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<document id="71BA07C9641200D505D30DBAD1B92D56" ID-CLB-Dataset="22250" ID-DOI="10.5852/ejt.2018.465" ID-GBIF-Dataset="64d84a28-6895-4f01-9ab7-6b4f37cec4ee" ID-Zenodo-Dep="3827023" ID-ZooBank="D73665B3-6043-4528-9A89-CCCBBC90D6B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589218847444" checkinUser="valdenar" docAuthor="Jocqué, Rudy &amp; Henrard, Arnaud" docDate="2018" docId="E16A87BDD01DFFBCFDB5FD917E0832C7" docLanguage="en" docName="ejt-465_jocque_henrard.pdf.imf" docOrigin="European Journal of Taxonomy 465" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cicynethus Simon 1910" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="1D53FFC5D01FFFBAFFEBFFFE7B7F333E" masterDocTitle="A revision of the genus Cicynethus Simon, 1910 (Araneae, Zodariidae), a tale of colour patterns" masterLastPageNumber="35" masterPageNumber="1" pageNumber="3" updateTime="1698839140968" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2B1758D8A366CDB0A2745FBB4C65C2D0">A revision of the genus Cicynethus Simon, 1910 (Araneae, Zodariidae), a tale of colour patterns</mods:title>
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<mods:namePart id="308776202D9B776326F3BDD1811E16D3">Jocqué, Rudy</mods:namePart>
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<mods:affiliation id="4D08869A1849984B73714C51584F100F">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. &amp; Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium.</mods:affiliation>
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<mods:namePart id="4E515EE4A246C30FFF7AA59ADD91454B">Henrard, Arnaud</mods:namePart>
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<mods:affiliation id="218DD835036502EBD4FAE1F31ABCFB43">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. &amp; Earth and Life Institute, Biodiversity Research Center, UCL- 17. 07. 04, Bâtiment Carnoy, Croix du Sud, 5, B- 1348, Louvain-la-Neuve, Belgium.</mods:affiliation>
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<treatment id="E16A87BDD01DFFBCFDB5FD917E0832C7" ID-DOI="http://doi.org/10.5281/zenodo.3843761" ID-GBIF-Taxon="164044209" ID-Zenodo-Dep="3843761" LSID="urn:lsid:plazi:treatment:E16A87BDD01DFFBCFDB5FD917E0832C7" httpUri="http://treatment.plazi.org/id/E16A87BDD01DFFBCFDB5FD917E0832C7" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
<subSubSection id="21D96520D01DFFB8FDB5FD9178AA31B7" box="[606,981,623,650]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="697C36ABD01DFFB8FDB5FD9178AA31B7" blockId="2.[606,981,623,650]" box="[606,981,623,650]" pageId="2" pageNumber="3">
<heading id="323481C7D01DFFB8FDB5FD9178AA31B7" box="[606,981,623,650]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus
<taxonomicName id="AEC34D28D01DFFB8FD5AFD9178AA31B7" ID-CoL="62LBJ" ID-ENA="2033237" authority="Simon, 1910" authorityName="Simon" authorityYear="1910" box="[689,981,623,650]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01DFFB8FD5AFD91784731B4" bold="true" box="[689,824,623,650]" italics="true" pageId="2" pageNumber="3">Cicynethus</emphasis>
Simon, 1910
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<subSubSection id="21D96520D01DFFB8FF56FD4C78CA3030" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="697C36ABD01DFFB8FF56FD4C7A2731F5" blockId="2.[189,344,689,716]" box="[189,344,689,716]" pageId="2" pageNumber="3">
<heading id="323481C7D01DFFB8FF56FD4C7A2731F5" bold="true" box="[189,344,689,716]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="5BB7EAB9D01DFFB8FF56FD4C7A2731F5" bold="true" box="[189,344,689,716]" pageId="2" pageNumber="3">
<typeStatus id="B6788809D01DFFB8FF56FD4C7B8531F2" box="[189,250,690,716]" pageId="2" pageNumber="3">Type</typeStatus>
species
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</heading>
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56FD0A78CA3030" blockId="2.[189,949,755,782]" box="[189,949,755,782]" pageId="2" pageNumber="3">
<taxonomicName id="AEC34D28D01DFFB8FF56FD0A791D3030" ID-CoL="VB67" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[189,610,755,782]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="5BB7EAB9D01DFFB8FF56FD0A7ABB3033" box="[189,452,756,782]" italics="true" pageId="2" pageNumber="3">Cicynethus peringueyi</emphasis>
Simon, 1893
</taxonomicName>
(known from juvenile only).
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<subSubSection id="21D96520D01DFFB8FF56FCCB796030D5" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="697C36ABD01DFFB8FF56FCCB7A493071" blockId="2.[189,310,821,847]" box="[189,310,821,847]" pageId="2" pageNumber="3">
<heading id="323481C7D01DFFB8FF56FCCB7A493071" bold="true" box="[189,310,821,847]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="5BB7EAB9D01DFFB8FF56FCCB7A493071" bold="true" box="[189,310,821,847]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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<taxonomicName id="AEC34D28D01DFFB8FF56FC9B7A403041" authorityName="Simon" authorityYear="1910" box="[189,319,869,895]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01DFFB8FF56FC9B7A403041" box="[189,319,869,895]" italics="true" pageId="2" pageNumber="3">Cicynethus</emphasis>
</taxonomicName>
are medium-sized, elongated spiders with eyes in two procurved rows and spination on legs reduced to a few small distal spines on the ventral side of the metatarsi. It differs from
<taxonomicName id="AEC34D28D01DFFB8FB4AFC777B8530F9" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Zodariidae" genus="Chariobas" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01DFFB8FB4AFC777E62309D" box="[1185,1309,905,931]" italics="true" pageId="2" pageNumber="3">Chariobas</emphasis>
Simon, 1893
</taxonomicName>
, which has the eyes in a close group, and from
<taxonomicName id="AEC34D28D01DFFB8FCDFFC537FE030F9" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[820,1183,941,967]" class="Arachnida" family="Zodariidae" genus="Thaumastochilus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01DFFB8FCDFFC53788330F9" box="[820,1020,941,967]" italics="true" pageId="2" pageNumber="3">Thaumastochilus</emphasis>
Simon, 1897
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, which has strong spines on the anterior leg pair.
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<subSubSection id="21D96520D01DFFBEFF56FBED7F923000" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="697C36ABD01DFFB8FF56FBED7A313713" blockId="2.[189,334,1043,1069]" box="[189,334,1043,1069]" pageId="2" pageNumber="3">
<heading id="323481C7D01DFFB8FF56FBED7A313713" bold="true" box="[189,334,1043,1069]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="5BB7EAB9D01DFFB8FF56FBED7A313713" bold="true" box="[189,334,1043,1069]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56FBBD7E2837D5" blockId="2.[189,1399,1090,1260]" pageId="2" pageNumber="3">
BODY. Medium-sized to large spiders (
<quantity id="AE3B9B4ED01DFFB8FD99FBBC78703763" box="[626,783,1090,1117]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.44" metricValueMin="0.9199999999999999" pageId="2" pageNumber="3" unit="mm" value="11.8" valueMax="14.4" valueMin="9.2">9.214.4 mm</quantity>
) with smooth to finely granulate teguments. Carapace longer than wide (L/W &lt;1.52.2) and fairly flat, protruding anteriorly (
<figureCitation id="F1F82A2ED01DFFB8FBECFB987F0937BE" box="[1031,1142,1126,1152]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[193,1398,725,1865]" captionTargetId="figure@20.[189,1398,725,1886]" captionTargetPageId="20" captionText="Fig. 12. Cicynethus mossambicus sp. nov. AB. ♂, holotype (RMCA_ARA_245528). CD. ♀, paratype (RMCA_ARA_245530). A. Habitus, dorsal view. B. As preceding, lateral view. C. Habitus, dorsal view. D. As preceding, ventral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827048" httpUri="https://zenodo.org/record/3827048/files/figure.png" pageId="2" pageNumber="3">Figs 12B</figureCitation>
,
<figureCitation id="F1F82A2ED01DFFB8FB69FB987FC737BF" box="[1154,1208,1126,1153]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[189,1398,942,1876]" captionTargetId="figure@25.[189,1398,942,1887]" captionTargetPageId="25" captionText="Fig. 16. Cicynethus subtropicalis (Lawrence, 1952) comb. nov. A. ♀, holotype (NM 1161). BC. ♀ (NCA 2005/1860). DE. ♀ (RMCA_ARA_211745). A. Habitus, ventral view. B. Prosoma, ventral view. C. Abdomen, ventral view. D. Prosoma, lateral view. E. Habitus, lateral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827056" httpUri="https://zenodo.org/record/3827056/files/figure.png" pageId="2" pageNumber="3">16D</figureCitation>
), widest at level of coxae IIIII, not much narrowed in front to about 0.60.65 times maximum width (cephalic width measured on strong frontal bend), with few hairs apart from a few longer hairs in front of fovea and on clypeus. Cervical grooves faint. Profile highest just behind eyes and with slight dip at level of fovea.
