple setae ventrally. Merus length 0.4 times ischium length, 1.6 times longer than wide; with 2 long robust setae distodorsally and with 3 slender setae of different size ventrally. Carpus length 2.3 times merus length, 5 times longer than wide; with 4 simple setae (3 broken off) dorsally and with 5 robust setae ventrally. Propodus length 1.1 times carpus length, 7.7 times longer than wide; with 1 small simple seta dorsally, with 2 long and 1 small simple setae and 1 long broom seta distally, with numerous small setae, membranously embedded, and 3 stout unequally bifid setae in between ventrally and 1 longer simple seta distally. Dactylus length half of propodus length, 6.1 times longer than wide; with 3 slender setae medially and with 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae ventrally.
</paragraph>
<paragraph id="DB1E36B2E06DFFD79DBB5B27FB61F877" blockId="8.[151,1437,152,1938]" pageId="8" pageNumber="475">
Basis of 
<emphasis id="E9D5EAA0E06DFFD79C535B27FE74F9CC" box="[302,435,1552,1577]" italics="true" pageId="8" pageNumber="475">pereopod 6</emphasis>
(
<figureCitation id="439A2A37E06DFFD79CB95B27FDCCF9CF" box="[452,523,1552,1578]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="8" pageNumber="475">Fig. 4</figureCitation>
B) 4.7 times longer than wide; with 5 slender, simple setae and 1 long broom seta dorsally, with 5 long, slender setae ventrally. Ischium length 0.7 times basis length, 3.7 times longer than wide; with 2 slender simple setae dorsally and 2 ventrally. Merus length 0.5 times ischium length, 1.8 times longer than wide; with 1 robust long setae distodorsally and 2 slender setae ventrally. Carpus length 2.5 times merus length, 7.6 times longer than wide; with 3 simple setae dorsally, and 5 simple setae ventrally. Propodus length about carpus length, 8.7 times longer than wide; with 2 long simple setae and 1 broom seta distodorsally, with numerous small setae, membranously embedded, 2 robust unequally bifid setae and 1 slender simple seta ventrally. Dactylus length half of propodus length, 5.4 times longer than wide; with 3 slender setae medially and with numerous small setae, membranously embedded, and 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae of similar size ventrally.
</paragraph>
<caption id="8FDE663AE06CFFD69DEA5AB9FA5BF843" box="[151,1436,1934,1958]" httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="9" pageNumber="476" targetBox="[187,1426,181,1933]" targetPageId="9">
<paragraph id="DB1E36B2E06CFFD69DEA5AB9FA5BF843" blockId="9.[151,1436,1934,1958]" box="[151,1436,1934,1958]" pageId="9" pageNumber="476">
<emphasis id="E9D5EAA0E06CFFD69DEA5AB9FEDEF843" bold="true" box="[151,281,1934,1958]" pageId="9" pageNumber="476">FIGURE 4.</emphasis>
<taxonomicName id="1CA14D31E06CFFD69C5C5AB8FDD7F843" box="[289,528,1935,1958]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="476" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E06CFFD69C5C5AB8FDD7F843" box="[289,528,1935,1958]" italics="true" pageId="9" pageNumber="476">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE06CFFD69F655AB8FDA9F843" box="[536,622,1935,1958]" pageId="9" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female (ZMH K–41262); A–C, P5–7. D, Plp 2 (operculum). E–G,
</paragraph>
</caption>
<paragraph id="DB1E36B2E06FFFD59DBB5DAFFCF7FDFF" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
Basis of 
<emphasis id="E9D5EAA0E06FFFD59C515DAFFE77FF54" box="[300,432,152,177]" italics="true" pageId="10" pageNumber="477">pereopod 7</emphasis>
(
<figureCitation id="439A2A37E06FFFD59CC25DAFFDC2FF57" box="[447,517,152,178]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
C) 5 times longer than wide; with 3 simple setae dorsally and 5 simple setae (1 broken off) ventrally. Ischium length 0.7 times basis length, 3.7 times longer than wide; with 2 slender simple setae dorsally and 2 ventrally. Merus length about half of ischium length, 1.8 times longer than wide; with 1 robust long and 1 smaller, slender seta distodorsally and with 2 slender simple setae ventrally. Carpus length 2.4 times merus length, 4.7 times longer than wide; with 4 simple setae dorsally, and 4 more robust simple setae ventrally. Propodus length 1.1 times carpus length, 7.3 times longer than wide; with 2 long slender setae and 1 broom seta distodorsally, with 2 robust unequally bifid setae and 2 simple setae ventrally. Dactylus length half of propodus length, 5.7 times longer than wide; with 4 slender setae medially and with numerous small setae, membranously embedded, and 1 spine-like appendix ventrally. Unguis with 1 robust cuspidate seta dorsally and 2 slender setae (1 broken off) ventrally.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5F1FFCE1FD8F" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5F1FFE86FDA4" box="[198,321,552,577]" italics="true" pageId="10" pageNumber="477">Pleopod 2</emphasis>
