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<document id="CF467D5396BA12402B9E9143624814E6" ID-DOI="10.11646/zootaxa.3688.1.1" ID-GBIF-Dataset="151481a0-71e1-4402-ab7e-8262caddf539" ID-ISSN="1175-5326" ID-Zenodo-Dep="284201" ID-ZooBank="DB84C679-C2AA-41CC-816E-83E2F6DCA391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460649331738" checkinUser="plazi" docAuthor="Haddad, Charles R." docDate="2013" docId="03DB8445FFBEFF94FF2AFA9F305BF918" docLanguage="en" docName="zt03688p079.pdf" docOrigin="Zootaxa 3688 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Copuetta kwamgumi Haddad, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="39" masterDocId="FFE2FC3DFF9AFFB2FFBDFFE6345DFFF0" masterDocTitle="Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae" masterLastPageNumber="79" masterPageNumber="1" pageNumber="37" updateTime="1698337697679" updateUser="plazi">
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<mods:title id="1453482538A2CC35FB80C8CB03304331">Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae</mods:title>
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<mods:namePart id="1501F551EB5ED1E8BDA80F36A2216A41">Haddad, Charles R.</mods:namePart>
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<mods:title id="F45704452083C7571BF963B07DB32991">Zootaxa</mods:title>
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<mods:date id="335BB723E88CE23068B59704D66A87E8">2013</mods:date>
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<mods:number id="D577FF52FF1D97AE541574D9FB751B72">3688</mods:number>
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<mods:classification id="8B02D63539827AC9C8F1350C06135C50">journal article</mods:classification>
<mods:identifier id="00FF4603A474BD024C4D22942D56392B" type="DOI">10.11646/zootaxa.3688.1.1</mods:identifier>
<mods:identifier id="67CDED96DCFC782E36EF261B1E4552F5" type="GBIF-Dataset">151481a0-71e1-4402-ab7e-8262caddf539</mods:identifier>
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<treatment id="03DB8445FFBEFF94FF2AFA9F305BF918" ID-DOI="http://doi.org/10.5281/zenodo.6153937" ID-GBIF-Taxon="127664911" ID-Zenodo-Dep="6153937" LSID="urn:lsid:plazi:treatment:03DB8445FFBEFF94FF2AFA9F305BF918" httpUri="http://treatment.plazi.org/id/03DB8445FFBEFF94FF2AFA9F305BF918" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37">
<subSubSection id="C36866D8FFBEFF96FF2AFA9F35E1FA44" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BCD3553FFBEFF96FF2AFA9F35A8FA63" blockId="36.[151,501,1401,1460]" box="[151,501,1401,1427]" pageId="36" pageNumber="37">
<heading id="D085823FFFBEFF96FF2AFA9F35A8FA63" bold="true" box="[151,501,1401,1427]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="B906E941FFBEFF96FF2AFA9F35A8FA63" bold="true" box="[151,501,1401,1427]" pageId="36" pageNumber="37">
<taxonomicName id="4C724ED0FFBEFF96FF2AFA9F35CDFA63" ID-CoL="5ZXYF" box="[151,400,1401,1427]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FF2AFA9F35CDFA63" bold="true" box="[151,400,1401,1427]" italics="true" pageId="36" pageNumber="37">Copuetta kwamgumi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235543AFFBEFF96FE25FA9F35A8FA63" box="[408,501,1401,1427]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3553FFBEFF96FF2AFA7A35E1FA44" blockId="36.[151,501,1401,1460]" box="[151,444,1436,1460]" pageId="36" pageNumber="37">
<figureCitation id="134929D6FFBEFF96FF2AFA7A354BFA44" box="[151,278,1436,1460]" captionStart="FIGURES 45 – 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 – 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="36" pageNumber="37">Figs 46, 47</figureCitation>
,