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56FAEA7A8236A7" blockId="2.[189,1398,1300,1434]" pageId="2" pageNumber="3">
COLOUR. Carapace medium to dark brown; chelicerae, mouthparts and sternum medium to orange brown; leg colour complex, frontal legs often darker and more uniform than posteriors; abdomen dorsum grey with species-specific colour pattern. Remark: colour pattern often darker in vivo (
<figureCitation id="F1F82A2ED01DFFB8FB65FAA57F9D3648" box="[1166,1250,1371,1398]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[189,1384,599,1839]" captionTargetId="figure@24.[189,1398,599,1849]" captionTargetPageId="24" captionText="Fig. 15. Cicynethus subtropicalis (Lawrence, 1952) comb. nov., habitus, dorsal views. AB. ♂, in vivo (Nambiti Game Reserve, Kwazulu Natal; courtesy of Bennie Kruger). C. ♂ (NCA 2007/3035). D. ♀, holotype (NM 1161). E. ♀ (NCA 2005/1820). F. ♀ (RMCA_ARA_211745). G. ♀ (NCA 2007/3556). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827054" httpUri="https://zenodo.org/record/3827054/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
AB), much more contrasted in alcohol.
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56FA3C7A0A351A" blockId="2.[189,1398,1473,1572]" pageId="2" pageNumber="3">
EYES. Eyes in two procurved rows (
<figureCitation id="F1F82A2ED01DFFB8FDB3FA3F79CB36E2" box="[600,692,1473,1500]" captionStart="Fig" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[189,1398,1157,1927]" captionTargetId="figure@9.[189,1398,1157,1927]" captionTargetPageId="9" captionText="Fig. 5. Cicynethus acer sp. nov., juvenile (RMCA_ARA_131816). A. Habitus, dorsal view. B. As preceding, lateral view. C. As preceding, frontal view. Scale bars: 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3827034" httpUri="https://zenodo.org/record/3827034/files/figure.png" pageId="2" pageNumber="3">Fig. 5C</figureCitation>
). All eyes pale and subequal, AME usually the largest. MOQ quadrangular or slightly longer than wide. Clypeus straight, height 0.8 to 2 times diameter of ALE, with dispersed setae.
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56F9B278353403" blockId="2.[189,1399,1611,1854]" pageId="2" pageNumber="3">
PROSOMA. Chilum double, poorly sclerotized, about or slightly more than twice as wide as high, lateral margin poorly defined, with a few setae. Chelicerae without teeth, broad from base to tip with evenly dispersed setae; fangs shorter than wide at base. Labium diamond-shaped, widest in distal half, narrowed base. Endites roughly triangular, converging, with basolateral extension accommodating palpal coxa; with field of thick setae in distal half. Sternum elongated, roughly oval (
<figureCitation id="F1F82A2ED01DFFB8FBF0F9257F0335CB" box="[1051,1148,1755,1781]" captionStart="Fig" captionStartId="13.[189,232,1931,1957]" captionTargetBox="[191,1398,818,1874]" captionTargetId="figure@13.[189,1398,818,1886]" captionTargetPageId="13" captionText="Fig. 8. Cicynethus decoratus (Lawrence, 1952) comb. nov. AD. ♀ (RMCA_ARA_131806). E. Subadult ♂ (same sample). A. Habitus, ventral view. B. As preceding, lateral view. C. Prosoma, ventral view. D. Female palp, prolateral view. E. Habitus, dorsal view. Scale bars: AB, E = 2 mm; CD = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3827040" httpUri="https://zenodo.org/record/3827040/files/figure.png" pageId="2" pageNumber="3">Figs 8C</figureCitation>
,
<figureCitation id="F1F82A2ED01DFFB8FB61F9257FAE35C8" box="[1162,1233,1755,1782]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[189,1398,612,1887]" captionTargetId="figure@15.[189,1398,612,1887]" captionTargetPageId="15" captionText="Fig. 9. Cicynethus floriumfontis Jocqué, 1991. AC. ♂ (NCA 95/385). DF. ♀ (NCA 95/243). A. Habitus, dorsal view. B. As preceding, ventral view. C. Carapace, dorsal view. D. Habitus, dorsal view. E. As preceding, ventral view. F. Carapace, dorsal view. Scale bars: AB, DE = 2 mm; C, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3827042" httpUri="https://zenodo.org/record/3827042/files/figure.png" pageId="2" pageNumber="3">9B, E</figureCitation>
,
<figureCitation id="F1F82A2ED01DFFB8FB34F9257E6B35CB" box="[1247,1300,1755,1781]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[193,1398,725,1865]" captionTargetId="figure@20.[189,1398,725,1886]" captionTargetPageId="20" captionText="Fig. 12. Cicynethus mossambicus sp. nov. AB. ♂, holotype (RMCA_ARA_245528). CD. ♀, paratype (RMCA_ARA_245530). A. Habitus, dorsal view. B. As preceding, lateral view. C. Habitus, dorsal view. D. As preceding, ventral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827048" httpUri="https://zenodo.org/record/3827048/files/figure.png" pageId="2" pageNumber="3">12D</figureCitation>
), longer than wide, widest at middle, without triangular extensions; sometimes with a few precoxal sclerites (
<figureCitation id="F1F82A2ED01DFFB8FF2DF8DD7A4E3400" box="[198,305,1827,1854]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[189,1398,942,1876]" captionTargetId="figure@25.[189,1398,942,1887]" captionTargetPageId="25" captionText="Fig. 16. Cicynethus subtropicalis (Lawrence, 1952) comb. nov. A. ♀, holotype (NM 1161). BC. ♀ (NCA 2005/1860). DE. ♀ (RMCA_ARA_211745). A. Habitus, ventral view. B. Prosoma, ventral view. C. Abdomen, ventral view. D. Prosoma, lateral view. E. Habitus, lateral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827056" httpUri="https://zenodo.org/record/3827056/files/figure.png" pageId="2" pageNumber="3">Fig. 16B</figureCitation>
); anterior margin straight or slightly concave.
</paragraph>
<paragraph id="697C36ABD01DFFB8FF56F8987E0934D5" blockId="2.[189,1398,1893,2027]" pageId="2" pageNumber="3">
LEGS. Fairly slender, anterior legs more robust than posterior ones. Formula 1243 or 1423. Spination reduced to a few short ventral spines on Mt, arranged as 13 along distal margin and 1 or 2 subdistal behind the middle distal one. Paired tarsal claws with 1014 teeth (
<figureCitation id="F1F82A2ED01DFFB8FC05F8537F3234F9" box="[1006,1101,1965,1991]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
); unpaired claw smooth, strongly reduced on hind legs. Tarsal organ capsulate (
<figureCitation id="F1F82A2ED01DFFB8FCDAF82F78F634D5" box="[817,905,2001,2027]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="2" pageNumber="3">Fig. 1E</figureCitation>
). Scopulae well developed. Metatarsi with
</paragraph>
<paragraph id="697C36ABD01CFFB9FF56FEF57FCC3277" blockId="3.[189,1399,267,329]" pageId="3" pageNumber="4">
distal hood (MtS) extended to ventrolateral side (
<figureCitation id="F1F82A2ED01CFFB9FD12FEF5782C321B" box="[761,851,267,293]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
). Tibial process well developed (
<figureCitation id="F1F82A2ED01CFFB9FB3FFEF57E4F321B" box="[1236,1328,267,293]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="3" pageNumber="4">Fig. 1D</figureCitation>
). One dorsal hinged hair on tibiae and metatarsi I and II. Bothria with one ridge (
<figureCitation id="F1F82A2ED01CFFB9FBC3FED07F163277" box="[1064,1129,302,329]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
FG).