(operculum, 
<figureCitation id="439A2A37E06FFFD59C985F1FFDEAFDA7" box="[485,557,552,578]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
D) almost as long as wide; ventral surface smooth. Lateral and posterior margins with several (&gt;20) medium-sized slender setae.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5F4FFEBDFCEF" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
Protopodite of 
<emphasis id="E9D5EAA0E06FFFD59C0B5F4FFE2CFD74" box="[374,491,632,657]" italics="true" pageId="10" pageNumber="477">pleopod 3</emphasis>
(
<figureCitation id="439A2A37E06FFFD59C865F4FFD85FD77" box="[507,578,632,658]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
E) almost as long as wide, length 0.6 times endopodite length. Exopodite about half of endopodite length, 1.3 times longer than wide, tapering in width distally; with numerous simple setae laterally and 1 very long, robust seta distally. Endopodite 1.5 times longer than wide, with 3 long plumose setae distally.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5E2FFCB3FC67" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
Protopodite of 
<emphasis id="E9D5EAA0E06FFFD59C015E2FFE33FCD4" box="[380,500,792,817]" italics="true" pageId="10" pageNumber="477">pleopod 4</emphasis>
(
<figureCitation id="439A2A37E06FFFD59F7B5E2FFD88FCD7" box="[518,591,792,818]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
F) rectangular, 1.5 times wider than long. Exopodite slender, about as long as endopodite, 6 times longer than wide; with several thin setules laterally and 1 long robust plumose seta distally. Endopodite ovoid-shaped, 1.7 times longer than wide.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5EA7FB42FC4F" blockId="10.[151,1436,152,2018]" box="[198,1157,912,938]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5EA7FEF9FC4C" box="[198,318,912,937]" italics="true" pageId="10" pageNumber="477">Pleopod 5</emphasis>
(
<figureCitation id="439A2A37E06FFFD59C335EA7FE53FC4F" box="[334,404,912,938]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
G) small oval lobe, without setation. 1.9 times longer than wide.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5E8FFC4BFBAF" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5E8FFEEBFC34" box="[198,300,952,977]" italics="true" pageId="10" pageNumber="477">Uropods</emphasis>
(
<figureCitation id="439A2A37E06FFFD59C415E8FFE44FC37" box="[316,387,952,978]" captionStart="FIGURE 4" captionStartId="9.[151,255,1934,1958]" captionTargetBox="[187,1426,181,1933]" captionTargetId="figure@9.[151,1436,108,1970]" captionTargetPageId="9" captionText="FIGURE 4. Hebefustis clareolithis sp. nov., paratype female (ZMH K – 41262); A – C, P 5 – 7. D, Plp 2 (operculum). E – G," httpUri="https://zenodo.org/record/183773/files/figure.png" pageId="10" pageNumber="477">Fig. 4</figureCitation>
H) biramous. Protopodite trapezoid, about as long as wide, as long as endopodite; with 2 robust setae (broken off). Exopodite 0.8 times endopodite length. 2.9 times longer than wide, with 2 setae (broken off) terminally. Endopodite twice as long as wide; with 1 robust simple seta laterally, with 4 broom setae (1 broken off) and 2 simple setae (1 broken off) terminally.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB596FFCFFFB27" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB596FFDDCFB94" box="[198,539,1112,1137]" italics="true" pageId="10" pageNumber="477">
Differences in 
<typeStatus id="041A8810E06FFFD59C0F596FFE1DFB94" box="[370,474,1112,1137]" pageId="10" pageNumber="477" type="paratype">paratype</typeStatus>
male
</emphasis>
(
<figureCitation id="439A2A37E06FFFD59F56596FFDB4FB97" box="[555,627,1112,1138]" captionStart="FIGURE 5" captionStartId="11.[151,255,1910,1934]" captionTargetBox="[235,1345,436,1895]" captionTargetId="figure@11.[222,1365,416,1905]" captionTargetPageId="11" captionText="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="10" pageNumber="477">Fig. 5</figureCitation>
). 