<figureCitation id="134929D6FFBEFF96FE9FFA7A3513FA44" box="[290,334,1436,1460]" captionStart="FIGURES 99 – 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 – 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="36" pageNumber="37">101</figureCitation>
,
<figureCitation id="134929D6FFBEFF96FEE7FA7A35E1FA44" box="[346,444,1436,1460]" captionStart="FIGURES 120 – 123" captionStartId="38.[151,264,1359,1382]" captionTargetBox="[167,1407,404,1328]" captionTargetId="figure@38.[151,1436,375,1334]" captionTargetPageId="38" captionText="FIGURES 120 – 123. Genitalic morphology of Copuetta kwamgumi sp. nov.: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="36" pageNumber="37">120–123</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFBEFF96FF2AFA03378EFA0C" box="[151,979,1508,1533]" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BCD3553FFBEFF96FF2AFA03378EFA0C" blockId="36.[151,1436,1508,2036]" box="[151,979,1508,1533]" pageId="36" pageNumber="37">
<emphasis id="B906E941FFBEFF96FF2AFA03357CFA0D" bold="true" box="[151,289,1509,1533]" pageId="36" pageNumber="37">Etymology:</emphasis>
The species name is a noun in apposition of the
<typeStatus id="54C98BF1FFBEFF96FCFFFA03372FFA0D" box="[834,882,1509,1533]" pageId="36" pageNumber="37">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFBEFF96FF7AF9EE3070F925" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BCD3553FFBEFF96FF7AF9EE3070F925" blockId="36.[151,1436,1508,2036]" pageId="36" pageNumber="37">
<emphasis id="B906E941FFBEFF96FF7AF9EE351EF9D1" bold="true" box="[199,323,1544,1569]" pageId="36" pageNumber="37">Diagnosis:</emphasis>
This species is closely related to
<taxonomicName id="4C724ED0FFBEFF96FD78F9EF3770F9D0" box="[709,813,1545,1568]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="erecta" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FD78F9EF3770F9D0" box="[709,813,1545,1568]" italics="true" pageId="36" pageNumber="37">C. erecta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBEFF96FC88F9EF37CDF9D1" bold="true" box="[821,912,1545,1569]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A235543AFFBEFF96FC88F9EF37CDF9D1" box="[821,912,1545,1569]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Males can be recognised by the tiny embolus with tip directed distally, which is the smallest in the genus (
<figureCitation id="134929D6FFBEFF96FCF3F9CA37ECF9B4" box="[846,945,1580,1604]" captionStart="FIGURES 99 – 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 – 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="36" pageNumber="37">Fig. 101</figureCitation>
), while that of
<taxonomicName id="4C724ED0FFBEFF96FBDCF9CB3095F9B5" box="[1121,1224,1581,1605]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="erecta" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FBDCF9CB3095F9B5" box="[1121,1224,1581,1605]" italics="true" pageId="36" pageNumber="37">C. erecta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBEFF96FB6DF9CB3171F9B5" bold="true" box="[1232,1324,1581,1605]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A235543AFFBEFF96FB6DF9CB3171F9B5" box="[1232,1324,1581,1605]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is clearly longer and straight, with the tip directed distally (
<figureCitation id="134929D6FFBEFF96FD7EF9B63778F998" box="[707,805,1616,1640]" captionStart="FIGURES 99 – 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 – 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="36" pageNumber="37">Fig. 100</figureCitation>
). Females are characterised by the small lateral commashaped sclerotised rims that surround the round copulatory openings (