</paragraph>
<paragraph id="697C36ABD01CFFBEFF56FE8E7E5A32AC" blockId="3.[189,1399,368,430]" lastBlockId="4.[189,1399,267,402]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
FEMALE PALP. Female palpal tarsus conical, with short but strong prolateral spines (
<figureCitation id="F1F82A2ED01CFFB9FB96FE8E7FA432B4" box="[1149,1243,368,394]" captionStart="Fig" captionStartId="13.[189,232,1931,1957]" captionTargetBox="[191,1398,818,1874]" captionTargetId="figure@13.[189,1398,818,1886]" captionTargetPageId="13" captionText="Fig. 8. Cicynethus decoratus (Lawrence, 1952) comb. nov. AD. ♀ (RMCA_ARA_131806). E. Subadult ♂ (same sample). A. Habitus, ventral view. B. As preceding, lateral view. C. Prosoma, ventral view. D. Female palp, prolateral view. E. Habitus, dorsal view. Scale bars: AB, E = 2 mm; CD = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3827040" httpUri="https://zenodo.org/record/3827040/files/figure.png" pageId="3" pageNumber="4">Fig. 8D</figureCitation>
); palpal claw with some small teeth at base (
<figureCitation id="F1F82A2ED01CFFB9FDCBFE6D79053293" box="[544,634,403,429]" captionStart="Fig" captionStartId="3.[189,232,1824,1850]" captionTargetBox="[189,1398,562,1782]" captionTargetId="figure@3.[189,1398,562,1782]" captionTargetPageId="3" captionText="Fig. 1. SEM micrographs of Cicynethus subtropicalis (Lawrence, 1952) comb. nov. AE. ♀ (NCA 2000/438). FG. ♂ (NCA 2008/4266). A. Paired tarsal claw II, retrolateral view. B. Palpal claw, prolateral view. C. Metatarsus II, apex, showing metatarsal stopper (MtS). D. Tibial process (arrow), left leg IV, dorsal view. E. Tarsal organ, leg IV, dorsal view. F. Bothrium, metatarsus IV, dorsal view. G. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="3" pageNumber="4">Fig. 1B</figureCitation>
); not turned inward; without distal patch of chemosensitive setae. ABDOMEN. Elongated oval or cylindrical, at least twice as long as wide; tracheal spiracle fairly small, somewhat advanced and provided with small rectangular scutellum. Both sexes with six spinnerets. ALS large, conical, biarticulate. PLS cylindrical, biarticulate, with 4 cylindrical gland spigots; PMS small, cylindrical, one-segmented with 2 cylindrical gland spigots. Colulus represented by haired field.
</paragraph>
<caption id="3DBC6623D01CFFB9FF56F8DE79CB34D4" ID-DOI="http://doi.org/10.5281/zenodo.3827026" ID-Zenodo-Dep="3827026" httpUri="https://zenodo.org/record/3827026/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1824,1850]" targetBox="[189,1398,562,1782]" targetPageId="3">
<paragraph id="697C36ABD01CFFB9FF56F8DE79CB34D4" blockId="3.[189,1399,1824,2027]" pageId="3" pageNumber="4">
<emphasis id="5BB7EAB9D01CFFB9FF56F8DE7A703404" bold="true" box="[189,271,1824,1850]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
SEM micrographs of
<taxonomicName id="AEC34D28D01CFFB9FDC8F8DE7F563405" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" box="[547,1065,1824,1851]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="subtropicalis" status="comb. nov.">
<emphasis id="5BB7EAB9D01CFFB9FDC8F8DE78363404" box="[547,841,1824,1850]" italics="true" pageId="3" pageNumber="4">Cicynethus subtropicalis</emphasis>
(
<bibRefCitation id="0D524B5AD01CFFB9FCB7F8DE7F5F3405" author="Lawrence R. F." box="[860,1056,1824,1851]" pageId="3" pageNumber="4" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D01CFFB9FBDFF8DE7FC53404" box="[1076,1210,1824,1850]" pageId="3" pageNumber="4" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="5BB7EAB9D01CFFB9FB2EF8DE7F833404" bold="true" box="[1221,1276,1824,1850]" pageId="3" pageNumber="4">AE</emphasis>
. ♀ (NCA 2000/438).
<emphasis id="5BB7EAB9D01CFFB9FEA1F8BD7AFE3463" bold="true" box="[330,385,1859,1885]" pageId="3" pageNumber="4">FG</emphasis>
. ♂ (NCA 2008/4266).
<emphasis id="5BB7EAB9D01CFFB9FD43F8BD79C23463" bold="true" box="[680,701,1859,1885]" pageId="3" pageNumber="4">A</emphasis>
. Paired tarsal claw II, retrolateral view.
<emphasis id="5BB7EAB9D01CFFB9FB50F8BD7FB03463" bold="true" box="[1211,1231,1859,1885]" pageId="3" pageNumber="4">B</emphasis>
. Palpal claw, prolateral view.
<emphasis id="5BB7EAB9D01CFFB9FE94F8987AEB34BE" bold="true" box="[383,404,1894,1920]" pageId="3" pageNumber="4">C</emphasis>
. Metatarsus II, apex, showing metatarsal stopper (MtS).
<emphasis id="5BB7EAB9D01CFFB9FBA9F8987F2834BE" bold="true" box="[1090,1111,1894,1920]" pageId="3" pageNumber="4">D</emphasis>
. Tibial process (arrow), left leg IV, dorsal view.
<emphasis id="5BB7EAB9D01CFFB9FE36F8747A8E349A" bold="true" box="[477,497,1930,1956]" pageId="3" pageNumber="4">E</emphasis>
. Tarsal organ, leg IV, dorsal view.
<emphasis id="5BB7EAB9D01CFFB9FC78F87478DA349A" bold="true" box="[915,933,1930,1956]" pageId="3" pageNumber="4">F</emphasis>
. Bothrium, metatarsus IV, dorsal view.
<emphasis id="5BB7EAB9D01CFFB9FF56F8527BAB34F8" bold="true" box="[189,212,1964,1990]" pageId="3" pageNumber="4">G</emphasis>
. Trichobothrium, base of RTA, retrolateral view. Abbreviation: MtS = metatarsal stopper. Scale bars: A = 0.1 mm; BD = 50 µm; EG = 10 µm.
