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<document id="957D0F2E35F47731F6DC6DA4C5858711" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA9AFEA9CBAFBD7FFEFAFD2F" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lissodrillia verrillii Dall 1881, new combination" docType="treatment" docVersion="13" lastPageNumber="258" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="256" updateTime="1698184378282" updateUser="plazi">
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<mods:title id="0C586DC3A8911BBB31C6E9F932FFDA08">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:name id="380DBEB63725426B4985901D226D568B" type="personal">
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<mods:namePart id="CD3BFE7278C6F7E58249C985131252E1">Fallon, Phillip J.</mods:namePart>
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<mods:typeOfResource id="ABF268890C953F947F89782AAC18352C">text</mods:typeOfResource>
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<mods:title id="5462BC91748C33F80B6F6307DFB96C22">Zootaxa</mods:title>
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<mods:part id="32D1926666488D2837F55254FA3642B4">
<mods:date id="8D081B2DB09A18884C7084D28D1B6C08">2016</mods:date>
<mods:detail id="40826B4D49AE35C1BC36A6569B2E4DBF" type="volume">
<mods:number id="608845CA2A0C0958B08E1BFBA0D44916">4090</mods:number>
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<mods:classification id="7318E02634EA975BBBC7C8488F0CADE0">journal article</mods:classification>
<mods:identifier id="0647B52D32F3F8E4EC4E0E56D9F3B833" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
<mods:identifier id="AFD19D4C46CDFAB73347BF15B3F5867E" type="GBIF-Dataset">e6b35f5a-435e-4473-b29e-1e4d842f84b0</mods:identifier>
<mods:identifier id="0E78B45E93B65C44B9924F22B5195848" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039F87C4FA9AFEA9CBAFBD7FFEFAFD2F" ID-DOI="http://doi.org/10.5281/zenodo.6076558" ID-GBIF-Taxon="117911608" ID-Zenodo-Dep="6076558" LSID="urn:lsid:plazi:treatment:039F87C4FA9AFEA9CBAFBD7FFEFAFD2F" httpUri="http://treatment.plazi.org/id/039F87C4FA9AFEA9CBAFBD7FFEFAFD2F" lastPageId="257" lastPageNumber="258" pageId="255" pageNumber="256">
<subSubSection id="C32C6559FA9AFF57CBAFBD7FFCFBFB4E" pageId="255" pageNumber="256" type="nomenclature">
<paragraph id="8B8936D2FA9AFF57CBAFBD7FFD16FBD4" blockId="255.[151,766,1124,1183]" box="[151,766,1124,1150]" pageId="255" pageNumber="256">
<heading id="D0C181BEFA9AFF57CBAFBD7FFD16FBD4" bold="true" box="[151,766,1124,1150]" fontSize="11" level="1" pageId="255" pageNumber="256" reason="1">
<emphasis id="B942EAC0FA9AFF57CBAFBD7FFD16FBD4" bold="true" box="[151,766,1124,1150]" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CBAFBD7FFDF2FBD4" ID-CoL="3VFFN" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[151,538,1124,1150]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii" status="comb. nov.">
<emphasis id="B942EAC0FA9AFF57CBAFBD7FFE69FBD4" bold="true" box="[151,385,1124,1150]" italics="true" pageId="255" pageNumber="256">Lissodrillia verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA9AFF57C91FBD7FFD16FBD4" box="[551,766,1124,1150]" pageId="255" pageNumber="256">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936D2FA9AFF57CBA6BD9DFEF2FB35" blockId="255.[151,766,1124,1183]" box="[158,282,1158,1183]" pageId="255" pageNumber="256">(Plate 130)</paragraph>
<paragraph id="8B8936D2FA9AFF57CBAFBDD5FCFBFB4E" blockId="255.[151,1329,1230,1406]" box="[151,787,1230,1252]" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CBAFBDD5FDB1FB4E" authority="Verrillii Dall, 1881: 68" authorityName="Verrillii Dall" authorityPageNumber="68" authorityYear="1881" box="[151,601,1230,1252]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="subGenus" subGenus="Drillia">
<emphasis id="B942EAC0FA9AFF57CBAFBDD5FE57FB4E" box="[151,447,1230,1252]" italics="true" pageId="255" pageNumber="256">Pleurotoma (Drillia) Verrillii</emphasis>
Dall, 1881: 68
</taxonomicName>
. Dall (1885: 239).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA9AFF57CBAFBDF6FB4EFAD4" pageId="255" pageNumber="256" type="description">
<paragraph id="8B8936D2FA9AFF57CBAFBDF6FD54FA88" blockId="255.[151,1329,1230,1406]" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CBAFBDF6FE5EFAA9" authority="Verrillii (Dall, 1881)" authorityName="Verrillii (Dall" authorityYear="1881" box="[151,438,1261,1283]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA9AFF57CBAFBDF6FED9FAA9" box="[151,305,1261,1283]" italics="true" pageId="255" pageNumber="256">Drillia Verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
: Tryon (1884: 213 [list]); Dall (1889a: 93, pl. 11, fig. 2); Dall (1889b: 98, pl. 11, fig. 2).