<emphasis id="E9D5EAA0E06FFFD59FF7596FFD20FB94" box="[650,743,1112,1137]" italics="true" pageId="10" pageNumber="477">Habitus</emphasis>
(
<figureCitation id="439A2A37E06FFFD59F8A596FFCFBFB97" box="[759,828,1112,1138]" captionStart="FIGURE 5" captionStartId="11.[151,255,1910,1934]" captionTargetBox="[235,1345,436,1895]" captionTargetId="figure@11.[222,1365,416,1905]" captionTargetPageId="11" captionText="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="10" pageNumber="477">Fig. 5</figureCitation>
A) very similar to 
<typeStatus id="041A8810E06FFFD5996B596FFB47FB97" box="[1046,1152,1112,1138]" pageId="10" pageNumber="477" type="holotype">holotype</typeStatus>
, dorsal cuticle also with numerous setules, but lateral margins of pereonites 1–3 more frontally directed, anterolateral spines on pereonite 2 more robust and pereonite 4 shorter than in female.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB59E7FCDFFB0F" blockId="10.[151,1436,152,2018]" box="[198,792,1232,1258]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB59E7FEA0FB0C" box="[198,359,1232,1257]" italics="true" pageId="10" pageNumber="477">Rostral crests</emphasis>
well developed compared to female.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB59CFFD28F9CF" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB59CFFEEFFAF4" box="[198,296,1272,1297]" italics="true" pageId="10" pageNumber="477">Antenna</emphasis>
(
<figureCitation id="439A2A37E06FFFD59C4559CFFEBBFAF7" box="[312,380,1272,1298]" captionStart="FIGURE 5" captionStartId="11.[151,255,1910,1934]" captionTargetBox="[235,1345,436,1895]" captionTargetId="figure@11.[222,1365,416,1905]" captionTargetPageId="11" captionText="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="10" pageNumber="477">Fig. 5</figureCitation>
E) with 6 peduncular and 9 flagellar articles. Articles 5–7 stouter than in female. Article 5 about 2.7 times longer than wide, with 4 simple setae and two broom setae laterally, with 1 broom seta and 1 simple seta medially. Article 6 length 1.2 times article 5 length, about 3.2 times longer than wide, with 4 simple setae laterally, with 2 broom setae and 2 small simple setae distally. Flagellar article 1 longest, length 1.1 times peduncular article 6 length, 3.2 times longer than wide, with 4 simple setae laterally and 4 long simple setae (1 broken off) distally. Flagellar articles 2–7 of similar length, width decreasing distally, each article with 1–2 short simple setae distally. Articles 8 and 9 somewhat shorter, article 8 with 2 short simple setae distally, article 9 with 6 long, slender setae terminally.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5B0FFEE8F99F" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5B0FFEF9F9B4" box="[198,318,1592,1617]" italics="true" pageId="10" pageNumber="477">Pleopod 1</emphasis>
(
<figureCitation id="439A2A37E06FFFD59C305B0FFE55F9B7" box="[333,402,1592,1618]" captionStart="FIGURE 5" captionStartId="11.[151,255,1910,1934]" captionTargetBox="[235,1345,436,1895]" captionTargetId="figure@11.[222,1365,416,1905]" captionTargetPageId="11" captionText="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="10" pageNumber="477">Fig. 5</figureCitation>
B, C) 1.7 times longer than distal width; tips slightly rounded in ventral view, with a few simple setae.
