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<document id="F0CCE0A7DDE88643E9855FEDE7588F2B" ID-CLB-Dataset="8891" ID-DOI="10.11646/zootaxa.4868.3.2" ID-GBIF-Dataset="03a9be04-f633-4adc-8397-058c282f216b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417761" ID-ZooBank="256D29E7-5148-415C-AD06-5F1F6D986887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609808401120" checkinUser="plazi" docAuthor="Cao, Yanghui, Dmitriev, Dmitry A., Dietrich, Christopher H. &amp; Zhang, Yalin" docDate="2020" docId="03998E57FFCFFFFE8B84FBC3FEA7BC45" docLanguage="en" docName="zootaxa.4868.3.2.pdf" docOrigin="Zootaxa 4868 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Etmaria triquetra Cao &amp; Zhang 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="347" masterDocId="FFA0F62FFFDFFFEE8B13FFAFFD37BB13" masterDocTitle="Review of the leafhopper genus Thaia Ghauri and related genera (Hemiptera: Cicadellidae: Typhlocybinae: Erythroneurini)" masterLastPageNumber="367" masterPageNumber="331" pageNumber="347" updateTime="1698852417663" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="308B9F984E1C15187A057F976007CCBC">Review of the leafhopper genus Thaia Ghauri and related genera (Hemiptera: Cicadellidae: Typhlocybinae: Erythroneurini)</mods:title>
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<mods:namePart id="CA03D01EB0634A6A764334931CA2E215">Cao, Yanghui</mods:namePart>
<mods:nameIdentifier id="5F4F044C3BAB9C5B9CC85B63A73D87F5" type="ORCID">0000-0002-0515-0767</mods:nameIdentifier>
<mods:affiliation id="B9409A322AD364F9579C8786623285FD">Key Laboratory of Plant Protection Resources and Pest Management, National Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, P. R. China &amp; Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; caoyh @ illinois. edu; https: // orcid. org / 0000 - 0002 - 0515 - 0767</mods:affiliation>
<mods:nameIdentifier id="8CC7C952C56E0CA33AEBD105658111A7" type="email">caoyh@illinois.edu</mods:nameIdentifier>
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<mods:namePart id="6D3D3ABF8F161636D85F4405C8B596DD">Dmitriev, Dmitry A.</mods:namePart>
<mods:nameIdentifier id="5A3DE1F87C317E39361E0AF42F5585F6" type="ORCID">0000-0002-3293-4794</mods:nameIdentifier>
<mods:affiliation id="F24D7340AAF791CB60ACC130B073EE36">Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; arboridia @ gmail. com; https: // orcid. org / 0000 - 0002 - 3293 - 4794</mods:affiliation>
<mods:nameIdentifier id="D9D4CEC87FE11C06B29B3EC5A6CD07C4" type="email">arboridia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="58D19051866BEC1DC4EEC60F8210A246">Dietrich, Christopher H.</mods:namePart>
<mods:nameIdentifier id="531B5C1AA2A59D9D39EC16AD9464D25B" type="ORCID">0000-0003-4005-4305</mods:nameIdentifier>
<mods:affiliation id="2C88A63E4FB232103969C62CEF5D43EF">Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; chdietri @ illinois. edu; https: // orcid. org / 0000 - 0003 - 4005 - 4305</mods:affiliation>
<mods:nameIdentifier id="1AE59973C49CA23FAEC3F7080C9077E8" type="email">chdietri@illinois.edu</mods:nameIdentifier>
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<mods:namePart id="E7331D6AA6614CA71D1715C68DDC2C18">Zhang, Yalin</mods:namePart>
<mods:affiliation id="8636071CB3058F69C9C659F0A3E2FA92">Key Laboratory of Plant Protection Resources and Pest Management, National Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, P. R. China</mods:affiliation>
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<mods:date id="8352C2F5F559064B927701F8E2F3DF7F">2020</mods:date>
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<mods:number id="8EB4B76C6424AA632962686CD7BFB1D2">2020-10-28</mods:number>
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<treatment id="03998E57FFCFFFFE8B84FBC3FEA7BC45" ID-DOI="http://doi.org/10.5281/zenodo.4417842" ID-GBIF-Taxon="176583721" ID-Zenodo-Dep="4417842" LSID="urn:lsid:plazi:treatment:03998E57FFCFFFFE8B84FBC3FEA7BC45" httpUri="http://treatment.plazi.org/id/03998E57FFCFFFFE8B84FBC3FEA7BC45" lastPageNumber="347" pageId="16" pageNumber="347">
<subSubSection id="C32A6CCAFFCFFFFE8B84FBC3FF98BF94" box="[151,687,1132,1159]" pageId="16" pageNumber="347" type="nomenclature">
<paragraph id="8B8F3F41FFCFFFFE8B84FBC3FF98BF94" blockId="16.[151,687,1132,1195]" box="[151,687,1132,1159]" pageId="16" pageNumber="347">
<heading id="D0C7882DFFCFFFFE8B84FBC3FF98BF94" bold="true" box="[151,687,1132,1159]" fontSize="11" level="1" pageId="16" pageNumber="347" reason="1">
<emphasis id="B944E353FFCFFFFE8B84FBC3FF98BF94" bold="true" box="[151,687,1132,1159]" pageId="16" pageNumber="347">
15.