</paragraph>
</caption>
<paragraph id="697C36ABD01BFFBEFF56FE4578533158" blockId="4.[189,1399,442,614]" pageId="4" pageNumber="5">
MALE PALP. Structure of male palp stable; RTA characteristic, with broad and long basal shaft ending in widened tip with two blades often perpendicular to each other (
<figureCitation id="F1F82A2ED01BFFBEFC52FE207F7E32C6" box="[953,1025,478,504]" captionStart="Fig" captionStartId="5.[189,232,1792,1818]" captionTargetBox="[189,1398,265,1768]" captionTargetId="figure@5.[189,1398,265,1768]" captionTargetPageId="5" captionText="Fig. 2. SEM micrographs of male palps and RTA. AC. Cicynethus floriumfontis Jocqué, 1991, ♂ (NCA 91/1051). DE. C. mossambicus sp. nov., ♂ (RMCA_ARA_245567). FH. C. subtropicalis (Lawrence, 1952) comb. nov., ♂ (NCA 2008/4266). A. Male palp, ventro-retrolateral view. B. RTA apex, ventroretrolateral view. C. As preceding, retrolateral view. D. Male palp, ventro-retrolateral view. E. RTA apex, ventro-retrolateral view. F. Male palp, retrolateral view. G. As preceding, ventro-retrolateral view. H. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
AH). Cymbium without distal claw (
<figureCitation id="F1F82A2ED01BFFBEFEE3FDFC7AEF3123" box="[264,400,514,541]" captionStart="Fig" captionStartId="5.[189,232,1792,1818]" captionTargetBox="[189,1398,265,1768]" captionTargetId="figure@5.[189,1398,265,1768]" captionTargetPageId="5" captionText="Fig. 2. SEM micrographs of male palps and RTA. AC. Cicynethus floriumfontis Jocqué, 1991, ♂ (NCA 91/1051). DE. C. mossambicus sp. nov., ♂ (RMCA_ARA_245567). FH. C. subtropicalis (Lawrence, 1952) comb. nov., ♂ (NCA 2008/4266). A. Male palp, ventro-retrolateral view. B. RTA apex, ventroretrolateral view. C. As preceding, retrolateral view. D. Male palp, ventro-retrolateral view. E. RTA apex, ventro-retrolateral view. F. Male palp, retrolateral view. G. As preceding, ventro-retrolateral view. H. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, D</figureCitation>
), with a few distal, slightly sinuous truncated spines (
<figureCitation id="F1F82A2ED01BFFBEFBC8FDFC7FFE3123" box="[1059,1153,514,541]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="4" pageNumber="5">Figs 3F</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFB7BFDFC7FCA3123" box="[1168,1205,514,541]" captionStart="Fig" captionStartId="10.[189,232,1929,1955]" captionTargetBox="[224,1359,856,1882]" captionTargetId="figure@10.[191,1396,818,1907]" captionTargetPageId="10" captionText="Fig. 6. Cicynethus acer sp. nov., genitalia drawings. A. Male palp, retrolateral view. B. As preceding, ventral view. C. Epigyne, ventral view. Abbreviations: E = embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; TS = tegular swelling. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827036" httpUri="https://zenodo.org/record/3827036/files/figure.png" pageId="4" pageNumber="5">6B</figureCitation>
); tegulum often strongly swollen; embolus fairly short, originating prolaterally on posterior part of tegulum (
<figureCitation id="F1F82A2ED01BFFBEFAE5FDD97E0D317F" box="[1294,1394,551,577]" captionStart="Fig" captionStartId="5.[189,232,1792,1818]" captionTargetBox="[189,1398,265,1768]" captionTargetId="figure@5.[189,1398,265,1768]" captionTargetPageId="5" captionText="Fig. 2. SEM micrographs of male palps and RTA. AC. Cicynethus floriumfontis Jocqué, 1991, ♂ (NCA 91/1051). DE. C. mossambicus sp. nov., ♂ (RMCA_ARA_245567). FH. C. subtropicalis (Lawrence, 1952) comb. nov., ♂ (NCA 2008/4266). A. Male palp, ventro-retrolateral view. B. RTA apex, ventroretrolateral view. C. As preceding, retrolateral view. D. Male palp, ventro-retrolateral view. E. RTA apex, ventro-retrolateral view. F. Male palp, retrolateral view. G. As preceding, ventro-retrolateral view. H. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="4" pageNumber="5">Figs 2D</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFF56FDB57BA3315B" box="[189,220,587,613]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="4" pageNumber="5">3F</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFF03FDB57A723158" box="[232,269,587,614]" captionStart="Fig" captionStartId="10.[189,232,1929,1955]" captionTargetBox="[224,1359,856,1882]" captionTargetId="figure@10.[191,1396,818,1907]" captionTargetPageId="10" captionText="Fig. 6. Cicynethus acer sp. nov., genitalia drawings. A. Male palp, retrolateral view. B. As preceding, ventral view. C. Epigyne, ventral view. Abbreviations: E = embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; TS = tegular swelling. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827036" httpUri="https://zenodo.org/record/3827036/files/figure.png" pageId="4" pageNumber="5">6B</figureCitation>
); MA strongly curved outwards (
<figureCitation id="F1F82A2ED01BFFBEFD79FDB57992315B" box="[658,749,587,613]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="4" pageNumber="5">Figs 3F</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFD12FDB578613158" box="[761,798,587,614]" captionStart="Fig" captionStartId="10.[189,232,1929,1955]" captionTargetBox="[224,1359,856,1882]" captionTargetId="figure@10.[191,1396,818,1907]" captionTargetPageId="10" captionText="Fig. 6. Cicynethus acer sp. nov., genitalia drawings. A. Male palp, retrolateral view. B. As preceding, ventral view. C. Epigyne, ventral view. Abbreviations: E = embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; TS = tegular swelling. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827036" httpUri="https://zenodo.org/record/3827036/files/figure.png" pageId="4" pageNumber="5">6B</figureCitation>
).
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FD707F9131F5" blockId="4.[189,1399,653,716]" pageId="4" pageNumber="5">
EPIGYNE. With two central depressions separated by septum, in front of transverse plate (
<figureCitation id="F1F82A2ED01BFFBEFB5DFD737E6C3199" box="[1206,1299,653,679]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[199,1398,702,1898]" captionTargetId="figure@8.[189,1398,702,1898]" captionTargetPageId="8" captionText="Fig. 4. Cicynethus acer sp. nov., paratype, ♀ (TM 13375). A. Habitus, dorsal view. B. As preceding, closer view. C. As preceding, ventral view. D. As preceding, lateral view. E. Epigyne, ventral view. Scale bars: AD = 5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827032" httpUri="https://zenodo.org/record/3827032/files/figure.png" pageId="4" pageNumber="5">Figs 4E</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFAF5FD707E533196" box="[1310,1324,654,680]" captionStart="Fig" captionStartId="12.[189,232,1929,1955]" captionTargetBox="[189,1398,720,1876]" captionTargetId="figure@12.[189,1398,707,1889]" captionTargetPageId="12" captionText="Fig. 7. Cicynethus decoratus (Lawrence, 1952) comb. nov. A, D. holotype,♀. B, E, G. Female specimen (NCA2014/3298). C, F, H. Female specimen (RMCA_ARA_131806). AC. Habitus, dorsal view. DH. Epigyne, ventral view. Scale bars: AC = 2 mm; DH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827038" httpUri="https://zenodo.org/record/3827038/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
DH). Spermathecae large, reniform, far apart (
<figureCitation id="F1F82A2ED01BFFBEFD72FD4F787E31F5" box="[665,769,689,715]" captionStart="Fig" captionStartId="21.[189,232,1858,1884]" captionTargetBox="[189,1398,889,1816]" captionTargetId="figure@21.[189,1398,889,1816]" captionTargetPageId="21" captionText="Fig. 13. Cicynethus mossambicus sp. nov. AC. ♂, holotype (RMCA_ARA_245528). DF. ♀, paratype (RMCA_ARA_245530). A. Male palp, ventral view. B. As preceding, retrolateral view. C. Distal tip of RTA, retrolateral view. D. Epigyne, ventral view. E. Epigyne, digested, ventral view. F. As preceding, dorsal view. Abbreviations: DTP = distal tegular protrusion; E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827050" httpUri="https://zenodo.org/record/3827050/files/figure.png" pageId="4" pageNumber="5">Fig. 13F</figureCitation>
); copulatory ducts short, with thick walls.