<figureCitation id="134929D6FFBEFF96FC1FF992305EF97C" box="[930,1027,1652,1676]" captionStart="FIGURES 120 – 123" captionStartId="38.[151,264,1359,1382]" captionTargetBox="[167,1407,404,1328]" captionTargetId="figure@38.[151,1436,375,1334]" captionTargetPageId="38" captionText="FIGURES 120 – 123. Genitalic morphology of Copuetta kwamgumi sp. nov.: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="36" pageNumber="37">Fig. 122</figureCitation>
), which differ from the small, coiled ridges of
<taxonomicName id="4C724ED0FFBEFF96FEBCF97F353AF940" box="[257,359,1689,1712]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="erecta" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FEBCF97F353AF940" box="[257,359,1689,1712]" italics="true" pageId="36" pageNumber="37">C. erecta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBEFF96FED3F97F3595F941" bold="true" box="[366,456,1689,1713]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A235543AFFBEFF96FED3F97F3595F941" box="[366,456,1689,1713]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="134929D6FFBEFF96FE65F97E3664F940" box="[472,569,1688,1712]" captionStart="FIGURES 114 – 117" captionStartId="33.[151,264,1205,1228]" captionTargetBox="[166,1421,208,1171]" captionTargetId="figure@33.[151,1436,195,1182]" captionTargetPageId="33" captionText="FIGURES 114 – 117. Genitalic morphology of Copuetta erecta sp. nov.: 114. Male palp, ventral view. 115. Same, retrolateral view. 116. Female epigyne, ventral view. 117. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284218/files/figure.png" pageId="36" pageNumber="37">Fig. 116</figureCitation>
). Furthermore,
<taxonomicName id="4C724ED0FFBEFF96FD5AF97F3710F940" box="[743,845,1689,1712]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="erecta" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FD5AF97F3710F940" box="[743,845,1689,1712]" italics="true" pageId="36" pageNumber="37">C. erecta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBEFF96FCE8F97F37F2F941" bold="true" box="[853,943,1689,1713]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A235543AFFBEFF96FCE8F97F37F2F941" box="[853,943,1689,1713]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
females have large semi-circular peripheral ridges in the posterior half of the epigyne that are absent in
<taxonomicName id="4C724ED0FFBEFF96FC8FF95B3791F925" box="[818,972,1725,1749]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">
<emphasis id="B906E941FFBEFF96FC8FF95B3791F925" box="[818,972,1725,1749]" italics="true" pageId="36" pageNumber="37">C. kwamgumi</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBEFF96FC6EF95B3070F925" bold="true" box="[979,1069,1725,1749]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A235543AFFBEFF96FC6EF95B3070F925" box="[979,1069,1725,1749]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFBEFF94FF7AF907365FFECF" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
<paragraph id="8BCD3553FFBEFF96FF7AF90730CAF8B0" blockId="36.[151,1436,1508,2036]" pageId="36" pageNumber="37">
<emphasis id="B906E941FFBEFF96FF7AF9073705F909" bold="true" box="[199,856,1760,1785]" pageId="36" pageNumber="37">
Male (
<typeStatus id="54C98BF1FFBEFF96FEABF90635DDF909" box="[278,384,1760,1785]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
,
<taxonomicName id="4C724ED0FFBEFF96FE2DF9063643F909" box="[400,542,1760,1785]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="76" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">Kwamgumi</taxonomicName>
, ZMUC). Measurements:
</emphasis>
CL 2.48, CW 2.00, AL 2.80, AW 1.70, TL 5.32 (5.00–5.90), FL 0.19, SL 1.11, SW 1.16, AME–AME 0.03, AME–ALE 0.01, ALE–ALE 0.44, PME–PME 0.11, PME–PLE 0.05, PLE–PLE 0.51,
<collectingCountry id="F36575C3FFBEFF96FDB0F8CF3604F8B0" box="[525,601,1833,1856]" name="Peru" pageId="36" pageNumber="37">PERW</collectingCountry>
0.79, MOQAW 0.44, MOQPW 0.44, MOQL 0.51.