</paragraph>
<paragraph id="DB1E36B2E06FFFD59DBB5BBFFAFEF917" blockId="10.[151,1436,152,2018]" pageId="10" pageNumber="477">
Sympod of 
<emphasis id="E9D5EAA0E06FFFD59C2C5BBFFE01F944" box="[337,454,1672,1697]" italics="true" pageId="10" pageNumber="477">pleopod 2</emphasis>
(
<figureCitation id="439A2A37E06FFFD59CAB5BBFFDDBF947" box="[470,540,1672,1698]" captionStart="FIGURE 5" captionStartId="11.[151,255,1910,1934]" captionTargetBox="[235,1345,436,1895]" captionTargetId="figure@11.[222,1365,416,1905]" captionTargetPageId="11" captionText="FIGURE 5. Hebefustis clareolithis sp. nov., paratype male (ZMH K – 41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A 2. Scale bar: A = 200 µm, B – E = 100 µm." httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="10" pageNumber="477">Fig. 5</figureCitation>
D) 2.4 times longer than wide; outer margin rounded; with 5 slender simple setae laterally, inner margin slightly concave. Endopod inserting 0.6 from distal tip of sympod; stylet about half of sympod length, slightly curved, distal end reaching tip of sympod. Exopod short and rounded.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E06FFFD59DBB5A37FD6FF8FF" box="[198,680,1792,1818]" pageId="10" pageNumber="477" type="materials_examined">
<paragraph id="DB1E36B2E06FFFD59DBB5A37FD6FF8FF" blockId="10.[151,1436,152,2018]" box="[198,680,1792,1818]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5A37FEA5F8FC" box="[198,354,1792,1817]" italics="true" pageId="10" pageNumber="477">
<typeStatus id="041A8810E06FFFD59DBB5A37FF3AF8FC" box="[198,253,1792,1817]" pageId="10" pageNumber="477">Type</typeStatus>
locality.
</emphasis>
Eastern Weddell Sea basin.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E06FFFD59DBB5A1FFB8FF8A7" box="[198,1096,1832,1858]" pageId="10" pageNumber="477" type="distribution">
<paragraph id="DB1E36B2E06FFFD59DBB5A1FFB8FF8A7" blockId="10.[151,1436,152,2018]" box="[198,1096,1832,1858]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5A1FFE9AF8A4" box="[198,349,1832,1857]" italics="true" pageId="10" pageNumber="477">Distribution.</emphasis>
The species is only known from the eastern Weddell Sea basin.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E06FFFD59DBB5A67FBD0F88F" box="[198,1047,1872,1898]" pageId="10" pageNumber="477" type="etymology">
<paragraph id="DB1E36B2E06FFFD59DBB5A67FBD0F88F" blockId="10.[151,1436,152,2018]" box="[198,1047,1872,1898]" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5A67FE1BF88C" box="[198,476,1872,1897]" italics="true" pageId="10" pageNumber="477">
Etymology. 
<taxonomicName id="1CA14D31E06FFFD59C2C5A67FE1BF88C" box="[337,476,1872,1897]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">Clareolithis</taxonomicName>
</emphasis>
(masculine) is dedicated to an invaluable friend.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E06FFFD49DBB5A4FFC5CFEB7" lastPageId="11" lastPageNumber="478" pageId="10" pageNumber="477" type="discussion">
<paragraph id="DB1E36B2E06FFFD49DBB5A4FFDFAFF3F" blockId="10.[151,1436,152,2018]" lastBlockId="11.[151,1436,152,338]" lastPageId="11" lastPageNumber="478" pageId="10" pageNumber="477">
<emphasis id="E9D5EAA0E06FFFD59DBB5A4FFD85F874" box="[198,578,1912,1937]" italics="true" pageId="10" pageNumber="477">
Remarks. 