<taxonomicName id="4C3044C2FFCFFFFE8BD0FBC3FF7CBF94" ID-CoL="8KJJN" authority="Cao &amp; Zhang" authorityName="Cao &amp; Zhang" authorityYear="2020" box="[195,587,1132,1159]" class="Insecta" family="Cicadellidae" genus="Etmaria" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="347" phylum="Arthropoda" rank="species" species="triquetra" status="sp. nov.">
<emphasis id="B944E353FFCFFFFE8BD0FBC3FCAFBF95" bold="true" box="[195,408,1132,1158]" italics="true" pageId="16" pageNumber="347">Etmaria triquetra</emphasis>
Cao &amp; Zhang
</taxonomicName>
<taxonomicNameLabel id="A2775E28FFCFFFFE8941FBC2FF98BF94" box="[594,687,1133,1159]" pageId="16" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFCFFFFE8B84FB3FFF45BD6C" pageId="16" pageNumber="347" type="description">
<paragraph id="8B8F3F41FFCFFFFE8B84FB3FFC71BFB8" blockId="16.[151,687,1132,1195]" box="[151,326,1168,1195]" pageId="16" pageNumber="347">
<figureCitation id="130B23C4FFCFFFFE8B84FB3FFDDBBFB8" box="[151,236,1168,1195]" captionStart="FIGURE 10" captionStartId="18.[151,250,1822,1846]" captionTargetBox="[153,1436,181,1787]" captionTargetId="figure-22@18.[151,1436,181,1799]" captionTargetPageId="18" captionText="FIGURE 10. Etmaria triquetra Cao &amp; Zhang sp. nov. a. abdominal apodemes 2S; b. genital capsule; c. anal tube appendage, lateral view; d. pygofer ventral appendage, lateral view; e. subgenital plate, style and connective; f. subgenital plate, dorsal view; g. style, dorsal view; h. style, lateral view; i. connective; j. aedeagus, lateral view; k. aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.4417783" httpUri="https://zenodo.org/record/4417783/files/figure.png" pageId="16" pageNumber="347">Figs 10</figureCitation>
,
<figureCitation id="130B23C4FFCFFFFE8BEBFB3EFC26BFB8" box="[248,273,1169,1195]" captionStart="FIGURE 19" captionStartId="32.[151,250,1876,1900]" captionTargetBox="[172,1413,190,1840]" captionTargetId="figure-22@32.[164,1423,181,1852]" captionTargetPageId="32" captionText="FIGURE 19. a–d. Etmaria infumata (Kuoh, 1982) comb. n.; e–h. Etmaria magna Cao &amp; Dietrich sp. nov.; i–l. Etmaria sinuata (Chiang &amp; Knight, 1990) comb. n.; m–p. Etmaria triquetra Cao &amp; Zhang sp. nov.; q–t. Etmaria ulterior Cao &amp; Zhang sp. nov.; u–x. Thaia (Thaia) barbata Dworakowska, 1979" figureDoi="http://doi.org/10.5281/zenodo.4417803" httpUri="https://zenodo.org/record/4417803/files/figure.png" pageId="16" pageNumber="347">19</figureCitation>
m–p
</paragraph>
<paragraph id="8B8F3F41FFCFFFFE8B84FB77FC80BE4D" blockId="16.[151,1437,1240,1879]" pageId="16" pageNumber="347">
<emphasis id="B944E353FFCFFFFE8B84FB77FC1EBFE1" bold="true" box="[151,297,1240,1266]" pageId="16" pageNumber="347">Description.</emphasis>
Vertex, face and mesonotum pale yellow with orange tint, vertex with pair of dark patches on transition to face, midline of frontoclypeal area and anteclypeus dark, eyes black. Pronotum dark orange. Basal triangles on mesonotum orange brown. Forewing light brown. Crown fore margin rounded apically, coronal suture distinct basally, anteclypeus long.