</paragraph>
<paragraph id="8BCD3553FFBEFF96FF7AF8AA3036F878" blockId="36.[151,1436,1508,2036]" pageId="36" pageNumber="37">Length of leg segments: I 2.23 + 0.88 + 1.88 + 1.95 + 1.10 = 8.04; II 2.15 + 0.85 + 1.78 + 1.92 + 1.07 = 7.77; III 2.08 + 0.80 + 1.60 + 1.95 + 0.90 = 7.33; IV 2.58 + 0.85 + 2.15 + 2.84 + 1.00 = 9.42.</paragraph>
<paragraph id="8BCD3553FFBEFF97FF7AF872348CFC0C" blockId="36.[151,1436,1508,2036]" lastBlockId="37.[151,1437,151,2010]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
General appearance as in
<figureCitation id="134929D6FFBEFF96FDBCF8723607F85C" box="[513,602,1940,1964]" captionStart="FIGURES 45 – 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 – 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="36" pageNumber="37">Fig. 46</figureCitation>
. Carapace yellow-orange, eye region dark grey; paired mottled black mediolateral markings from palpal coxae to posterior slope, separated by broad pale median line from PER to posterior of carapace; small mottled black marking in front of fovea; orange-brown striae with faint black mottling radiating from fovea, directed between coxae; lateral margins with black fringe, expanded between leg coxae; clypeus mottled black between ALE; markings covered with black feathery setae, areas between them covered in white feathery and short straight setae. AME separated by distance equal to 2⁄5 their diameter; AME separated from ALE by distance less than 1⁄5 AME diameter; clypeus height slightly larger than 4⁄5 AME diameter; PLE larger than PME; PME separated by distance equal to ½ their diameter; PME separated from PLE by distance slightly less than ½ PME diameter; CW:
<collectingCountry id="F36575C3FFBFFF97FE25FEB135BAFE9F" box="[408,487,343,367]" name="Peru" pageId="37" pageNumber="38">PERW</collectingCountry>
= 2.53:1. Chelicerae yellow-brown with black mottling, yellow distally; promargin with two teeth separated by basal width of proximal tooth, distal tooth largest; retromargin with two subequal teeth separated by ½ their basal width, distal tooth close to fang base. Endites yellow, cream prolaterally distally; labium yellow-brown, cream distally; sternum bright yellow, darker around margins. Legs bright yellow, markings densely covered in black feathery setae, with yellowish feathery setae between them; femora I–IV with broad brown band in distal half; patellae without markings; tibiae and metatarsi I and II without markings, III and IV with faint brown broad median band; tarsi without markings. Leg spination: femora: I pl 3 do 3 rl 1, II pl 2 do 3 rl 2, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 2; all femora with scattered erect ventral setae; patellae: I and II with fine proximal and distal do setae, III and IV with fine proximal seta and distal spine dorsally; tibiae: I pl 2 plv 2-3 rlv 2, II pl 2 plv 2 rlv 2, III pl 2 do 1 rl 2 plv 2 rlv 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I plv 2 rlv 2, II pl 0-1 plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 3 rlv 1. Abdomen with yellow anterior dorsal scutum extending to 4⁄5 abdomen length; dorsum with oval marking anteriorly, laterally and posteriorly, comprising black, yellow-brown and white feathery setae, with small faint black spot anteriorly; dorsum pale along midline, covered in white feathery setae; spinnerets surrounded by narrow black ring dorsally and laterally, absent ventrally; venter cream, pale grey between ventral sclerite and spinnerets; epigastric scutum, ventral sclerite and inframamillary sclerite yellow. Male palp yellow, cymbium yellow-brown; tegulum orange, with orange-brown ducts; embolus tiny, with narrow stalk-like base and single coil, distal end slightly curved, directed at distal end of cymbium (
<figureCitation id="134929D6FFBFFF97FB47FC263102FC28" box="[1274,1375,960,984]" captionStart="FIGURES 99 – 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 – 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="37" pageNumber="38">Figs 101</figureCitation>
,
<figureCitation id="134929D6FFBFFF97FAD6FC263498FC0C" captionStart="FIGURES 120 – 123" captionStartId="38.[151,264,1359,1382]" captionTargetBox="[167,1407,404,1328]" captionTargetId="figure@38.[151,1436,375,1334]" captionTargetPageId="38" captionText="FIGURES 120 – 123. Genitalic morphology of Copuetta kwamgumi sp. nov.: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="37" pageNumber="38">120, 121</figureCitation>
).
