treatments-xml/data/03/DC/9D/03DC9D25FFBBFFA0FF27FCE4FB83FCF1.xml

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<document id="5A2C858264E67846AC68FAEF0FDD2E14" ID-CLB-Dataset="27294" ID-DOI="10.11646/zootaxa.4585.3.9" ID-GBIF-Dataset="9b404efd-c9b4-4d83-bc6d-1fa39672a9c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="2640279" ID-ZooBank="E6BBA677-EB76-461E-ACBC-FE57E1E5926E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555312417773" checkinUser="plazi" docAuthor="Teslenko, Valentina A., Palatov, Dmitry M. &amp; Semenchenko, Alexander A." docDate="2019" docId="03DC9D25FFBBFFA0FF27FCE4FB83FCF1" docLanguage="en" docName="zootaxa.4585.3.9.pdf" docOrigin="Zootaxa 4585 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lectra adjariae Teslenko 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="552" masterDocId="FFE5E55DFFB9FFA6FFB0FFADFFF7FFC6" masterDocTitle="Description of new apterous winter species of Leuctra (Plecoptera: Leuctridae) based morphology and DNA barcoding and further records to stonefly fauna of the Caucasus, Georgia" masterLastPageNumber="560" masterPageNumber="546" pageNumber="548" updateTime="1698721741559" updateUser="ExternalLinkService">
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<mods:title id="CA72BE946C6CEF1DF42D60FBBD618948">Description of new apterous winter species of Leuctra (Plecoptera: Leuctridae) based morphology and DNA barcoding and further records to stonefly fauna of the Caucasus, Georgia</mods:title>
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<mods:namePart id="DBB8EB92F6143A39030EB202C915ADDA">Teslenko, Valentina A.</mods:namePart>
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<mods:namePart id="EC8EDB0B5BFABD734AA749E69A3D4F5B">Palatov, Dmitry M.</mods:namePart>
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<mods:namePart id="EEC7683E74660E5214BA5151608B2142">Semenchenko, Alexander A.</mods:namePart>
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<treatment id="03DC9D25FFBBFFA0FF27FCE4FB83FCF1" ID-DOI="http://doi.org/10.5281/zenodo.5941115" ID-GBIF-Taxon="156199730" ID-Zenodo-Dep="5941115" LSID="urn:lsid:plazi:treatment:03DC9D25FFBBFFA0FF27FCE4FB83FCF1" httpUri="http://treatment.plazi.org/id/03DC9D25FFBBFFA0FF27FCE4FB83FCF1" lastPageId="6" lastPageNumber="552" pageId="2" pageNumber="548">
<subSubSection id="C36F7FB8FFBBFFA4FF27FCE4FDF9FCA5" box="[151,526,841,867]" pageId="2" pageNumber="548" type="nomenclature">
<paragraph id="8BCA2C33FFBBFFA4FF27FCE4FDF9FCA5" blockId="2.[151,984,841,936]" box="[151,526,841,867]" pageId="2" pageNumber="548">
<heading id="D0829B5FFFBBFFA4FF27FCE4FDF9FCA5" bold="true" box="[151,526,841,867]" fontSize="11" level="1" pageId="2" pageNumber="548" reason="1">
<emphasis id="B901F021FFBBFFA4FF27FCE4FDF9FCA5" bold="true" box="[151,526,841,867]" pageId="2" pageNumber="548">
<taxonomicName id="4C7557B0FFBBFFA4FF27FCE4FE32FCA5" authority="Teslenko" authorityName="Teslenko" authorityYear="2019" box="[151,453,841,867]" class="Insecta" family="Leuctridae" genus="Lectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="548" phylum="Arthropoda" rank="species" species="adjariae" status="sp. nov.">
<emphasis id="B901F021FFBBFFA4FF27FCE4FEB8FCA5" bold="true" box="[151,335,841,867]" italics="true" pageId="2" pageNumber="548">Lectra adjariae</emphasis>
Teslenko
</taxonomicName>
<taxonomicNameLabel id="A2324D5AFFBBFFA4FE7CFCE4FDF9FCA5" box="[460,526,841,867]" pageId="2" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="C36F7FB8FFBBFFA4FF27FCC1FC2FFC6E" pageId="2" pageNumber="548" type="description">
<paragraph id="8BCA2C33FFBBFFA4FF27FCC1FEEEFC42" blockId="2.[151,984,841,936]" box="[151,281,876,901]" pageId="2" pageNumber="548">
(