<taxonomicName id="4C364D51FA9AFF57CBAFBC17FE5DFA88" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[151,437,1292,1314]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FA9AFF57CBAFBC17FEC7FA88" box="[151,303,1292,1314]" italics="true" pageId="255" pageNumber="256">Drillia verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
: Maury (1922: 151 [list]).
</paragraph>
<paragraph id="8B8936D2FA9AFF57CBAFBC31FCBAFAF5" blockId="255.[151,1329,1230,1406]" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CBAFBC31FE34FAEA" box="[151,476,1322,1344]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrilli" subGenus="Cerodrillia">
<emphasis id="B942EAC0FA9AFF57CBAFBC31FE34FAEA" box="[151,476,1322,1344]" italics="true" pageId="255" pageNumber="256">Cerodrillia (Cerodrillia) verrilli</emphasis>
</taxonomicName>
[sic] (Dall, 1881): Abbott (1974: 272, spec. &amp; fig. 3030).
<taxonomicName id="4C364D51FA9AFF57CBAFBC52FEBCFAF5" box="[151,340,1353,1375]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrilli">
<emphasis id="B942EAC0FA9AFF57CBAFBC52FEBCFAF5" box="[151,340,1353,1375]" italics="true" pageId="255" pageNumber="256">Cerodrillia verrilli</emphasis>
</taxonomicName>
[sic] (Dall, 1881): Turgeon
<emphasis id="B942EAC0FA9AFF57C94ABC51FD4DFAF5" box="[626,677,1353,1375]" italics="true" pageId="255" pageNumber="256">et al.</emphasis>
(1988: 96 [list]).
</paragraph>
<paragraph id="8B8936D2FA9AFF57CBAFBC73FB4EFAD4" blockId="255.[151,1329,1230,1406]" box="[151,1190,1384,1406]" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CBAFBC73FE08FAD4" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[151,480,1384,1406]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FA9AFF57CBAFBC73FEB3FAD4" box="[151,347,1384,1406]" italics="true" pageId="255" pageNumber="256">Cerodrillia verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
: Turgeon
<emphasis id="B942EAC0FA9AFF57C97DBC72FD91FAD4" box="[581,633,1384,1406]" italics="true" pageId="255" pageNumber="256">et al.</emphasis>
(1998: 102 [list]); Williams (2005; 2006; 2009: 1534).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA9AFEA8CBAFBCB0FB19FF7F" lastPageId="256" lastPageNumber="257" pageId="255" pageNumber="256" type="materials_examined">
<paragraph id="8B8936D2FA9AFF57CBAFBCB0FC75F931" blockId="255.[151,1436,1450,1691]" pageId="255" pageNumber="256">
<emphasis id="B942EAC0FA9AFF57CBAFBCB0FEACFA69" bold="true" box="[151,324,1450,1475]" pageId="255" pageNumber="256">
<typeStatus id="548D8870FA9AFF57CBAFBCB0FF3AFA68" box="[151,210,1451,1474]" pageId="255" pageNumber="256">Type</typeStatus>
material.
</emphasis>
Dall (1881: 68) mentioned only a single 5.5 x
<quantity id="4CCE9B37FA9AFF57C876BCB0FC59FA68" box="[846,945,1451,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="255" pageNumber="256" unit="mm" value="1.75">1.75 mm</quantity>
specimen from R/V
<emphasis id="B942EAC0FA9AFF57CFA2BCB1FB32FA69" box="[1178,1242,1450,1475]" italics="true" pageId="255" pageNumber="256">Blake</emphasis>
Sta. 19 off Bahia Honda,
<collectingCountry id="F3217642FA9AFF57CBCABCD5FED8FA4D" box="[242,304,1486,1511]" name="Cuba" pageId="255" pageNumber="256">Cuba</collectingCountry>
, in his description. Later, he identified a lot from R/V
<emphasis id="B942EAC0FA9AFF57C89EBCD5FC0EFA4D" box="[934,998,1486,1511]" italics="true" pageId="255" pageNumber="256">Blake</emphasis>
Sta. 20, also off Bahia Honda, as this species (Dall, 1889a: 93). Since lots were sometimes split between the Smithsonian and Museum of Comparative Zoology, both museums were searched for
<emphasis id="B942EAC0FA9AFF57C941BF0DFD51F985" box="[633,697,1558,1583]" italics="true" pageId="255" pageNumber="256">Blake</emphasis>
specimens from both stations. An on-line database search yielded 3 lots at the MCZ (MCZ 7083, 72401 &amp; 7084), and a drawer search yielded a single lot at the Smithsonian (USNM 87457), all of which were collected at either locality and identified as this species. The following table is a summary of the known