</paragraph>
<paragraph id="8BCD3553FFBFFF97FF7AFBEF30CAFB98" blockId="37.[151,1437,151,2010]" pageId="37" pageNumber="38">
<emphasis id="B906E941FFBFFF97FF7AFBEF3733FBD1" bold="true" box="[199,878,1032,1057]" pageId="37" pageNumber="38">
Female (
<typeStatus id="54C98BF1FFBFFF97FE93FBEE35C1FBD1" box="[302,412,1032,1057]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
,
<taxonomicName id="4C724ED0FFBFFF97FE16FBEE3664FBD1" box="[427,569,1032,1057]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="76" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">Kwamgumi</taxonomicName>
, ZMUC). Measurements:
</emphasis>
CL 3.27, CW 2.48, AL 3.88, AW 2.59, TL 6.75 (5.70–7.25), FL 0.28, SL 1.39, SW 1.40, AME–AME 0.07, AME–ALE 0.01, ALE–ALE 0.52, PME–PME 0.13, PME–PLE 0.06, PLE–PLE 0.61,
<collectingCountry id="F36575C3FFBFFF97FDB0FBB73604FB98" box="[525,601,1105,1128]" name="Peru" pageId="37" pageNumber="38">PERW</collectingCountry>
0.94, MOQAW 0.55, MOQPW 0.54, MOQL 0.60.
</paragraph>
<paragraph id="8BCD3553FFBFFF97FF7AFB92302AFB40" blockId="37.[151,1437,151,2010]" pageId="37" pageNumber="38">Length of leg segments: I 2.78 + 1.13 + 2.35 + 2.45 + 1.40 = 10.11; II 2.62 + 1.10 + 2.20 + 2.40 + 1.33 = 9.65; III 2.53 + 1.05 + 1.98 + 2.40 + 1.08 = 9.04; IV 3.20 + 1.13 + 2.69 + 3.56 + 1.15 = 11.73.</paragraph>
<paragraph id="8BCD3553FFBFFF94FF7AFB5A365FFECF" blockId="37.[151,1437,151,2010]" lastBlockId="38.[151,1437,151,319]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
General appearance as in
<figureCitation id="134929D6FFBFFF97FDBCFB5A3604FB24" box="[513,601,1212,1236]" captionStart="FIGURES 45 – 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 – 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="37" pageNumber="38">Fig. 47</figureCitation>
, female more robustly built than male. Carapace deep yellow-brown, markings and setae as for male but darker and with more distinct black mottling. AME separated by distance slightly more than ¼ their diameter; AME separated from ALE by distance less than 1⁄10 AME diameter; clypeus height slightly more than 4⁄5 AME diameter; PLE very slightly larger than PME; PME separated by distance slightly less than ½ their diameter; PME separated from PLE by distance slightly less than ½ PME diameter; CW:
<collectingCountry id="F36575C3FFBFFF97FA88FAB131D8FA9F" box="[1333,1413,1367,1391]" name="Peru" pageId="37" pageNumber="38">PERW</collectingCountry>
= 2.64:1. Chelicerae brown with black mottling, yellow prolaterally distally; promargin with two teeth separated by ½ the basal width of proximal tooth, distal tooth largest; retromargin with two teeth separated by ½ the basal width of proximal tooth, distal tooth slightly smaller than proximal tooth, close to fang base. Endites yellow-orange, cream distally; labium yellow-brown, cream distally; sternum yellow-brown, orange-brown along margins. Legs densely covered in black feathery setae on markings, with yellowish feathery setae between them; femora I–IV yellow with orange-brown band in distal half, marking covered in faint black mottling; patellae I–IV yellow-brown with faint black mottling laterally and ventrally, with faint median band dorsally; tibiae I–IV yellow with broad median band, absent proximally and distally, faint on tibiae I and II, darker brown on III and IV; metatarsi I–IV yellow-orange, posteriors slightly darker; tarsi I–IV