<taxonomicName id="1CA14D31E06FFFD59C465A4FFD85F874" box="[315,578,1912,1937]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="477" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">Hebefustis clareolithis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="F2E657DBE06FFFD59F355A4FFD65F877" box="[584,674,1912,1938]" pageId="10" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
belongs to a cluster of species in the genus 
<taxonomicName id="1CA14D31E06FFFD599E15A4FFADFF874" box="[1180,1304,1912,1937]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="477" phylum="Arthropoda" rank="genus">
<emphasis id="E9D5EAA0E06FFFD599E15A4FFADFF874" box="[1180,1304,1912,1937]" italics="true" pageId="10" pageNumber="477">Hebefustis</emphasis>
</taxonomicName>
possessing acute, distally directed posterolateral margins of the sixth and seventh pereonite (see 
<bibRefCitation id="BF304B43E06FFFD599EB5A97FF10F807" author="Siebenaller" pageId="10" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler 1977</bibRefCitation>
:30). This cluster comprises, besides the new species, 
<taxonomicName id="1CA14D31E06FFFD59E295AFFFA50F807" authority="Siebenaller &amp; Hessler, 1977" authorityName="Siebenaller &amp; Hessler" authorityYear="1977" box="[852,1431,1992,2018]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="477" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="E9D5EAA0E06FFFD59E295AFFFBF8F804" box="[852,1087,1992,2017]" italics="true" pageId="10" pageNumber="477">Hebefustis cornutus</emphasis>
<bibRefCitation id="BF304B43E06FFFD5993A5AFFFA50F807" author="Siebenaller" box="[1095,1431,1992,2018]" pageId="10" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler, 1977</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="1CA14D31E06EFFD49DEA5DAFFDA3FF57" authority="Siebenaller &amp; Hessler, 1977" authorityName="Siebenaller &amp; Hessler" authorityYear="1977" box="[151,612,152,178]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="E9D5EAA0E06EFFD49DEA5DAFFECFFF54" box="[151,264,152,177]" italics="true" pageId="11" pageNumber="478">H. dispar</emphasis>
<bibRefCitation id="BF304B43E06EFFD49C6F5DAFFDA3FF57" author="Siebenaller" box="[274,612,152,178]" pageId="11" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler, 1977</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="1CA14D31E06EFFD49F0E5DAFFB4CFF57" authority="Siebenaller &amp; Hessler, 1977" authorityName="Siebenaller &amp; Hessler" authorityYear="1977" box="[627,1163,152,178]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="hexadentium">
<emphasis id="E9D5EAA0E06EFFD49F0E5DAFFCE8FF54" box="[627,815,152,177]" italics="true" pageId="11" pageNumber="478">H. hexadentium</emphasis>
<bibRefCitation id="BF304B43E06EFFD49E455DAFFB4CFF57" author="Siebenaller" box="[824,1163,152,178]" pageId="11" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler, 1977</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="1CA14D31E06EFFD499E45DAFFF19FF3F" authority="Menzies, 1962" authorityName="Menzies" authorityYear="1962" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="E9D5EAA0E06EFFD499E45DAFFAD9FF54" box="[1177,1310,152,177]" italics="true" pageId="11" pageNumber="478">H. hirsutus</emphasis>
(
<bibRefCitation id="BF304B43E06EFFD4984D5DAFFF11FF3F" author="Menzies" pageId="11" pageNumber="480" refString="Menzies, R. J. (1962) The isopods of abyssal depths in the Atlantic Ocean. Vema Research Series, 1, 79 - 206." type="journal article" year="1962">Menzies, 1962</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="1CA14D31E06EFFD49C615DF7FDFEFF3F" authority="Mezhov, 1986" authorityName="Mezhov" authorityYear="1986" box="[284,569,192,218]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="vitjazi">
<emphasis id="E9D5EAA0E06EFFD49C615DF7FE40FF3C" box="[284,391,192,217]" italics="true" pageId="11" pageNumber="478">H. vitjazi</emphasis>
<bibRefCitation id="BF304B43E06EFFD49CF35DF7FDFEFF3F" author="Mezhov" box="[398,569,192,218]" pageId="11" pageNumber="480" refString="Mezhov, B. V. (1986) Bathyal and abyssal Nannoniscidae and Desmosomatidae (Isopoda, Asellota) from Alaska Bay. Archives of the Zoological Museum, Moscow State Unversity 24, 126 - 167. [In Russian]" type="journal article" year="1986">Mezhov, 1986</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="DB1E36B2E06EFFD49DBB5DDFFC5CFEB7" blockId="11.[151,1436,152,338]" pageId="11" pageNumber="478">
<taxonomicName id="1CA14D31E06EFFD49DBB5DDFFEA9FEE4" box="[198,366,232,257]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E06EFFD49DBB5DDFFEA9FEE4" box="[198,366,232,257]" italics="true" pageId="11" pageNumber="478">H. clareolithis</emphasis>
</taxonomicName>
differs from all other species of this cluster by the following characters: robust spine on the anterolateral margins of pereonite 
<quantity id="1C599B57E06EFFD49F535C27FD9BFECF" box="[558,604,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="478" unit="in" value="2.0">2 in</quantity>
male and female; female antenna bearing 12 flagellar articles; second article of antennula with 4 broom setae; pleopod 1 widest distally.