</paragraph>
<paragraph id="8B8F3F41FFCFFFFE8BD4FAC7F83EBE91" blockId="16.[151,1437,1240,1879]" box="[199,1289,1384,1411]" pageId="16" pageNumber="347">Anal tube very short, appendage extended from base, broad, slightly serrated on cephalic margin.</paragraph>
<paragraph id="8B8F3F41FFCFFFFE8BD4FA23F81BBD49" blockId="16.[151,1437,1240,1879]" pageId="16" pageNumber="347">Boundary between basal and distal pygofer lobes oblique and rounded; ventral appendage tubular, slightly sinuated, directed caudad, surpassing hind margin of lobe, without rigid setae at base. Subgenital plate with large distal disc, with about 3 macrosetae. Style with apical part thick and long, straight in dorsal view, apex pointed and curved in lateral view. Connective with manubrium long and narrow, margin of central lobe almost straight. Aedeagal shaft short and very slim, directed dorsad, with pair of tiny triangular processes near base, atrium very long in lateral view, nearly rectangular in caudal view; dorsal apodeme well developed; preatrial processes short and straight.</paragraph>
<paragraph id="8B8F3F41FFCFFFFE8BD4F9CAFF45BD6C" blockId="16.[151,1437,1240,1879]" box="[199,626,1636,1663]" pageId="16" pageNumber="347">
<emphasis id="B944E353FFCFFFFE8BD4F9CAFC44BD6C" bold="true" box="[199,371,1637,1663]" pageId="16" pageNumber="347">Measurement.</emphasis>
Male length 3.30 mm.
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFCFFFFE8BD4F927FFC8BDD4" pageId="16" pageNumber="347" type="materials_examined">
<paragraph id="8B8F3F41FFCFFFFE8BD4F927FFC8BDD4" blockId="16.[151,1437,1240,1879]" pageId="16" pageNumber="347">
<emphasis id="B944E353FFCFFFFE8BD4F927FC8EBDB1" bold="true" box="[199,441,1672,1698]" pageId="16" pageNumber="347">Material examined.</emphasis>
<materialsCitation id="3B58351CFFCFFFFE8ADBF927FFC8BDD4" ID-GBIF-Occurrence="3015520322" collectingDate="2012-04-30" collectionCode="NWAFU" collectorName="Yanghui Cao." country="China" elevation="1089" latitude="24.475834" location="Mangxian" longLatPrecision="21" longitude="97.75056" municipality="Yingjiang" pageId="16" pageNumber="347" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="548B81E3FFCFFFFE8ADBF927FF02BDB1" box="[456,565,1672,1698]" pageId="16" pageNumber="347" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D36F4C8FFCFFFFE8954F926FF69BDB0" box="[583,606,1673,1699]" pageId="16" pageNumber="347" type="male">♂</specimenCount>
,
<collectingCountry id="F3277FD1FFCFFFFE8961F926FFF9BDB0" box="[626,718,1673,1699]" name="China" pageId="16" pageNumber="347">CHINA</collectingCountry>
,
<collectingRegion id="49F4F1A3FFCFFFFE89F3F926FEB5BDB0" box="[736,898,1673,1699]" country="China" name="Yunnan" pageId="16" pageNumber="347">Yunnan Prov.</collectingRegion>
,
<collectingMunicipality id="6BEBA53BFFCFFFFE8880F927F933BDB1" box="[915,1028,1672,1698]" pageId="16" pageNumber="347">Yingjiang</collectingMunicipality>
,