<typeStatus id="548D8870FA9AFF57CA99BF98FDEBF930" box="[417,515,1667,1690]" pageId="255" pageNumber="256" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C364D51FA9AFF57C910BF99FC71F931" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[552,921,1666,1691]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FA9AFF57C910BF99FCEDF931" box="[552,773,1666,1691]" italics="true" pageId="255" pageNumber="256">Lissodrillia verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
:
</paragraph>
<paragraph id="8B8936D2FA9AFF57CBA7BFADFB25F83A" pageId="255" pageNumber="256">
<table id="F936C472FA9A0057CBA7BFADFACFF83A" box="[159,1319,1718,1936]" gridcols="5" gridrows="5" pageId="255" pageNumber="256">
<tr id="35063490FA9A0057CBA7BFADFACFF941" box="[159,1319,1718,1771]" gridrow="0" pageId="255" pageNumber="256">
<th id="76D75DECFA9A0057CBA7BFADFF0EF941" box="[159,230,1718,1771]" gridcol="0" gridrow="0" pageId="255" pageNumber="256">Blake Station</th>
<th id="76D75DECFA9A0057CA24BFADFE78F941" box="[284,400,1718,1771]" gridcol="1" gridrow="0" pageId="255" pageNumber="256">Depth(fms) [meters]</th>
<th id="76D75DECFA9A0057CAE8BFADFD91F941" box="[464,633,1718,1771]" gridcol="2" gridrow="0" pageId="255" pageNumber="256">Locality</th>
<th id="76D75DECFA9A0057C9DCBFADFC7FF941" box="[740,919,1718,1771]" gridcol="3" gridrow="0" pageId="255" pageNumber="256">Catalog number (count)</th>
<th id="76D75DECFA9A0057CF39BFADFACFF941" box="[1025,1319,1718,1771]" gridcol="4" gridrow="0" pageId="255" pageNumber="256">Confirmed Identification (length in mm)</th>
</tr>
<tr id="35063490FA9A0057CBA7BFE5FACFF8BE" box="[159,1319,1790,1812]" gridrow="1" pageId="255" pageNumber="256">
<th id="76D75DECFA9A0057CBA7BFE5FF0EF8BE" box="[159,230,1790,1812]" gridcol="0" gridrow="1" pageId="255" pageNumber="256">19</th>
<td id="76D75DECFA9A0057CA24BFE5FE78F8BE" box="[284,400,1790,1812]" gridcol="1" gridrow="1" pageId="255" pageNumber="256">310 [567]</td>
<td id="76D75DECFA9A0057CAE8BFE5FD91F8BE" box="[464,633,1790,1812]" gridcol="2" gridrow="1" pageId="255" pageNumber="256">off Bahia Honda</td>
<td id="76D75DECFA9A0057C9DCBFE5FC7FF8BE" box="[740,919,1790,1812]" gridcol="3" gridrow="1" pageId="255" pageNumber="256">MCZ 7083 (1)</td>
<td id="76D75DECFA9A0057CF39BFE5FACFF8BE" box="[1025,1319,1790,1812]" gridcol="4" gridrow="1" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CF39BFE4FB80F8BE" box="[1025,1128,1790,1812]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FA9AFF57CF39BFE4FB80F8BE" box="[1025,1128,1790,1812]" italics="true" pageId="255" pageNumber="256">L. verrillii</emphasis>
</taxonomicName>
(4.4 mm)
</td>
</tr>
<tr id="35063490FA9A0057CBA7BE33FACFF894" box="[159,1319,1832,1854]" gridrow="2" pageId="255" pageNumber="256">
<th id="76D75DECFA9A0057CBA7BE33FF0EF894" box="[159,230,1832,1854]" gridcol="0" gridrow="2" pageId="255" pageNumber="256">19</th>
<td id="76D75DECFA9A0057CA24BE33FE78F894" box="[284,400,1832,1854]" gridcol="1" gridrow="2" pageId="255" pageNumber="256">310 [567]</td>
<td id="76D75DECFA9A0057CAE8BE33FD91F894" box="[464,633,1832,1854]" gridcol="2" gridrow="2" pageId="255" pageNumber="256">off Bahia Honda</td>
<td id="76D75DECFA9A0057C9DCBE33FC7FF894" box="[740,919,1832,1854]" gridcol="3" gridrow="2" pageId="255" pageNumber="256">MCZ 72401 (2)</td>
<td id="76D75DECFA9A0057CF39BE33FACFF894" box="[1025,1319,1832,1854]" gridcol="4" gridrow="2" pageId="255" pageNumber="256">not examined</td>
</tr>
<tr id="35063490FA9A0057CBA7BE4AFACFF8CD" box="[159,1319,1873,1895]" gridrow="3" pageId="255" pageNumber="256">
<th id="76D75DECFA9A0057CBA7BE4AFF0EF8CD" box="[159,230,1873,1895]" gridcol="0" gridrow="3" pageId="255" pageNumber="256">20</th>
<td id="76D75DECFA9A0057CA24BE4AFE78F8CD" box="[284,400,1873,1895]" gridcol="1" gridrow="3" pageId="255" pageNumber="256">220 [402]</td>
<td id="76D75DECFA9A0057CAE8BE4AFD91F8CD" box="[464,633,1873,1895]" gridcol="2" gridrow="3" pageId="255" pageNumber="256">off Bahia Honda</td>
<td id="76D75DECFA9A0057C9DCBE4AFC7FF8CD" box="[740,919,1873,1895]" gridcol="3" gridrow="3" pageId="255" pageNumber="256">MCZ 7084 (1)</td>