</paragraph>
</subSubSection>
<caption id="8FDE663AE06EFFD49DEA5A41FC56F855" httpUri="https://zenodo.org/record/183774/files/figure.png" pageId="11" pageNumber="478" targetBox="[235,1345,436,1895]" targetPageId="11">
<paragraph id="DB1E36B2E06EFFD49DEA5A41FC56F855" blockId="11.[151,1436,1910,1968]" pageId="11" pageNumber="478">
<emphasis id="E9D5EAA0E06EFFD49DEA5A41FEDAF86B" bold="true" box="[151,285,1910,1934]" pageId="11" pageNumber="478">FIGURE 5.</emphasis>
<taxonomicName id="1CA14D31E06EFFD49C585A41FDDFF868" box="[293,536,1910,1933]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="478" phylum="Arthropoda" rank="species" species="clareolithis" status="sp. nov.">
<emphasis id="E9D5EAA0E06EFFD49C585A41FDDFF868" box="[293,536,1910,1933]" italics="true" pageId="11" pageNumber="478">Hebefustis clareolithis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E657DBE06EFFD49F5F5A41FDBEF868" box="[546,633,1910,1933]" pageId="11" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
, paratype male (ZMH K–41263); A, habitus, dorsal view. B, Plt, ventral view. C, Plp 1. D, Plp 2. E, A2. Scale bar: A = 200 µm, B–E = 100 µm.
</paragraph>
</caption>
<subSubSection id="93BB6539E069FFD39DBB5DAFFB61FE17" pageId="12" pageNumber="479" type="reference_group">
<paragraph id="DB1E36B2E069FFD39DBB5DAFFCAAFEB7" blockId="12.[151,1437,152,1058]" pageId="12" pageNumber="479">
<taxonomicName id="1CA14D31E069FFD39DBB5DAFFEA9FF54" ID-CoL="3JXH2" box="[198,366,152,177]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E069FFD39DBB5DAFFEA9FF54" box="[198,366,152,177]" italics="true" pageId="12" pageNumber="479">H. clareolithis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="1CA14D31E069FFD39F3A5DAFFB43FF57" ID-CoL="3JXH3" authority="Siebenaller &amp; Hessler, 1977" authorityName="Siebenaller &amp; Hessler" authorityYear="1977" box="[583,1156,152,178]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="E9D5EAA0E069FFD39F3A5DAFFCF7FF54" box="[583,816,152,177]" italics="true" pageId="12" pageNumber="479">Hebefustis cornutus</emphasis>
<bibRefCitation id="BF304B43E069FFD39E455DAFFB43FF57" author="Siebenaller" box="[824,1156,152,178]" pageId="12" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler, 1977</bibRefCitation>
</taxonomicName>
; the latter can be distinguished from the new species by the following characters: female antenna with 9 flagellar articles; second article of antennula with just 2 broom setae distally; male antenna more robust: peduncular article 5 about 1.6 times longer than wide, article 6 about 2.3 times longer than wide, first flagellar article 1.8 times longer than wide; posterior margin of male pleotelson with double angles.