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FD0A7A013030" blockId="4.[189,382,756,782]" box="[189,382,756,782]" pageId="4" pageNumber="5">
<heading id="323481C7D01BFFBEFF56FD0A7A013030" bold="true" box="[189,382,756,782]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="5BB7EAB9D01BFFBEFF56FD0A7A013030" bold="true" box="[189,382,756,782]" pageId="4" pageNumber="5">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FCDD7F923000" blockId="4.[189,1261,803,830]" box="[189,1261,803,830]" pageId="4" pageNumber="5">
See the Discussion for the few observations concerning the habits of species of
<taxonomicName id="AEC34D28D01BFFBEFB88FCDD7F9A3003" authorityName="Simon" authorityYear="1910" box="[1123,1253,803,829]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01BFFBEFB88FCDD7F9A3003" box="[1123,1253,803,829]" italics="true" pageId="4" pageNumber="5">Cicynethus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="21D96520D01BFFBEFF56FC9B788A3603" pageId="4" pageNumber="5" type="distribution">
<paragraph id="697C36ABD01BFFBEFF56FC9B7A293041" blockId="4.[189,342,869,895]" box="[189,342,869,895]" pageId="4" pageNumber="5">
<heading id="323481C7D01BFFBEFF56FC9B7A293041" bold="true" box="[189,342,869,895]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="5BB7EAB9D01BFFBEFF56FC9B7A293041" bold="true" box="[189,342,869,895]" pageId="4" pageNumber="5">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FC6B79F230ED" blockId="4.[189,1399,917,979]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FC6B7A403091" authorityName="Simon" authorityYear="1910" box="[189,319,917,943]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="5BB7EAB9D01BFFBEFF56FC6B7A403091" box="[189,319,917,943]" italics="true" pageId="4" pageNumber="5">Cicynethus</emphasis>
</taxonomicName>
is found in
<collectingCountry id="11D4763BD01BFFBEFE31FC6B790C3091" box="[474,627,917,943]" name="South Africa" pageId="4" pageNumber="5">South Africa</collectingCountry>
and
<collectingCountry id="11D4763BD01BFFBEFD58FC6B782E3091" box="[691,849,917,943]" name="Mozambique" pageId="4" pageNumber="5">Mozambique</collectingCountry>
, with its northernmost locality in the Nyassa Province of
<collectingCountry id="11D4763BD01BFFBEFEA7FC477A9830ED" box="[332,487,953,979]" name="Mozambique" pageId="4" pageNumber="5">Mozambique</collectingCountry>
(
<figureCitation id="F1F82A2ED01BFFBEFE1CFC4779FF30ED" box="[503,640,953,979]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="30.[189,232,1929,1955]" captionStartId-1="31.[189,232,2000,2026]" captionTargetBox-0="[189,1398,1107,1888]" captionTargetBox-1="[189,1398,931,1954]" captionTargetId-0="figure@30.[189,1398,1107,1888]" captionTargetId-1="figure@31.[189,1398,931,1954]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="Fig. 21. Distribution map: Cicynethus acer sp. nov. (●); C. decoratus (Lawrence, 1952) comb. nov. (★); C. floriumfontis Jocqué, 1991 (■); C. peringueyi Simon, 1910 (▲); C. subtropicalis (Lawrence, 1952) comb. nov. (Ä)." captionText-1="Fig. 22. Distribution of Cicynethus mossambicus sp. nov. (A)." figureDoi-0="http://doi.org/10.5281/zenodo.3827066" figureDoi-1="http://doi.org/10.5281/zenodo.3827070" httpUri-0="https://zenodo.org/record/3827066/files/figure.png" httpUri-1="https://zenodo.org/record/3827070/files/figure.png" pageId="4" pageNumber="5">Figs 2122</figureCitation>
).
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FC057AF3372B" blockId="4.[189,396,1019,1045]" box="[189,396,1019,1045]" pageId="4" pageNumber="5">
<heading id="323481C7D01BFFBEFF56FC057AF3372B" bold="true" box="[189,396,1019,1045]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="5BB7EAB9D01BFFBEFF56FC057AF3372B" bold="true" box="[189,396,1019,1045]" pageId="4" pageNumber="5">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FBD5795C377B" blockId="4.[189,902,1067,1275]" box="[189,547,1067,1094]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FBD57A04377B" authority="Jocqué &amp; Henrard, 2018" authorityName="Jocqué &amp; Henrard" authorityYear="2018" box="[189,379,1067,1093]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acer" status="sp. nov.">
<emphasis id="5BB7EAB9D01BFFBEFF56FBD57A04377B" box="[189,379,1067,1093]" italics="true" pageId="4" pageNumber="5">Cicynethus acer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFE69FBD27AA23778" box="[386,477,1068,1094]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(♂
<specimenCount id="7FC5FD22D01BFFBEFDEDFBD57966377B" box="[518,537,1067,1093]" pageId="4" pageNumber="5" type="female"></specimenCount>
)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FBB178343757" blockId="4.[189,902,1067,1275]" box="[189,843,1103,1130]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FBB179ED3754" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" box="[189,658,1103,1130]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="decoratus" status="comb. nov.">
<emphasis id="5BB7EAB9D01BFFBEFF56FBB17AC43757" box="[189,443,1103,1129]" italics="true" pageId="4" pageNumber="5">Cicynethus decoratus</emphasis>
(
<bibRefCitation id="0D524B5AD01BFFBEFE21FBB179F53754" author="Lawrence R. F." box="[458,650,1103,1130]" pageId="4" pageNumber="5" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFD71FBB178643754" box="[666,795,1103,1130]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
(
<specimenCount id="7FC5FD22D01BFFBEFCC6FBB1783F3757" box="[813,832,1103,1129]" pageId="4" pageNumber="5" type="female"></specimenCount>
)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FB8D79B937B0" blockId="4.[189,902,1067,1275]" box="[189,710,1139,1166]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FB8D790037B0" authority="Jocque, 1991" authorityName="Jocque" authorityYear="1991" box="[189,639,1139,1166]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="floriumfontis">
<emphasis id="5BB7EAB9D01BFFBEFF56FB8D7AA237B3" box="[189,477,1139,1165]" italics="true" pageId="4" pageNumber="5">Cicynethus floriumfontis</emphasis>
<bibRefCitation id="0D524B5AD01BFFBEFE0FFB8D790037B0" author="Jocque R." box="[484,639,1139,1166]" pageId="4" pageNumber="5" pagination="1 - 160" refId="ref11880" refString="Jocque R. 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1 - 160." type="journal article" year="1991">Jocqué, 1991</bibRefCitation>
</taxonomicName>
(♂
<specimenCount id="7FC5FD22D01BFFBEFD43FB8A79C437B0" box="[680,699,1140,1166]" pageId="4" pageNumber="5" type="female"></specimenCount>
)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FB6679F4378C" blockId="4.[189,902,1067,1275]" box="[189,651,1176,1202]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FB667A9C378C" authority="Jocqué &amp; Henrard, 2018" authorityName="Jocqué &amp; Henrard" authorityYear="2018" box="[189,483,1176,1202]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mossambicus" status="sp. nov.">
<emphasis id="5BB7EAB9D01BFFBEFF56FB667A9C378C" box="[189,483,1176,1202]" italics="true" pageId="4" pageNumber="5">Cicynethus mossambicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFE01FB66793A378C" box="[490,581,1176,1202]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(♂
<specimenCount id="7FC5FD22D01BFFBEFD85FB6679FE378C" box="[622,641,1176,1202]" pageId="4" pageNumber="5" type="female"></specimenCount>
)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FB42799937E9" blockId="4.[189,902,1067,1275]" box="[189,742,1212,1239]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FB42791D37E8" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[189,610,1212,1238]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="5BB7EAB9D01BFFBEFF56FB427ABB37E8" box="[189,452,1212,1238]" italics="true" pageId="4" pageNumber="5">Cicynethus peringueyi</emphasis>
Simon, 1893
</taxonomicName>
(juveniles)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FB1E78F937C4" blockId="4.[189,902,1067,1275]" box="[189,902,1248,1275]" pageId="4" pageNumber="5">
<taxonomicName id="AEC34D28D01BFFBEFF56FB1E79C937C5" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" box="[189,694,1248,1275]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subtropicalis" status="comb. nov.">
<emphasis id="5BB7EAB9D01BFFBEFF56FB1E7AA037C4" box="[189,479,1248,1274]" italics="true" pageId="4" pageNumber="5">Cicynethus subtropicalis</emphasis>
(
<bibRefCitation id="0D524B5AD01BFFBEFE05FB1E79D137C5" author="Lawrence R. F." box="[494,686,1248,1275]" pageId="4" pageNumber="5" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFD55FB1E783F37C4" box="[702,832,1248,1274]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
(♂