<location id="8EEF699AFFCFFFFE8F0BF927F9BBBDB1" LSID="urn:lsid:plazi:treatment:03998E57FFCFFFFE8B84FBC3FEA7BC45:8EEF699AFFCFFFFE8F0BF927F9BBBDB1" box="[1048,1164,1672,1698]" country="China" latitude="24.475834" longLatPrecision="21" longitude="97.75056" municipality="Yingjiang" name="Mangxian" pageId="16" pageNumber="347" stateProvince="Yunnan">Mangxian</location>
,
<quantity id="4CC892A4FFCFFFFE8FB3F926F9C8BDB0" box="[1184,1279,1673,1699]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="16" pageNumber="347" unit="m" value="1089.0">
<elevation id="001DD872FFCFFFFE8FB3F926F9C8BDB0" box="[1184,1279,1673,1699]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="16" pageNumber="347" unit="m" value="1089.0">1089 m</elevation>
</quantity>
,
<geoCoordinate id="EE045986FFCFFFFE8E00F926F8AFBDB0" box="[1299,1432,1673,1699]" degrees="24" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="347" precision="15" seconds="33" value="24.475834">24°28'33''N</geoCoordinate>
,
<geoCoordinate id="EE045986FFCFFFFE8B84F903FC2EBDD4" box="[151,281,1708,1735]" degrees="97" direction="east" minutes="45" orientation="longitude" pageId="16" pageNumber="347" precision="15" seconds="02" value="97.75056">97°45'02''E</geoCoordinate>
,
<date id="FF8E1981FFCFFFFE8A36F902FCA8BDD4" box="[293,415,1708,1735]" pageId="16" pageNumber="347" value="2012-04-30">
<collectingDate id="EFCAE069FFCFFFFE8A36F902FCA8BDD4" box="[293,415,1708,1735]" pageId="16" pageNumber="347" value="2012-04-30">30 iv 2012</collectingDate>
</date>
, coll.
<collectorName id="26C55A97FFCFFFFE8AF2F903FF4CBDD4" box="[481,635,1708,1735]" pageId="16" pageNumber="347">Yanghui Cao.</collectorName>
[
<collectionCode id="ED21A784FFCFFFFE8994F903FFC3BDD4" box="[647,756,1708,1735]" pageId="16" pageNumber="347">NWAFU</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFCFFFFE8BD4F97EFE40BC1D" pageId="16" pageNumber="347" type="etymology">
<paragraph id="8B8F3F41FFCFFFFE8BD4F97EFE40BC1D" blockId="16.[151,1437,1240,1879]" pageId="16" pageNumber="347">
<emphasis id="B944E353FFCFFFFE8BD4F97EFC79BDF8" bold="true" box="[199,334,1745,1771]" pageId="16" pageNumber="347">Etymology.</emphasis>
The new specific epithet is derived from Latin adjective “
<emphasis id="B944E353FFCFFFFE88DDF97EF90BBDF9" box="[974,1084,1745,1770]" italics="true" pageId="16" pageNumber="347">triquetrus</emphasis>
” which means triangular, referring to the pair of basal triangular processes of the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFCFFFFE8BD4F8B7FEA7BC45" pageId="16" pageNumber="347" type="diagnosis">
<paragraph id="8B8F3F41FFCFFFFE8BD4F8B7FEA7BC45" blockId="16.[151,1437,1240,1879]" pageId="16" pageNumber="347">
<emphasis id="B944E353FFCFFFFE8BD4F8B7FC75BC21" bold="true" box="[199,322,1816,1842]" pageId="16" pageNumber="347">Diagnosis.</emphasis>
The new species can be distinguished by the long anal tube appendage shifted basad and the presence of tiny triangular processes near the base of the aedeagal shaft.
</paragraph>
</subSubSection>
</treatment>
</document>
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