</paragraph>
<paragraph id="DB1E36B2E069FFD39DBB5C57FB61FE17" blockId="12.[151,1437,152,1058]" pageId="12" pageNumber="479">
The new species also resembles 
<taxonomicName id="1CA14D31E069FFD39F3B5C57FB93FE9F" ID-CoL="3JXH5" authority="Siebenaller &amp; Hessler, 1977" authorityName="Siebenaller &amp; Hessler" authorityYear="1977" box="[582,1108,352,378]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hexadentium">
<emphasis id="E9D5EAA0E069FFD39F3B5C57FCC7FE9C" box="[582,768,352,377]" italics="true" pageId="12" pageNumber="479">H. hexadentium</emphasis>
<bibRefCitation id="BF304B43E069FFD39E755C57FB93FE9F" author="Siebenaller" box="[776,1108,352,378]" pageId="12" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler, 1977</bibRefCitation>
</taxonomicName>
which can be distinguished from 
<taxonomicName id="1CA14D31E069FFD39DAA5CBFFEB9FE44" ID-CoL="3JXH2" box="[215,382,392,417]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E069FFD39DAA5CBFFEB9FE44" box="[215,382,392,417]" italics="true" pageId="12" pageNumber="479">H. clareolithis</emphasis>
</taxonomicName>
as follows: second article of antennula with 2 broom setae distally; female antenna with 7 flagellar articles, first flagellar article relatively short, about 0.25 times peduncular article 6 length; pereonites 2 and 3 long compared to pereonite 1; male posterior margin of pleotelson softly curved.
</paragraph>
</subSubSection>
<subSubSection id="93BB6539E069FFD39DBB5F37FA5AFBC7" pageId="12" pageNumber="479" type="description">
<paragraph id="DB1E36B2E069FFD39DBB5F37FEFFFCD7" blockId="12.[151,1437,152,1058]" pageId="12" pageNumber="479">
<bibRefCitation id="BF304B43E069FFD39DBB5F37FDDBFDFF" author="Siebenaller" box="[198,540,512,538]" pageId="12" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler (1977)</bibRefCitation>
pointed out remarkable similarities between 
<taxonomicName id="1CA14D31E069FFD399545F37FB24FDFC" box="[1065,1251,512,537]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hexadentium">
<emphasis id="E9D5EAA0E069FFD399545F37FB24FDFC" box="[1065,1251,512,537]" italics="true" pageId="12" pageNumber="479">H. hexadentium</emphasis>
</taxonomicName>
and 
<taxonomicName id="1CA14D31E069FFD398675F37FE9BFDA7" authority="Menzies, 1962" authorityName="Menzies" authorityYear="1962" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="E9D5EAA0E069FFD398675F37FA5AFDFC" box="[1306,1437,512,537]" italics="true" pageId="12" pageNumber="479">H. hirsutus</emphasis>
(
<bibRefCitation id="BF304B43E069FFD39DE25F1FFE94FDA7" author="Menzies" box="[159,339,552,578]" pageId="12" pageNumber="480" refString="Menzies, R. J. (1962) The isopods of abyssal depths in the Atlantic Ocean. Vema Research Series, 1, 79 - 206." type="journal article" year="1962">Menzies, 1962</bibRefCitation>
)
</taxonomicName>
, which were both first recorded from the South Atlantic (
<tableCitation id="96230309E069FFD3996F5F1FFBAAFDA7" box="[1042,1133,552,578]" captionStart="TABLE 1" captionStartId="1.[151,239,822,846]" captionTargetBox="[156,1424,947,2008]" captionTargetPageId="1" captionText="TABLE 1. Species composition and distribution of the genus Hebefustis Siebenaller &amp; Hessler, 1977." httpUri="http://table.plazi.org/id/8FDE663AE064FFDE9DEA5E01FB12FCAB" pageId="12" pageNumber="479" tableUuid="8FDE663AE064FFDE9DEA5E01FB12FCAB">Table 1</tableCitation>
). The male 
<typeStatus id="041A8810E069FFD3987F5F1FFAAEFDA7" box="[1282,1385,552,578]" pageId="12" pageNumber="479" type="holotype">holotype</typeStatus>
and only specimen of 
<taxonomicName id="1CA14D31E069FFD39C175F67FE29FD8C" box="[362,494,592,617]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="E9D5EAA0E069FFD39C175F67FE29FD8C" box="[362,494,592,617]" italics="true" pageId="12" pageNumber="479">H. hirsutus</emphasis>
</taxonomicName>
is no longer extant (