<figureCitation id="134E30B6FFBBFFA4FF2FFCC1FEE7FC42" box="[159,272,876,901]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 24" captionStart-2="FIGURES 59" captionStart-3="FIGURES 1013" captionStartId-0="2.[151,250,677,699]" captionStartId-1="3.[151,264,1555,1578]" captionStartId-2="4.[151,264,1975,1998]" captionStartId-3="5.[151,264,1435,1458]" captionTargetBox-0="[176,1411,205,645]" captionTargetBox-1="[300,1392,193,1525]" captionTargetBox-2="[170,1400,340,1946]" captionTargetBox-3="[150,1421,193,1407]" captionTargetId-0="figure@2.[164,1412,195,662]" captionTargetId-1="figure@3.[162,1416,193,1538]" captionTargetId-2="figure@4.[170,1417,337,1955]" captionTargetId-3="figure@5.[150,1436,193,1415]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Maximum Likelihood (ML) tree (-Ln likelihood = 1785.88) of Leuctra adjariae sp. n., Leuctra georgiae sp. n. and outgroup inferred from the cytochrome c oxidase I (COI) nucleotide sequence data (658 bp). Numbers are bootstrap support of 500 replicates. Specimens obtained in this study are in BOLD." captionText-1="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." captionText-2="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." captionText-3="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri-0="https://zenodo.org/record/2640281/files/figure.png" httpUri-1="https://zenodo.org/record/2640283/files/figure.png" httpUri-2="https://zenodo.org/record/2640285/files/figure.png" httpUri-3="https://zenodo.org/record/2640287/files/figure.png" pageId="2" pageNumber="548">Figs 113</figureCitation>
)
</paragraph>
<paragraph id="8BCA2C33FFBBFFA4FF27FC22FC2FFC6E" blockId="2.[151,984,841,936]" box="[151,984,911,936]" pageId="2" pageNumber="548">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C36F7FB8FFBBFFA4FF27FC78FACDFBF0" pageId="2" pageNumber="548" type="materials_examined">
<paragraph id="8BCA2C33FFBBFFA4FF27FC78FACDFBF0" blockId="2.[151,1437,981,2031]" pageId="2" pageNumber="548">
<emphasis id="B901F021FFBBFFA4FF27FC78FE83FC28" bold="true" box="[151,372,981,1006]" pageId="2" pageNumber="548">Material examined.</emphasis>
<materialsCitation id="3B1D266EFFBBFFA4FEC9FC78FAFBFBD4" ID-GBIF-Occurrence="2235669110" collectionCode="FSC, EATB, FEB" collectorName="D. Palatov" country="Georgia" latitude="41.786533" location="Kintrishi River" longLatPrecision="1" longitude="41.9565" pageId="2" pageNumber="548" specimenCount="1" stateProvince="Ajaria" typeStatus="holotype">
<typeStatus id="54CE9291FFBBFFA4FEC9FC78FE2BFC28" box="[377,476,981,1006]" pageId="2" pageNumber="548" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F3626CA3FFBBFFA4FD92FC78FD8DFC28" box="[546,634,981,1006]" name="Georgia" pageId="2" pageNumber="548">Georgia</collectingCountry>
.
<collectingRegion id="49B1E2D1FFBBFFA4FD33FC78FD22FC28" box="[643,725,981,1006]" country="Georgia" name="Ajaria" pageId="2" pageNumber="548">Adjaria</collectingRegion>
.
<location id="8EAA7AE8FFBBFFA4FD6EFC78FC8AFC28" LSID="urn:lsid:plazi:treatment:03DC9D25FFBBFFA0FF27FCE4FB83FCF1:8EAA7AE8FFBBFFA4FD6EFC78FC8AFC28" box="[734,893,981,1006]" country="Georgia" latitude="41.786533" longLatPrecision="1" longitude="41.9565" name="Kintrishi River" pageId="2" pageNumber="548" stateProvince="Ajaria">Kintrishi River</location>
,
<locationDeviation id="9A783511FFBBFFA4FC37FC78FB3AFC28" box="[903,1229,981,1006]" location="Kobuleti" pageId="2" pageNumber="548">
<quantity id="4C8D81D6FFBBFFA4FC37FC78FC3DFC28" box="[903,970,981,1006]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="548" unit="km" value="16.0">16 km</quantity>
upstream from Kobuleti
</locationDeviation>
and
<locationDeviation id="9A783511FFBBFFA4FB4FFC7BFE64FBD4" location="Tchakhati Village" pageId="2" pageNumber="548">
<quantity id="4C8D81D6FFBBFFA4FB4FFC7BFAC2FC28" box="[1279,1333,981,1006]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="548" unit="km" value="4.0">4 km</quantity>
upstream from Tchakhati Village
</locationDeviation>
,
<geoCoordinate id="EE414AF4FFBBFFA4FE10FC57FDDDFBD7" box="[416,554,1018,1042]" direction="north" orientation="latitude" pageId="2" pageNumber="548" precision="1" value="41.786533">41°47.192 N</geoCoordinate>
<geoCoordinate id="EE414AF4FFBBFFA4FD84FC54FD4AFBD7" box="[564,701,1017,1042]" direction="east" orientation="longitude" pageId="2" pageNumber="548" precision="1" value="41.9565">41°57.390 E</geoCoordinate>
, 0 6.02.2017, coll.