<td id="76D75DECFA9A0057CF39BE4AFACFF8CD" box="[1025,1319,1873,1895]" gridcol="4" gridrow="3" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CF39BE4AFAECF8CD" authority="(Dall, 1881)" authorityName="Dall" authorityYear="1881" baseAuthorityName="Dall" baseAuthorityYear="1881" box="[1025,1284,1873,1895]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FA9AFF57CF39BE4AFB6BF8CD" box="[1025,1155,1873,1895]" italics="true" pageId="255" pageNumber="256">S. lissotropis</emphasis>
(Dall, 1881)
</taxonomicName>
</td>
</tr>
<tr id="35063490FA9A0057CBA7BE61FACFF83A" box="[159,1319,1914,1936]" gridrow="4" pageId="255" pageNumber="256">
<th id="76D75DECFA9A0057CBA7BE61FF0EF83A" box="[159,230,1914,1936]" gridcol="0" gridrow="4" pageId="255" pageNumber="256">20</th>
<td id="76D75DECFA9A0057CA24BE61FE78F83A" box="[284,400,1914,1936]" gridcol="1" gridrow="4" pageId="255" pageNumber="256">220 [402]</td>
<td id="76D75DECFA9A0057CAE8BE61FD91F83A" box="[464,633,1914,1936]" gridcol="2" gridrow="4" pageId="255" pageNumber="256">off Bahia Honda</td>
<td id="76D75DECFA9A0057C9DCBE61FC7FF83A" box="[740,919,1914,1936]" gridcol="3" gridrow="4" pageId="255" pageNumber="256">USNM 87457 (1)</td>
<td id="76D75DECFA9A0057CF39BE61FACFF83A" box="[1025,1319,1914,1936]" gridcol="4" gridrow="4" pageId="255" pageNumber="256">
<taxonomicName id="4C364D51FA9AFF57CF39BE60FB80F83A" box="[1025,1128,1914,1936]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="255" pageNumber="256" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FA9AFF57CF39BE60FB80F83A" box="[1025,1128,1914,1936]" italics="true" pageId="255" pageNumber="256">L. verrillii</emphasis>
</taxonomicName>
(5.1 mm)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFB983FB19FF7F" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">
MCZ 7083: out on loan but a length measurement and photograph of the specimen in this lot was provided by Peter Stahlschmidt. It agrees with Dall’s description but is smaller, 4.4, not
<quantity id="4CCE9B37FB65FEA8C8DBB9A6FBD1FF7E" box="[995,1081,189,212]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="256" pageNumber="257" unit="mm" value="5.5">5.5 mm</quantity>
(Pl. 130
<figureCitation id="130D2A57FB65FEA8CF98B9A7FB37FF7F" box="[1184,1247,188,213]" captionStart="Fig. 1" captionStartId="66.[151,188,1159,1181]" captionTargetBox="[151,1436,375,1146]" captionTargetId="figure@66.[151,1436,375,1146]" captionTargetPageId="66" captionText="Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip." httpUri="https://zenodo.org/record/263325/files/figure.png" pageId="256" pageNumber="257">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB65FEA8CBFFB9FBFCC0FD02" pageId="256" pageNumber="257" type="description">
<paragraph id="8B8936D2FB65FEA8CBFFB9FBFC8CFEEA" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">MCZ 72401: also from R/V Blake Station 19, was out on loan and could not be examined. The on-line database lists two specimens in this lot. It is believed they were originally in MCZ 7083, which was recorded as having three specimens in the MCZ register, but later separated.</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFB857FAF9FE06" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">
MCZ 7084: out on loan but a photograph (no measurement) was provided by John Tucker. The specimen in the photo does not match Dall’s description of
<taxonomicName id="4C364D51FB65FEA8C9BFB86BFD14FE22" box="[647,764,367,392]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="256" pageNumber="257" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FB65FEA8C9BFB86BFD14FE22" box="[647,764,367,392]" italics="true" pageId="256" pageNumber="257">L. verrillii</emphasis>
</taxonomicName>
but is instead
<taxonomicName id="4C364D51FB65FEA8C89EB874FA88FE22" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[934,1376,367,392]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="256" pageNumber="257" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB65FEA8C89EB874FB22FE22" box="[934,1226,367,392]" italics="true" pageId="256" pageNumber="257">Syntomodrillia lissotropis</emphasis>
(Dall, 1881)
</taxonomicName>
. R/V
<emphasis id="B942EAC0FB65FEA8CBAFB88FFF3FFE07" box="[151,215,404,429]" italics="true" pageId="256" pageNumber="257">Blake</emphasis>