yellow-orange. Leg spination: femora: I pl 3 do 3 rl 1, II pl 2 do 3 rl 1, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 2; all femora with scattered erect ventral setae; patellae: I and II with fine proximal and distal do setae, III and IV with fine proximal seta and distal spine dorsally; tibiae: I pl 2 plv 2 rlv 2, II pl 2 plv 2 rlv 2, III pl 2 do 1 rl 2 plv 2 rlv 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 2 plv 1; tarsi: pl 1 plv 3 rlv 1. Abdomen with yellow-orange dorsal scutum extending to 4⁄5 abdomen length; dorsum mottled grey with cream chevrons along midline, sides with three white stripes at midpoint; mottling densely covered in yellow-brown and scattered white feathery setae, chevrons covered in white feathery and short straight setae; venter cream, pale mottled grey in front of spinnerets; spinnerets with narrow black ring dorsally and laterally, absent ventrally; epigastric scutum and inframamillary sclerite yellow. Epigyne with small lateral comma-shaped ridges at midpoint of epigyne with copulatory openings situated in middle of ridges, apparently entering directly into ST II posterolaterally (
<figureCitation id="134929D6FFBCFF94FC60FF39301CFF07" box="[989,1089,223,247]" captionStart="FIGURES 120 – 123" captionStartId="38.[151,264,1359,1382]" captionTargetBox="[167,1407,404,1328]" captionTargetId="figure@38.[151,1436,375,1334]" captionTargetPageId="38" captionText="FIGURES 120 – 123. Genitalic morphology of Copuetta kwamgumi sp. nov.: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="38" pageNumber="39">Fig. 122</figureCitation>
); ST II n-shaped with anterior bend, connected broadly to posterior kidney-shaped ST I; ST I slightly narrower than ST II; both ST with many folds on their surface (
<figureCitation id="134929D6FFBCFF94FE2EFEC135ABFECF" box="[403,502,295,319]" captionStart="FIGURES 120 – 123" captionStartId="38.[151,264,1359,1382]" captionTargetBox="[167,1407,404,1328]" captionTargetId="figure@38.[151,1436,375,1334]" captionTargetPageId="38" captionText="FIGURES 120 – 123. Genitalic morphology of Copuetta kwamgumi sp. nov.: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="38" pageNumber="39">Fig. 123</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0D65DBFFBCFF94FF2AFAA930D0FA74" httpUri="https://zenodo.org/record/284220/files/figure.png" pageId="38" pageNumber="39" targetBox="[167,1407,404,1328]" targetPageId="38">
<paragraph id="8BCD3553FFBCFF94FF2AFAA930D0FA74" blockId="38.[151,1436,1359,1412]" pageId="38" pageNumber="39">
<emphasis id="B906E941FFBCFF94FF2AFAA9352EFA94" bold="true" box="[151,371,1359,1382]" pageId="38" pageNumber="39">FIGURES 120–123.</emphasis>
Genitalic morphology of
<taxonomicName id="4C724ED0FFBCFF94FD32FAB6373BFA95" box="[655,870,1359,1381]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">
<emphasis id="B906E941FFBCFF94FD32FAB6373BFA95" box="[655,870,1359,1381]" italics="true" pageId="38" pageNumber="39">Copuetta kwamgumi</emphasis>
</taxonomicName>
<emphasis id="B906E941FFBCFF94FCCFFAB63795FA95" bold="true" box="[882,968,1360,1381]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A235543AFFBCFF94FCCFFAB63795FA95" box="[882,968,1360,1381]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 120. Male palp, ventral view. 121. Same, retrolateral view. 122. Female epigyne, ventral view. 123. Same, dorsal view. Scale bars = 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C36866D8FFBCFF94FF7AFA5736DCF950" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BCD3553FFBCFF94FF7AFA57378FFA1C" blockId="38.[151,1436,1456,1769]" pageId="38" pageNumber="39">