<specimenCount id="7FC5FD22D01BFFBEFC83FB1E780437C4" box="[872,891,1248,1274]" pageId="4" pageNumber="5" type="female"></specimenCount>
)
</paragraph>
<paragraph id="697C36ABD01BFFBEFF56FADC788A3603" blockId="4.[189,1013,1314,1341]" box="[189,1013,1314,1341]" pageId="4" pageNumber="5">
<emphasis id="5BB7EAB9D01BFFBEFF56FADC7AE43602" bold="true" box="[189,411,1314,1340]" pageId="4" pageNumber="5">Key to the species</emphasis>
(specimens in ethanol;
<taxonomicName id="AEC34D28D01BFFBEFD59FADD782E3602" authorityName="Simon" authorityYear="1893" box="[690,849,1315,1341]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="5BB7EAB9D01BFFBEFD59FADD782E3602" box="[690,849,1315,1341]" italics="true" pageId="4" pageNumber="5">C. peringueyi</emphasis>
</taxonomicName>
not included)
</paragraph>
</subSubSection>
<subSubSection id="21D96520D01BFFBCFF56FAAC7E0832C7" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="key">
<key id="1D52619AD01BFFBCFF56FAAC7E0832C7" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<keyStep id="D2372E0ED01BFFBEFF56FAAC7E0835B3" pageId="4" pageNumber="5">
<paragraph id="697C36ABD01BFFBEFF56FAAC7E0836C3" blockId="4.[189,1399,1362,1678]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF56FAAC7E0836C3" pageId="4" pageNumber="5">
1. Cephalothorax uniformly dark chestnut brown, with small reddish spot in front of posterior margin (
<figureCitation id="F1F82A2ED01BFFBEFF18FA887AF436AE" box="[243,395,1398,1424]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[193,1398,725,1865]" captionTargetId="figure@20.[189,1398,725,1886]" captionTargetPageId="20" captionText="Fig. 12. Cicynethus mossambicus sp. nov. AB. ♂, holotype (RMCA_ARA_245528). CD. ♀, paratype (RMCA_ARA_245530). A. Habitus, dorsal view. B. As preceding, lateral view. C. Habitus, dorsal view. D. As preceding, ventral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827048" httpUri="https://zenodo.org/record/3827048/files/figure.png" pageId="4" pageNumber="5">Fig. 12A, C</figureCitation>
); legs covered with numerous long hairs (
<figureCitation id="F1F82A2ED01BFFBEFC73FA88788E36AE" box="[920,1009,1398,1424]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[193,1398,725,1865]" captionTargetId="figure@20.[189,1398,725,1886]" captionTargetPageId="20" captionText="Fig. 12. Cicynethus mossambicus sp. nov. AB. ♂, holotype (RMCA_ARA_245528). CD. ♀, paratype (RMCA_ARA_245530). A. Habitus, dorsal view. B. As preceding, lateral view. C. Habitus, dorsal view. D. As preceding, ventral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827048" httpUri="https://zenodo.org/record/3827048/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
AD); male palpal bulbus, with well developed distal tegular protrusion (DTP), without retrolateral boss and not strongly swollen (
<figureCitation id="F1F82A2ED01BFFBEFF18FA407A2B36E6" box="[243,340,1470,1496]" captionStart="Fig" captionStartId="5.[189,232,1792,1818]" captionTargetBox="[189,1398,265,1768]" captionTargetId="figure@5.[189,1398,265,1768]" captionTargetPageId="5" captionText="Fig. 2. SEM micrographs of male palps and RTA. AC. Cicynethus floriumfontis Jocqué, 1991, ♂ (NCA 91/1051). DE. C. mossambicus sp. nov., ♂ (RMCA_ARA_245567). FH. C. subtropicalis (Lawrence, 1952) comb. nov., ♂ (NCA 2008/4266). A. Male palp, ventro-retrolateral view. B. RTA apex, ventroretrolateral view. C. As preceding, retrolateral view. D. Male palp, ventro-retrolateral view. E. RTA apex, ventro-retrolateral view. F. Male palp, retrolateral view. G. As preceding, ventro-retrolateral view. H. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="4" pageNumber="5">Figs 2D</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFE8BFA407A0336E6" box="[352,380,1470,1496]" captionStart="Fig" captionStartId="21.[189,232,1858,1884]" captionTargetBox="[189,1398,889,1816]" captionTargetId="figure@21.[189,1398,889,1816]" captionTargetPageId="21" captionText="Fig. 13. Cicynethus mossambicus sp. nov. AC. ♂, holotype (RMCA_ARA_245528). DF. ♀, paratype (RMCA_ARA_245530). A. Male palp, ventral view. B. As preceding, retrolateral view. C. Distal tip of RTA, retrolateral view. D. Epigyne, ventral view. E. Epigyne, digested, ventral view. F. As preceding, dorsal view. Abbreviations: DTP = distal tegular protrusion; E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827050" httpUri="https://zenodo.org/record/3827050/files/figure.png" pageId="4" pageNumber="5">13</figureCitation>
AB); epigyne with thick transverse plate and short, well-defined thick median septum (
<figureCitation id="F1F82A2ED01BFFBEFF18FA1C7A3B36C2" box="[243,324,1506,1532]" captionStart="Fig" captionStartId="21.[189,232,1858,1884]" captionTargetBox="[189,1398,889,1816]" captionTargetId="figure@21.[189,1398,889,1816]" captionTargetPageId="21" captionText="Fig. 13. Cicynethus mossambicus sp. nov. AC. ♂, holotype (RMCA_ARA_245528). DF. ♀, paratype (RMCA_ARA_245530). A. Male palp, ventral view. B. As preceding, retrolateral view. C. Distal tip of RTA, retrolateral view. D. Epigyne, ventral view. E. Epigyne, digested, ventral view. F. As preceding, dorsal view. Abbreviations: DTP = distal tegular protrusion; E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827050" httpUri="https://zenodo.org/record/3827050/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
DF) ................................................................................................
<taxonomicName id="AEC34D28D01BFFBEFBBAFA1C7E6A36C3" ID-CoL="VB66" authority="Jocqué &amp; Henrard, 2018" authorityName="Jocqué &amp; Henrard" authorityYear="2018" box="[1105,1301,1506,1533]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mossambicus" status="sp. nov.">
<emphasis id="5BB7EAB9D01BFFBEFBBAFA1C7E6A36C3" bold="true" box="[1105,1301,1506,1533]" italics="true" pageId="4" pageNumber="5">C. mossambicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFAF7FA1D7E0836C3" box="[1308,1399,1507,1533]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF2AF9F97E0835B3" blockId="4.[189,1399,1362,1678]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF2AF9F97E0835B3" pageId="4" pageNumber="5">
Cephalothorax differently coloured and with pattern; legs covered with shorter hairs; male palpal bulbus strongly swollen (
<figureCitation id="F1F82A2ED01BFFBEFDF9F9D5790C357B" box="[530,627,1579,1605]" captionStart="Fig" captionStartId="5.[189,232,1792,1818]" captionTargetBox="[189,1398,265,1768]" captionTargetId="figure@5.[189,1398,265,1768]" captionTargetPageId="5" captionText="Fig. 2. SEM micrographs of male palps and RTA. AC. Cicynethus floriumfontis Jocqué, 1991, ♂ (NCA 91/1051). DE. C. mossambicus sp. nov., ♂ (RMCA_ARA_245567). FH. C. subtropicalis (Lawrence, 1952) comb. nov., ♂ (NCA 2008/4266). A. Male palp, ventro-retrolateral view. B. RTA apex, ventroretrolateral view. C. As preceding, retrolateral view. D. Male palp, ventro-retrolateral view. E. RTA apex, ventro-retrolateral view. F. Male palp, retrolateral view. G. As preceding, ventro-retrolateral view. H. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="4" pageNumber="5">Figs 2A</figureCitation>
, FG, 10AB, 18A, CD, F) or with retrolateral boss (
<figureCitation id="F1F82A2ED01BFFBEFB18F9D57E44357B" box="[1267,1339,1579,1605]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="4" pageNumber="5">Figs 3</figureCitation>
DF, 6AB), DTP reduced or inconspicuous; epigyne with transverse plate narrower and with longer or faint median septum........................................................................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D2372E0ED01BFFBEFF56F94B7E083426" pageId="4" pageNumber="5">
<paragraph id="697C36ABD01BFFBEFF56F94B7E0835CA" blockId="4.[189,1399,1717,1816]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF56F94B7E0835CA" pageId="4" pageNumber="5">
2. Entire body yellowish brown; base of setae with black spots throughout; legs spotted and with stripes on femora and patellae (
<figureCitation id="F1F82A2ED01BFFBEFDB1F92779E135CA" box="[602,670,1753,1780]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[189,1398,612,1887]" captionTargetId="figure@15.[189,1398,612,1887]" captionTargetPageId="15" captionText="Fig. 9. Cicynethus floriumfontis Jocqué, 1991. AC. ♂ (NCA 95/385). DF. ♀ (NCA 95/243). A. Habitus, dorsal view. B. As preceding, ventral view. C. Carapace, dorsal view. D. Habitus, dorsal view. E. As preceding, ventral view. F. Carapace, dorsal view. Scale bars: AB, DE = 2 mm; C, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3827042" httpUri="https://zenodo.org/record/3827042/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
AF) .......................................