Station 20 is the
<typeStatus id="548D8870FB65FEA8CAA1B88EFE21FE06" box="[409,457,405,428]" pageId="256" pageNumber="257">type</typeStatus>
locality of
<taxonomicName id="4C364D51FB65FEA8C972B88FFD32FE07" box="[586,730,404,429]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="256" pageNumber="257" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB65FEA8C972B88FFD32FE07" box="[586,730,404,429]" italics="true" pageId="256" pageNumber="257">S. lissotropis</emphasis>
</taxonomicName>
. This specimen is a
<typeStatus id="548D8870FB65FEA8C886B88FFBBDFE07" box="[958,1109,404,429]" pageId="256" pageNumber="257" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C364D51FB65FEA8CF42B88FFAE2FE07" box="[1146,1290,404,429]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="256" pageNumber="257" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB65FEA8CF42B88FFAE2FE07" box="[1146,1290,404,429]" italics="true" pageId="256" pageNumber="257">S. lissotropis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFB8A3FEB8FE5E" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">
USNM 87457: was examined directly and matches Dall’s description (Pl. 130,
<figureCitation id="130D2A57FB65FEA8CF03B8ACFB96FE7A" box="[1083,1150,439,464]" captionStart="Fig. 2" captionStartId="210.[151,188,1695,1717]" captionTargetBox="[305,1256,193,1592]" captionTargetId="figure@210.[305,1257,193,1592]" captionText="Fig. 2. Holotype, 18.2 x 7.3 mm, F. alba, new species (USNM 1291342). Fig. 3. Paratype, 17.3 x 7.6 mm, F. chaaci (Espinosa &amp; Rolán, 1995) (MNHN 2857). Fig. 4. Holotype, 15.6 x 6.0 mm, F. crocea, new species (UF 155640). Fig. 5. Holotype, 15.3 x 5.4 mm, F. contracta, new species (UF 170316). Fig. 6. Holotype, 13.2 x 5.1 mm, F. petiti Tippett, 1995 (USNM 880073). Fig. 7. Holotype, 27.5 x 11.0 mm, F. phasma (Schwengel, 1940) (ANSP 176453). Fig. 8. Neotype, 25.5 x 9.5 mm, F. fucata (Reeve, 1845) (USNM 1291346). Fig. 9. Holotype, 25.0 x 7.4 mm, F. elongata, new species (UF 155626). Fig. 10. Holotype, 21.2 x 7.4 mm, F. tippetti, new species (UF 155639). Fig. 11. Holotype, 16.8 x 7.4 mm, F. marmarina (Watson, 1881) (NHMUK 1887.2.9.103). Fig. 12. Lectotype, 17.8 x 5.8 mm, F. pagodula (Dall, 1889) (USNM 87471)." pageId="256" pageNumber="257">Fig. 2</figureCitation>
), except that it is smaller, 5.1, not
<quantity id="4CCE9B37FB65FEA8CBCBB8C6FEA4FE5E" box="[243,332,477,500]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="256" pageNumber="257" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFBB1BFCC0FD02" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">
None of the examined specimens exactly match the
<quantity id="4CCE9B37FB65FEA8C81CBB1BFC95FDBD" box="[804,893,512,535]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="256" pageNumber="257" unit="mm" value="5.5">5.5 mm</quantity>
length described by Dall. The drawing in Dall (1889a: pl. 11, fig. 2) is very poor, but as explained by Dall, “These shells are so very small and polished that it is extremely difficult for an artist in pure line-work to represent them adequately.” (Dall, 1889a: 92). Since the length measurements of the examined shells do not agree precisely with Dall’s measurement, and inasmuch as all lots were not examined, no
<typeStatus id="548D8870FB65FEA8CAA5BB94FDECFD02" box="[413,516,655,680]" pageId="256" pageNumber="257" type="lectotype">lectotype</typeStatus>
designation is made here.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB65FEA8CBFFBBAEFC60F8AB" pageId="256" pageNumber="257" type="materials_examined">
<paragraph id="8B8936D2FB65FEA8CBFFBBAEFD6AFD5A" blockId="256.[151,1437,151,752]" pageId="256" pageNumber="257">
<emphasis id="B942EAC0FB65FEA8CBFFBBAEFE8FFD67" bold="true" box="[199,359,692,717]" pageId="256" pageNumber="257">
<typeStatus id="548D8870FB65FEA8CBFFBBAEFEEAFD66" box="[199,258,693,716]" pageId="256" pageNumber="257">Type</typeStatus>
locality.
</emphasis>
R/V
<emphasis id="B942EAC0FB65FEA8CA9FBBAFFE0FFD67" box="[423,487,692,717]" italics="true" pageId="256" pageNumber="257">Blake</emphasis>
Station 19, off Bahia Honda, [Artemisa Prov., formerly Pinar del Rio Prov.]