<emphasis id="B906E941FFBCFF94FF7AFA573520FA39" bold="true" box="[199,381,1456,1481]" pageId="38" pageNumber="39">
<typeStatus id="54C98BF1FFBCFF94FF7AFA57355FFA39" box="[199,258,1457,1481]" pageId="38" pageNumber="39">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54C98BF1FFBCFF94FE36FA5635AEFA38" box="[395,499,1456,1480]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
3:
<emphasis id="B906E941FFBCFF94FD97FA56369AFA39" bold="true" box="[554,711,1456,1481]" pageId="38" pageNumber="39">
<collectingCountry id="F36575C3FFBCFF94FD97FA56369FFA39" box="[554,706,1456,1481]" name="Tanzania" pageId="38" pageNumber="39">TANZANIA</collectingCountry>
:
</emphasis>
Tanga Region: Muheza district,
<taxonomicName id="4C724ED0FFBCFF94FBE8FA563084FA38" box="[1109,1241,1456,1480]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="76" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">Kwamgumi</taxonomicName>
Forest Reserve,
<geoCoordinate id="EE465394FFBCFF94FF2AFA3234ACFA1C" box="[151,241,1492,1516]" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="-4.95">04°57'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFBCFF94FF41FA323505FA1C" box="[252,344,1492,1516]" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="38.733334">38°44'E</geoCoordinate>
, leg. S. McKamey
<emphasis id="B906E941FFBCFF94FD92FA30363AFA1D" box="[559,615,1493,1517]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
,
<date id="FFCC1393FFBCFF94FDC8FA3236A3FA1C" box="[629,766,1492,1516]" pageId="38" pageNumber="39" value="1995-07-26">26.VII.1995</date>
(fog 12) (
<collectionCode id="ED63AD96FFBCFF94FCD2FA32379BFA1C" box="[879,966,1492,1516]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="8BCD3553FFBCFF94FF7AFA1F3678F98C" blockId="38.[151,1436,1456,1769]" pageId="38" pageNumber="39">
<typeStatus id="54C98BF1FFBCFF94FF7AFA1F3565F9E1" box="[199,312,1529,1553]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B906E941FFBCFF94FEFFFA1E3582F9E1" bold="true" box="[322,479,1528,1553]" pageId="38" pageNumber="39">
<collectingCountry id="F36575C3FFBCFF94FEFFFA1E3587F9E1" box="[322,474,1528,1553]" name="Tanzania" pageId="38" pageNumber="39">TANZANIA</collectingCountry>
:
</emphasis>
Tanga Region: Muheza district,
<taxonomicName id="4C724ED0FFBCFF94FCF7FA1E3793F9E0" box="[842,974,1528,1552]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="76" phylum="Arthropoda" rank="species" species="kwamgumi" status="sp. nov.">Kwamgumi</taxonomicName>
Forest Reserve,
<geoCoordinate id="EE465394FFBCFF94FB37FA1E30B9F9E0" box="[1162,1252,1528,1552]" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="-4.95">04°57'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFBCFF94FB52FA1E3116F9E0" box="[1263,1355,1528,1552]" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="38.733334">38°44'E</geoCoordinate>
, leg. S. McKamey
<emphasis id="B906E941FFBCFF94FEABF9F8350CF9C5" box="[278,337,1565,1589]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
,
<date id="FFCC1393FFBCFF94FEDCF9FA35B7F9C4" box="[353,490,1564,1588]" pageId="38" pageNumber="39" value="1995-07-20">20.VII.1995</date>
(fog 8), 23 4Ƥ (
<collectionCode id="ED63AD96FFBCFF94FD09F9FA3756F9C4" box="[692,779,1564,1588]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