<taxonomicName id="AEC34D28D01BFFBEFBFFF9247E0835CA" ID-CoL="VB65" authority="Jocque, 1991" authorityName="Jocque" authorityYear="1991" box="[1044,1399,1753,1780]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="floriumfontis">
<emphasis id="5BB7EAB9D01BFFBEFBFFF9247FAB35CA" bold="true" box="[1044,1236,1753,1780]" italics="true" pageId="4" pageNumber="5">C. floriumfontis</emphasis>
<bibRefCitation id="0D524B5AD01BFFBEFB37F9247E0835CA" author="Jocque R." box="[1244,1399,1754,1780]" pageId="4" pageNumber="5" pagination="1 - 160" refId="ref11880" refString="Jocque R. 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1 - 160." type="journal article" year="1991">Jocqué, 1991</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF2AF9007E083426" blockId="4.[189,1399,1717,1816]" box="[193,1399,1790,1816]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF2AF9007E083426" box="[193,1399,1790,1816]" pageId="4" pageNumber="5"> Differently coloured; base of setae without black spots; leg pattern different ................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="D2372E0ED01BFFBEFF56F8BE7E0834D5" pageId="4" pageNumber="5">
<paragraph id="697C36ABD01BFFBEFF56F8BE7E08349D" blockId="4.[189,1399,1856,2027]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF56F8BE7E08349D" pageId="4" pageNumber="5">
3. Abdomen dorsum with large anterior diamond-shaped spot, without white longitudinal stripe in front of spinnerets (
<figureCitation id="F1F82A2ED01BFFBEFD8FF89A79CE3440" box="[612,689,1892,1918]" captionStart="Fig" captionStartId="12.[189,232,1929,1955]" captionTargetBox="[189,1398,720,1876]" captionTargetId="figure@12.[189,1398,707,1889]" captionTargetPageId="12" captionText="Fig. 7. Cicynethus decoratus (Lawrence, 1952) comb. nov. A, D. holotype,♀. B, E, G. Female specimen (NCA2014/3298). C, F, H. Female specimen (RMCA_ARA_131806). AC. Habitus, dorsal view. DH. Epigyne, ventral view. Scale bars: AC = 2 mm; DH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827038" httpUri="https://zenodo.org/record/3827038/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
AC); femora (F) and tibiae (T) IIIV with black ring (
<figureCitation id="F1F82A2ED01BFFBEFF18F8767A32349C" box="[243,333,1928,1954]" captionStart="Fig" captionStartId="13.[189,232,1931,1957]" captionTargetBox="[191,1398,818,1874]" captionTargetId="figure@13.[189,1398,818,1886]" captionTargetPageId="13" captionText="Fig. 8. Cicynethus decoratus (Lawrence, 1952) comb. nov. AD. ♀ (RMCA_ARA_131806). E. Subadult ♂ (same sample). A. Habitus, ventral view. B. As preceding, lateral view. C. Prosoma, ventral view. D. Female palp, prolateral view. E. Habitus, dorsal view. Scale bars: AB, E = 2 mm; CD = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3827040" httpUri="https://zenodo.org/record/3827040/files/figure.png" pageId="4" pageNumber="5">Fig. 8B</figureCitation>
) ..........................................................................
<taxonomicName id="AEC34D28D01BFFBEFC95F8767F91349D" ID-CoL="VB64" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" box="[894,1262,1928,1955]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="decoratus" status="comb. nov.">
<emphasis id="5BB7EAB9D01BFFBEFC95F8767F69349D" bold="true" box="[894,1046,1928,1955]" italics="true" pageId="4" pageNumber="5">C. decoratus</emphasis>
(
<bibRefCitation id="0D524B5AD01BFFBEFBCEF8767F9A349D" author="Lawrence R. F." box="[1061,1253,1928,1955]" pageId="4" pageNumber="5" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D01BFFBEFB1EF8767E08349D" box="[1269,1399,1928,1955]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="697C36ABD01BFFBEFF2AF8537E0834D5" blockId="4.[189,1399,1856,2027]" pageId="4" pageNumber="5">
<keyLead id="D232959ED01BFFBEFF2AF8537E0834D5" pageId="4" pageNumber="5">
Abdomen dorsum with simple longitudinal median stripe ending in short white stretch in front of spinnerets (
<figureCitation id="F1F82A2ED01BFFBEFE9AF82F7AAD34D5" box="[369,466,2001,2027]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="4" pageNumber="5">Figs 3A</figureCitation>
,
<figureCitation id="F1F82A2ED01BFFBEFE34F82F7A9134D5" box="[479,494,2001,2027]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[199,1398,702,1898]" captionTargetId="figure@8.[189,1398,702,1898]" captionTargetPageId="8" captionText="Fig. 4. Cicynethus acer sp. nov., paratype, ♀ (TM 13375). A. Habitus, dorsal view. B. As preceding, closer view. C. As preceding, ventral view. D. As preceding, lateral view. E. Epigyne, ventral view. Scale bars: AD = 5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827032" httpUri="https://zenodo.org/record/3827032/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
AB, 5A, 15AG); F and T without ring ...................................................... 4
</keyLead>
</paragraph>
</keyStep>
<caption id="3DBC6623D01AFFBFFF56F8FE7A2034D3" ID-DOI="http://doi.org/10.5281/zenodo.3827028" ID-Zenodo-Dep="3827028" httpUri="https://zenodo.org/record/3827028/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1792,1818]" targetBox="[189,1398,265,1768]" targetPageId="5">
<paragraph id="697C36ABD01AFFBFFF56F8FE7A2034D3" blockId="5.[189,1399,1792,2029]" pageId="5" pageNumber="6">
<emphasis id="5BB7EAB9D01AFFBFFF56F8FE7A763424" bold="true" box="[189,265,1792,1818]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
SEM micrographs of male palps and RTA.
<emphasis id="5BB7EAB9D01AFFBFFCE9F8FE78443424" bold="true" box="[770,827,1792,1818]" pageId="5" pageNumber="6">AC</emphasis>
.
<taxonomicName id="AEC34D28D01AFFBFFCA3F8FE7E763424" ID-CoL="VB65" authority="Jocque, 1991" authorityName="Jocque" authorityYear="1991" box="[840,1289,1792,1818]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="floriumfontis">
<emphasis id="5BB7EAB9D01AFFBFFCA3F8FE7F183424" box="[840,1127,1792,1818]" italics="true" pageId="5" pageNumber="6">Cicynethus floriumfontis</emphasis>
<bibRefCitation id="0D524B5AD01AFFBFFB86F8FE7E763424" author="Jocque R." box="[1133,1289,1792,1818]" pageId="5" pageNumber="6" pagination="1 - 160" refId="ref11880" refString="Jocque R. 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1 - 160." type="journal article" year="1991">Jocqué, 1991</bibRefCitation>
</taxonomicName>
, ♂ (NCA 91/1051).
<emphasis id="5BB7EAB9D01AFFBFFEDEF8DD7A133403" bold="true" box="[309,364,1827,1853]" pageId="5" pageNumber="6">DE</emphasis>
.
<taxonomicName id="AEC34D28D01AFFBFFE91F8DD79483403" ID-CoL="VB66" authority="Jocqué &amp; Henrard, 2018" authorityName="Jocqué &amp; Henrard" authorityYear="2018" box="[378,567,1827,1853]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mossambicus" status="sp. nov.">
<emphasis id="5BB7EAB9D01AFFBFFE91F8DD79483403" box="[378,567,1827,1853]" italics="true" pageId="5" pageNumber="6">C. mossambicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="408457C2D01AFFBFFDD5F8DD79E53403" box="[574,666,1827,1853]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (RMCA_ARA_245567).
<emphasis id="5BB7EAB9D01AFFBFFC06F8DD7F5B3403" bold="true" box="[1005,1060,1827,1853]" pageId="5" pageNumber="6">FH</emphasis>
.
<taxonomicName id="AEC34D28D01AFFBFFBD9F8DD7A7E345F" ID-CoL="VB68" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="subtropicalis" status="comb. nov.">
<emphasis id="5BB7EAB9D01AFFBFFBD9F8DD7F943403" box="[1074,1259,1827,1853]" italics="true" pageId="5" pageNumber="6">C. subtropicalis</emphasis>
(
<bibRefCitation id="0D524B5AD01AFFBFFB11F8DD7B86345F" author="Lawrence R. F." pageId="5" pageNumber="6" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D01AFFBFFEE0F8B87AEF345F" box="[267,400,1862,1889]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
, ♂ (NCA 2008/4266).