<collectingCountry id="F3217642FB65FEA8CE62BBAFFA70FD67" box="[1370,1432,692,717]" name="Cuba" pageId="256" pageNumber="257">Cuba</collectingCountry>
,
<geoCoordinate id="EE025015FB65FEA8CBAFBBC3FEE9FD5A" box="[151,257,728,752]" direction="north" orientation="latitude" pageId="256" pageNumber="257" precision="15" value="23.05">23°3'0&quot;N</geoCoordinate>
,
<geoCoordinate id="EE025015FB65FEA8CA35BBC3FE7EFD5A" box="[269,406,728,752]" direction="west" orientation="longitude" pageId="256" pageNumber="257" precision="15" value="-83.174995">83°10'30&quot;W</geoCoordinate>
, in 310 fms [
<quantity id="4CCE9B37FB65FEA8C910BBC3FD9EFD5A" box="[552,630,728,752]" metricMagnitude="2" metricUnit="m" metricValue="5.67" pageId="256" pageNumber="257" unit="m" value="567.0">567 m</quantity>
].
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBAFBF39FED5F9DF" pageId="256" pageNumber="257">
<table id="F936C472FB650057CBAFBF39FA74F9DF" box="[151,1436,1570,1653]" gridcols="3" gridrows="3" pageId="256" pageNumber="257">
<tr id="35063490FB650057CBAFBF39FA74F992" box="[151,1436,1570,1592]" gridrow="0" pageId="256" pageNumber="257">
<th id="76D75DECFB650057CBAFBF39FD02F992" box="[151,746,1570,1592]" gridcol="0" gridrow="0" pageId="256" pageNumber="257">
<emphasis id="B942EAC0FB65FEA8CBAFBF39FEF7F992" bold="true" box="[151,287,1570,1592]" pageId="256" pageNumber="257">PLATE 130.</emphasis>
<taxonomicName id="4C364D51FB65FEA8CA11BF39FD97F992" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[297,639,1570,1592]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="256" pageNumber="257" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FB65FEA8CA11BF39FE76F992" box="[297,414,1570,1592]" italics="true" pageId="256" pageNumber="257">Lissodrillia</emphasis>
<emphasis id="B942EAC0FB65FEA8CA9FBF39FE1CF992" box="[423,500,1570,1592]" italics="true" pageId="256" pageNumber="257">verrillii</emphasis>
(Dall, 1881)
</taxonomicName>
. Syntypes
</th>
<th id="76D75DECFB650057C9C9BF39FB43F992" box="[753,1195,1570,1592]" gridcol="1" gridrow="0" pageId="256" pageNumber="257">from off Bahia Honda, Cuba. Fig. 1: R/V</th>
<th id="76D75DECFB650057CF8BBF39FA74F992" box="[1203,1436,1570,1592]" gridcol="2" gridrow="0" pageId="256" pageNumber="257">
<emphasis id="B942EAC0FB65FEA8CF8BBF39FB05F992" box="[1203,1261,1570,1592]" italics="true" pageId="256" pageNumber="257">Blake</emphasis>
Sta. 19, ventral,
</th>
</tr>
<tr id="35063490FB650057CBAFBF5BFA74F9FC" box="[151,1436,1600,1622]" gridrow="1" pageId="256" pageNumber="257">
<th id="76D75DECFB650057CBAFBF5BFD02F9FC" box="[151,746,1600,1622]" gridcol="0" gridrow="1" pageId="256" pageNumber="257">lateral &amp; dorsal views (MCZ 7083) (Photographs courtesy</th>
<td id="76D75DECFB650057C9C9BF5BFB43F9FC" box="[753,1195,1600,1622]" gridcol="1" gridrow="1" pageId="256" pageNumber="257">
P. Stahlschmidt); Fig. 2: R/V
<emphasis id="B942EAC0FB65FEA8CF26BF5BFBB0F9FC" box="[1054,1112,1600,1622]" italics="true" pageId="256" pageNumber="257">Blake</emphasis>
Sta. 20,
</td>
<td id="76D75DECFB650057CF8BBF5BFA74F9FC" box="[1203,1436,1600,1622]" gridcol="2" gridrow="1" pageId="256" pageNumber="257">ventral &amp; lateral views</td>
</tr>
<tr id="35063490FB650057CBAFBF44FA74F9DF" box="[151,1436,1631,1653]" gridrow="2" pageId="256" pageNumber="257" rowspan-1="1" rowspan-2="1">
<th id="76D75DECFB650057CBAFBF44FD02F9DF" box="[151,746,1631,1653]" gridcol="0" gridrow="2" pageId="256" pageNumber="257">(USNM 87457).</th>
</tr>
</table>
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFBFBAFDE3F974" blockId="256.[151,1436,1697,2010]" pageId="256" pageNumber="257">
Information on
<typeStatus id="548D8870FB65FEA8CA46BFB9FE08F913" box="[382,480,1698,1721]" pageId="256" pageNumber="257" type="syntype">syntypes</typeStatus>
from
<emphasis id="B942EAC0FB65FEA8C914BFBAFD84F910" box="[556,620,1697,1722]" italics="true" pageId="256" pageNumber="257">Blake</emphasis>
Station 20 is: off Bahia Honda, [Artemisa Prov.]