); same data,
<date id="FFCC1393FFBCFF94FC1EF9FA3071F9C4" box="[931,1068,1564,1588]" pageId="38" pageNumber="39" value="1995-07-22">22.VII.1995</date>
(fog 9), 13 1Ƥ (
<collectionCode id="ED63AD96FFBCFF94FB4BF9FA3110F9C4" box="[1270,1357,1564,1588]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
); same data,
<date id="FFCC1393FFBCFF94FF64F9A6353CF9A8" box="[217,353,1600,1624]" pageId="38" pageNumber="39" value="1995-07-23">23.VII.1995</date>
(fog 10), 13 1Ƥ (
<collectionCode id="ED63AD96FFBCFF94FDF4F9A636FDF9A8" box="[585,672,1600,1624]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
); same data,
<date id="FFCC1393FFBCFF94FCFCF9A63797F9A8" box="[833,970,1600,1624]" pageId="38" pageNumber="39" value="1995-07-25">25.VII.1995</date>
(fog 11), 13 1Ƥ (
<collectionCode id="ED63AD96FFBCFF94FB0DF9A63155F9A8" box="[1200,1288,1600,1624]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
); same data,
<date id="FFCC1393FFBCFF94FF2AF982357DF98C" box="[151,288,1636,1660]" pageId="38" pageNumber="39" value="1995-07-26">26.VII.1995</date>
(fog 12), 1Ƥ (
<collectionCode id="ED63AD96FFBCFF94FE7CF9823645F98C" box="[449,536,1636,1660]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="38" pageNumber="39">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="8BCD3553FFBCFF94FF7AF96E36DCF950" blockId="38.[151,1436,1456,1769]" box="[199,641,1672,1697]" pageId="38" pageNumber="39">
<emphasis id="B906E941FFBCFF94FF7AF96E366BF951" bold="true" box="[199,566,1672,1697]" pageId="38" pageNumber="39">Additional material examined:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFBCFF94FF7AF94A3731F934" box="[199,876,1708,1733]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BCD3553FFBCFF94FF7AF94A3731F934" blockId="38.[151,1436,1456,1769]" box="[199,876,1708,1733]" pageId="38" pageNumber="39">
<emphasis id="B906E941FFBCFF94FF7AF94A353FF935" bold="true" box="[199,354,1708,1733]" pageId="38" pageNumber="39">Distribution:</emphasis>
Known only from the
<typeStatus id="54C98BF1FFBCFF94FDDFF94B36CFF935" box="[610,658,1709,1733]" pageId="38" pageNumber="39">type</typeStatus>
locality (
<figureCitation id="134929D6FFBCFF94FD40F94A3702F934" box="[765,863,1708,1732]" captionStart="FIGURE 132" captionStartId="46.[151,250,1450,1473]" captionTargetBox="[151,1425,193,1417]" captionTargetId="figure@46.[151,1436,193,1429]" captionTargetPageId="46" captionText="FIGURE 132. Distribution of Copuetta comorica sp. nov. (black circles), C. erecta sp. nov. (white circles), C. kakamega sp. nov. (black star), C. kwamgumi sp. nov. (white star), C. lacustris (Strand, 1916) (black triangles), C. lesnei sp. nov. (white triangle) and C. litipo sp. nov. (cross) in the Afrotropical Region." httpUri="https://zenodo.org/record/284223/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFBCFF94FF7AF936305BF918" box="[199,1030,1744,1769]" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="8BCD3553FFBCFF94FF7AF936305BF918" blockId="38.[151,1436,1456,1769]" box="[199,1030,1744,1769]" pageId="38" pageNumber="39">
<emphasis id="B906E941FFBCFF94FF7AF9363577F919" bold="true" box="[199,298,1744,1769]" pageId="38" pageNumber="39">Biology:</emphasis>
All of the specimens were collected by canopy fogging in forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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