<emphasis id="5BB7EAB9D01AFFBFFD4FF8B879C6345E" bold="true" box="[676,697,1862,1888]" pageId="5" pageNumber="6">A</emphasis>
. Male palp, ventro-retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFB93F8B87FF3345E" bold="true" box="[1144,1164,1862,1888]" pageId="5" pageNumber="6">B</emphasis>
. RTA apex, ventroretrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFE79F8977AD834BD" bold="true" box="[402,423,1897,1923]" pageId="5" pageNumber="6">C</emphasis>
. As preceding, retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFCD5F894782C34BA" bold="true" box="[830,851,1898,1924]" pageId="5" pageNumber="6">D</emphasis>
. Male palp, ventro-retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFAF7F8947E4F34BA" bold="true" box="[1308,1328,1898,1924]" pageId="5" pageNumber="6">E</emphasis>
. RTA apex, ventro-retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFDC8F873794A3499" bold="true" box="[547,565,1933,1959]" pageId="5" pageNumber="6">F</emphasis>
. Male palp, retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFC7AF87278D73498" bold="true" box="[913,936,1932,1958]" pageId="5" pageNumber="6">G</emphasis>
. As preceding, ventro-retrolateral view.
<emphasis id="5BB7EAB9D01AFFBFFF56F84E7BAB34F4" bold="true" box="[189,212,1968,1994]" pageId="5" pageNumber="6">H</emphasis>
. RTA apex, ventro-retrolateral view. Abbreviations: E = embolus; MA = median apophysis. Scale bars: 0.2 mm.
</paragraph>
</caption>
<keyStep id="D2372E0ED019FFBCFF56FEF57E0832C7" pageId="6" pageNumber="7">
<paragraph id="697C36ABD019FFBCFF56FEF57E083252" blockId="6.[189,1399,267,505]" pageId="6" pageNumber="7">
<keyLead id="D232959ED019FFBCFF56FEF57E083252" pageId="6" pageNumber="7">
4. Tip of RTA with sharp dorsal appendage (
<figureCitation id="F1F82A2ED019FFBCFD2CFEF57860321B" box="[711,799,267,293]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="6" pageNumber="7">Fig. 3F</figureCitation>
); tegulum not strongly swollen, but with retrolateral boss (
<figureCitation id="F1F82A2ED019FFBCFEDFFED07A033277" box="[308,380,302,329]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
DF); MA smoothly curved; embolus long, almost reaching distal tip of cymbium (
<figureCitation id="F1F82A2ED019FFBCFF18FEAC7A333252" box="[243,332,338,364]" captionStart="Fig" captionStartId="7.[189,232,1896,1922]" captionTargetBox="[199,1378,688,1854]" captionTargetId="figure@7.[189,1398,683,1854]" captionTargetPageId="7" captionText="Fig. 3. Cicynethus acer sp. nov., holotype, ♂ (TM 6162). A. Habitus, dorsal view. B. As preceding, ventral view. C. As preceding, lateral view. D. Palp, retrolateral view. E. As preceding, detail of RTA. F. Palp, ventral view. Abbreviations: E = embolus; MA = median apophysis; TS = tegular swelling. Scale bars: AC = 5 mm; D, F = 0.5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827030" httpUri="https://zenodo.org/record/3827030/files/figure.png" pageId="6" pageNumber="7">Fig. 3F</figureCitation>
); epigyne with well defined, Y-shaped median septum (
<figureCitation id="F1F82A2ED019FFBCFC3DFEAC7F4A3252" box="[982,1077,338,364]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[199,1398,702,1898]" captionTargetId="figure@8.[189,1398,702,1898]" captionTargetPageId="8" captionText="Fig. 4. Cicynethus acer sp. nov., paratype, ♀ (TM 13375). A. Habitus, dorsal view. B. As preceding, closer view. C. As preceding, ventral view. D. As preceding, lateral view. E. Epigyne, ventral view. Scale bars: AD = 5 mm; E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827032" httpUri="https://zenodo.org/record/3827032/files/figure.png" pageId="6" pageNumber="7">Figs 4E</figureCitation>
,
<figureCitation id="F1F82A2ED019FFBCFBAAFEAC7F1B3253" box="[1089,1124,338,365]" captionStart="Fig" captionStartId="10.[189,232,1929,1955]" captionTargetBox="[224,1359,856,1882]" captionTargetId="figure@10.[191,1396,818,1907]" captionTargetPageId="10" captionText="Fig. 6. Cicynethus acer sp. nov., genitalia drawings. A. Male palp, retrolateral view. B. As preceding, ventral view. C. Epigyne, ventral view. Abbreviations: E = embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; TS = tegular swelling. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827036" httpUri="https://zenodo.org/record/3827036/files/figure.png" pageId="6" pageNumber="7">6C</figureCitation>
) .........
<taxonomicName id="AEC34D28D019FFBCFB54FEAC7E6A3252" ID-CoL="VB63" authority="Jocqué &amp; Henrard, 2018" authorityName="Jocqué &amp; Henrard" authorityYear="2018" box="[1215,1301,338,364]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acer" status="sp. nov.">
<emphasis id="5BB7EAB9D019FFBCFB54FEAC7E6A3252" bold="true" box="[1215,1301,338,364]" italics="true" pageId="6" pageNumber="7">C. acer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="408457C2D019FFBCFAF7FEAC7E083252" box="[1308,1399,338,364]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="697C36ABD019FFBCFF2AFE8B7E0832C7" blockId="6.[189,1399,267,505]" pageId="6" pageNumber="7">
<keyLead id="D232959ED019FFBCFF2AFE8B7E0832C7" pageId="6" pageNumber="7">
Palp with dorsal excrescence of RTA tip flat, lamellar; tegulum strongly swollen and bulging (
<figureCitation id="F1F82A2ED019FFBCFF18FE667A1B328C" box="[243,356,408,434]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,269,1845]" captionTargetId="figure@27.[189,1398,269,1845]" captionTargetPageId="27" captionText="Fig. 18. Cicynethus subtropicalis (Lawrence, 1952) comb. nov., male palps. AC. NCA 2007/3035. DI. NCA 2016/842. A, D. Retrolateral views. B, E. Apex of RTA, retrolateral views. C, F. Ventral views. G. RTA, dorsal, slightly lateral view. H. Apex of RTA, frontal view. I. As preceding, ventral view. Scale bars: A, CD, F = 0.5 mm; B, E, GI = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827060" httpUri="https://zenodo.org/record/3827060/files/figure.png" pageId="6" pageNumber="7">Fig. 18A</figureCitation>
, CD, F); MA frontal margin with boss; embolus reaching distal margin of cymbial alveolus; epigyne with faint median septum (
<figureCitation id="F1F82A2ED019FFBCFD10FE45783232EB" box="[763,845,443,469]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[189,1398,907,1871]" captionTargetId="figure@26.[189,1398,907,1887]" captionTargetPageId="26" captionText="Fig. 17. Cicynethus subtropicalis (Lawrence, 1952) comb. nov., ♀, epigynes. A. Holotype (NM 1161). B. NCA 2005/1860. C, H. RMCA_ARA_211745. D, G. NCA 2007/3556. EF. NCA 2000/438. AE, GH = ventral views; F = dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3827058" httpUri="https://zenodo.org/record/3827058/files/figure.png" pageId="6" pageNumber="7">Fig. 17</figureCitation>
AH) ................................................................ ...................................................................................
<taxonomicName id="AEC34D28D019FFBCFCB2FE217F9132C7" ID-CoL="VB68" authority="(Lawrence, 1952)" authorityName="Jocqué &amp; Henrard" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1952" box="[857,1262,478,505]" class="Arachnida" family="Zodariidae" genus="Cicynethus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="subtropicalis" status="comb. nov.">
<emphasis id="5BB7EAB9D019FFBCFCB2FE217F6932C7" bold="true" box="[857,1046,478,505]" italics="true" pageId="6" pageNumber="7">C. subtropicalis</emphasis>
(
<bibRefCitation id="0D524B5AD019FFBCFBCEFE217F9A32C7" author="Lawrence R. F." box="[1061,1253,478,505]" pageId="6" pageNumber="7" pagination="183 - 226" refId="ref12014" refString="Lawrence R. F. 1952. New spiders from the eastern half of South Africa. Annals of the Natal Museum 12: 183 - 226." type="journal article" year="1952">Lawrence, 1952</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="408457C2D019FFBCFB1EFE207E0832C7" box="[1269,1399,478,505]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>