<collectingCountry id="F3217642FB65FEA8CF81BFBAFB1FF910" box="[1209,1271,1697,1722]" name="Cuba" pageId="256" pageNumber="257">Cuba</collectingCountry>
,
<geoCoordinate id="EE025015FB65FEA8CE3EBFB9FA7FF913" box="[1286,1431,1698,1722]" direction="north" orientation="latitude" pageId="256" pageNumber="257" precision="15" value="23.041666">23°02'30&quot; N</geoCoordinate>
,
<geoCoordinate id="EE025015FB65FEA8CBAFBFDDFEF7F974" box="[151,287,1734,1758]" direction="west" orientation="longitude" pageId="256" pageNumber="257" precision="15" value="-83.183334">83°11'00&quot;W</geoCoordinate>
, in 220 fms [
<quantity id="4CCE9B37FB65FEA8CA89BFDDFDE8F974" box="[433,512,1734,1758]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="256" pageNumber="257" unit="m" value="402.0">402 m</quantity>
].
</paragraph>
<paragraph id="8B8936D2FB65FEA8CBFFBFF1FC60F8AB" blockId="256.[151,1436,1697,2010]" box="[199,904,1769,1794]" pageId="256" pageNumber="257">
<emphasis id="B942EAC0FB65FEA8CBFFBFF1FE43F8A8" bold="true" box="[199,427,1769,1794]" pageId="256" pageNumber="257">Range and habitat.</emphasis>
<collectingCountry id="F3217642FB65FEA8CA8ABFF2FE05F8A8" box="[434,493,1769,1794]" name="Cuba" pageId="256" pageNumber="257">Cuba</collectingCountry>
[Artemisa Prov.], in 402 and
<specimenCount id="9D30FD5BFB65FEA8C801BFF1FC60F8AB" box="[825,904,1770,1794]" pageId="256" pageNumber="257" type="generic">
<quantity id="4CCE9B37FB65FEA8C801BFF1FC6BF8AB" box="[825,899,1770,1794]" metricMagnitude="2" metricUnit="m" metricValue="5.67" pageId="256" pageNumber="257" unit="m" value="567.0">567 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB65FEA9CBFFBE16FAE7FECF" lastPageId="257" lastPageNumber="258" pageId="256" pageNumber="257" type="description">
<paragraph id="8B8936D2FB65FEA9CBFFBE16FAE7FECF" blockId="256.[151,1436,1697,2010]" lastBlockId="257.[151,1437,151,645]" lastPageId="257" lastPageNumber="258" pageId="256" pageNumber="257">
<emphasis id="B942EAC0FB65FEA8CBFFBE16FE77F88C" bold="true" box="[199,415,1805,1830]" pageId="256" pageNumber="257">
Description.
<emphasis id="B942EAC0FB65FEA8CA5CBE16FE77F88C" bold="true" box="[356,415,1805,1830]" italics="true" pageId="256" pageNumber="257">Shell</emphasis>
</emphasis>
very small (to
<quantity id="4CCE9B37FB65FEA8C963BE14FD5EF88C" box="[603,694,1807,1830]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="256" pageNumber="257" unit="mm" value="5.5">5.5 mm</quantity>
in total length), fusiform, glossy smooth, of about 6½ whorls, teleoconch whorls angular at their peripheries; the last whorl about 55% of total shell length. Sculpture of ribs; aperture narrow, anterior canal moderately long.
<emphasis id="B942EAC0FB65FEA8C984BE4EFCAAF8C4" bold="true" box="[700,834,1877,1902]" italics="true" pageId="256" pageNumber="257">Protoconch</emphasis>
of 1½ smooth round inflated whorls.
<emphasis id="B942EAC0FB65FEA8CFD3BE4EFA74F8C4" bold="true" box="[1259,1436,1877,1902]" italics="true" pageId="256" pageNumber="257">Axial sculpture</emphasis>
of ribs, obsolete to nearly obsolete on shoulder near suture, strong at periphery and to suture of succeeding whorls, and to about ⅔ the distance to anterior canal on last whorl; ribs number 12 on penultimate and 5 to varix on last whorl. Ribs angular at periphery, about mid-whorl. Growth striae microscopic.
<emphasis id="B942EAC0FB65FEA8CF24BEDAFBB0F870" bold="true" box="[1052,1112,1985,2010]" italics="true" pageId="256" pageNumber="257">Varix</emphasis>
a thickened rib about ⅓–½- turn from edge of outer lip.
<emphasis id="B942EAC0FB64FEA9CAF3B98CFD6BFF1A" bold="true" box="[459,643,151,176]" italics="true" pageId="257" pageNumber="258">Spiral sculpture</emphasis>
absent.
<emphasis id="B942EAC0FB64FEA9C9E6B98CFCC4FF1A" bold="true" box="[734,812,151,176]" italics="true" pageId="257" pageNumber="258">Sulcus</emphasis>
marked by the absence or near absence of ribs on whorl shoulder.