<bibRefCitation id="BF304B43E069FFD39F9C5F67FBEBFD8F" author="Siebenaller" box="[737,1068,592,618]" pageId="12" pageNumber="480" refString="Siebenaller, J. F. &amp; Hessler, R. R. (1977) The Nannoniscidae (Isopoda, Asellota): Hebefustis n. gen. and Nannoniscoides Hansen. Transactions of the San Diego Society of Natural History, 19, 17 - 44." type="journal article" year="1977">Siebenaller &amp; Hessler 1977</bibRefCitation>
), and the drawings by 
<bibRefCitation id="BF304B43E069FFD398445F67FF23FD77" author="Menzies" pageId="12" pageNumber="480" refString="Menzies, R. J. (1962) The isopods of abyssal depths in the Atlantic Ocean. Vema Research Series, 1, 79 - 206." type="journal article" year="1962">Menzies (1962)</bibRefCitation>
are rudimentary, i.e. appendices were just partly drawn. Therefore, the comparison of this species to both 
<taxonomicName id="1CA14D31E069FFD39DAA5F97FE44FD5C" box="[215,387,672,697]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E069FFD39DAA5F97FE44FD5C" box="[215,387,672,697]" italics="true" pageId="12" pageNumber="479">H. clareolithis</emphasis>
</taxonomicName>
and 
<taxonomicName id="1CA14D31E069FFD39CBB5F97FD43FD5C" box="[454,644,672,697]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hexadentium">
<emphasis id="E9D5EAA0E069FFD39CBB5F97FD43FD5C" box="[454,644,672,697]" italics="true" pageId="12" pageNumber="479">H. hexadentium</emphasis>
</taxonomicName>
remains difficult (cf. Key). However, 
<taxonomicName id="1CA14D31E069FFD3991F5F97FB2EFD5C" box="[1122,1257,672,697]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="E9D5EAA0E069FFD3991F5F97FB2EFD5C" box="[1122,1257,672,697]" italics="true" pageId="12" pageNumber="479">H. hirsutus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="1CA14D31E069FFD39C465FFFFE23FD04" box="[315,484,712,737]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E069FFD39C465FFFFE23FD04" box="[315,484,712,737]" italics="true" pageId="12" pageNumber="479">H. clareolithis</emphasis>
</taxonomicName>
by the following characters: pereonite 2 with a robust seta on anterolateral margins; maxilliped with 2 coupling hooks, incisor of right mandible with 4 teeth, posterior margin of pleotelson softly curved.
</paragraph>
<paragraph id="DB1E36B2E069FFD39DBB5E77FD54FC4F" blockId="12.[151,1437,152,1058]" pageId="12" pageNumber="479">
<taxonomicName id="1CA14D31E069FFD39DBB5E77FEF2FCBC" box="[198,309,832,857]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="E9D5EAA0E069FFD39DBB5E77FEF2FCBC" box="[198,309,832,857]" italics="true" pageId="12" pageNumber="479">H. dispar</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="1CA14D31E069FFD39F0F5E77FCDDFCBC" box="[626,794,832,857]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="E9D5EAA0E069FFD39F0F5E77FCDDFCBC" box="[626,794,832,857]" italics="true" pageId="12" pageNumber="479">H. clareolithis</emphasis>
</taxonomicName>
as follows: anterolateral margins of pereonites 6 and 7 more rounded; pleotelson 1.2 times wider than long, posterior margin softly curved, posterolateral spines 0.7 from anterior margin, spines less produced.
</paragraph>
<paragraph id="DB1E36B2E069FFD39DBB5E8FFA5AFBC7" blockId="12.[151,1437,152,1058]" pageId="12" pageNumber="479">
<taxonomicName id="1CA14D31E069FFD39DBB5E8FFEF4FC34" box="[198,307,952,977]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="479" phylum="Arthropoda" rank="species" species="vitjazi">
<emphasis id="E9D5EAA0E069FFD39DBB5E8FFEF4FC34" box="[198,307,952,977]" italics="true" pageId="12" pageNumber="479">H. vitjazi</emphasis>
</taxonomicName>
differs from the new species by the following characters: operculum relatively long, length 0.9 times pleotelson length (measured in ventral view), posterolateral spines about 0.8 from anterior margin; sympod of pleopod 1 with numerous setae (&gt;15), endopodal stylet overlapping distal end of sympod about a third.
</paragraph>
</subSubSection>
</treatment>
</document>