<emphasis id="B942EAC0FB64FEA9CA3BB9A7FE82FF7F" bold="true" box="[259,362,188,213]" italics="true" pageId="257" pageNumber="258">Outer lip</emphasis>
thin, without axial folds, smooth, edge including anal sinus forms an uninterrupted reversed “S” curve from suture to stromboid notch, which is weak.
<emphasis id="B942EAC0FB64FEA9C82EB9C4FC7EFF52" bold="true" box="[790,918,223,248]" italics="true" pageId="257" pageNumber="258">Anal sinus</emphasis>
a deep notch on whorl shoulder adjacent to suture, its outer edge divergent, such that its edge in congruent with that of the outer lip. Inner edge of sinus flared out, directed laterally, spout-like.
<emphasis id="B942EAC0FB64FEA9C92EB833FD95FEEA" bold="true" box="[534,637,295,320]" italics="true" pageId="257" pageNumber="258">Inner lip</emphasis>
thin, margined, extends roof-like over apex of aperture, callus at anal sinus weak.
<emphasis id="B942EAC0FB64FEA9CA18B857FE21FECF" bold="true" box="[288,457,332,357]" italics="true" pageId="257" pageNumber="258">Anterior canal</emphasis>
moderately long for the genus, open, unnotched.
<emphasis id="B942EAC0FB64FEA9C8CDB857FBDEFECF" bold="true" box="[1013,1078,332,357]" italics="true" pageId="257" pageNumber="258">Color</emphasis>
white to off-white.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB64FEA9CBFFB86BFEFAFD2F" pageId="257" pageNumber="258" type="discussion">
<paragraph id="8B8936D2FB64FEA9CBFFB86BFEFAFD2F" blockId="257.[151,1437,151,645]" pageId="257" pageNumber="258">
<emphasis id="B942EAC0FB64FEA9CBFFB86BFE2AFE2D" bold="true" box="[199,450,368,391]" pageId="257" pageNumber="258">
Remarks.
<emphasis id="B942EAC0FB64FEA9CA7BB86BFE2AFE2D" box="[323,450,368,391]" italics="true" pageId="257" pageNumber="258">Taxonomy.</emphasis>
</emphasis>
<taxonomicName id="4C364D51FB64FEA9CAF2B874FD41FE22" box="[458,681,367,392]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="verrillii">Lissodrillia verrillii</taxonomicName>
has the characteristics of the genus: a very small glossy shell that lacks spiral sculpture, and with inner edge of anal sinus and edge of outer lip that trace a reverse “S”. Unlike its congeners, its ribs are weak to absent in the sulcus, and the inflated spire whorls are angular at their peripheries.
<emphasis id="B942EAC0FB64FEA9CBAFB8C7FDC8FE5F" box="[151,544,476,501]" italics="true" pageId="257" pageNumber="258">
<emphasis id="B942EAC0FB64FEA9CBAFB8C7FED3FE5F" bold="true" box="[151,315,476,501]" italics="true" pageId="257" pageNumber="258">Identification.</emphasis>
<taxonomicName id="4C364D51FB64FEA9CA7AB8C7FDC8FE5F" box="[322,544,476,501]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="verrillii">Lissodrillia verrillii</taxonomicName>
</emphasis>
can be distinguished by its small size and inflated, angular whorls. It is similar to
<taxonomicName id="4C364D51FB64FEA9CB8EBB1BFECFFDB2" box="[182,295,511,536]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="turgida" status="sp. nov.">
<emphasis id="B942EAC0FB64FEA9CB8EBB1BFECFFDB2" box="[182,295,511,536]" italics="true" pageId="257" pageNumber="258">L. turgida</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFB64FEA9CA00BB1BFE2DFDB2" box="[312,453,511,536]" pageId="257" pageNumber="258" rank="species">new species</taxonomicNameLabel>
, which differs in possessing evenly rounded, not angular whorls.
<taxonomicName id="4C364D51FB64FEA9CF84B8E4FA73FDB2" box="[1212,1435,511,536]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="verrillii">
<emphasis id="B942EAC0FB64FEA9CF84B8E4FA73FDB2" box="[1212,1435,511,536]" italics="true" pageId="257" pageNumber="258">Lissodrillia verrillii</emphasis>
</taxonomicName>
also has a narrower aperture, and a varix positioned ⅓-turn from the edge of the outer lip. It is closer to the edge of the outer lip in
<taxonomicName id="4C364D51FB64FEA9CA07BB53FE46FDCA" box="[319,430,583,608]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="turgida">
<emphasis id="B942EAC0FB64FEA9CA07BB53FE46FDCA" box="[319,430,583,608]" italics="true" pageId="257" pageNumber="258">L. turgida</emphasis>
</taxonomicName>
. Most of the other members of the genus have rather flat whorl profiles when compared to
<emphasis id="B942EAC0FB64FEA9CBAFBB76FEFAFD2F" box="[151,274,620,645]" italics="true" pageId="257" pageNumber="258">
<taxonomicName id="4C364D51FB64FEA9CBAFBB76FEE6FD2F" box="[151,270,620,645]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="257" pageNumber="258" phylum="Mollusca" rank="species" species="verrillii">L. verrillii</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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