<collectorName id="268049E5FFBBFFA4FC38FC57FC0DFBD4" box="[904,1018,1017,1042]" pageId="2" pageNumber="548">D. Palatov</collectorName>
(
<collectionCode id="ED64B4F6FFBBFFA4FBBAFC57FBCCFBD4" box="[1034,1083,1018,1042]" country="Canada" httpUri="http://grbio.org/cool/q309-g0ck" name="Fredericton Stock Culture Collection" pageId="2" pageNumber="548">FSC</collectionCode>
<collectionCode id="ED64B4F6FFBBFFA4FBF5FC57FB35FBD7" box="[1093,1218,1018,1042]" pageId="2" pageNumber="548">EATB FEB</collectionCode>
RAS)
</materialsCitation>
.
<typeStatus id="54CE9291FFBBFFA4FAA8FC57FA74FBD4" box="[1304,1411,1018,1042]" pageId="2" pageNumber="548" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D73E7BAFFBBFFA4FA3FFC57FF22FBF0" pageId="2" pageNumber="548" type="male">2 males</specimenCount>
(one mounted),
<materialsCitation id="3B1D266EFFBBFFA4FE33FBB3FAC1FBF0" ID-GBIF-Occurrence="2235669108" box="[387,1334,1053,1078]" collectionCode="FSC, EATB, FEB" collectorName="D. Palatov" country="Georgia" latitude="41.786533" location="Kintrishi River" longLatPrecision="1" longitude="41.9565" pageId="2" pageNumber="548" specimenCount="3" specimenCount-female="3" stateProvince="Ajaria" typeStatus="paratype">
<specimenCount id="9D73E7BAFFBBFFA4FE33FBB3FE1FFBF0" box="[387,488,1053,1078]" pageId="2" pageNumber="548" type="female">3 females</specimenCount>
(one mounted), same locality and date, coll.
<collectorName id="268049E5FFBBFFA4FC71FBB3FBC7FBF0" box="[961,1072,1053,1078]" pageId="2" pageNumber="548">D. Palatov</collectorName>
(
<collectionCode id="ED64B4F6FFBBFFA4FB8EFBB3FB98FBF0" box="[1086,1135,1054,1078]" country="Canada" httpUri="http://grbio.org/cool/q309-g0ck" name="Fredericton Stock Culture Collection" pageId="2" pageNumber="548">FSC</collectionCode>
<collectionCode id="ED64B4F6FFBBFFA4FBC5FBB3FB07FBF3" box="[1141,1264,1054,1078]" pageId="2" pageNumber="548">EATB FEB</collectionCode>
RAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F7FB8FFBBFFA2FF77FBECFDEDFEDA" lastPageId="4" lastPageNumber="550" pageId="2" pageNumber="548" type="description">
<paragraph id="8BCA2C33FFBBFFA4FF77FBECFAB4FB00" blockId="2.[151,1437,981,2031]" pageId="2" pageNumber="548">
<emphasis id="B901F021FFBBFFA4FF77FBECFEA6FB9C" bold="true" box="[199,337,1089,1114]" pageId="2" pageNumber="548">Description</emphasis>
. Body length of males 3.54.0 mm, females 4.05.0 mm. Apterous species in both sexes. General color brown, sclerotization heavily, body covered with pilosity, visible mainly in lateral view (
<figureCitation id="134E30B6FFBBFFA4FB0EFBC8FAF4FBB8" box="[1214,1283,1125,1150]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 4</figureCitation>
). Palpi, head, pronotum, meso-, metanotum and legs uniformly brown, dark brown rugosities on the occiput and pronotum (
<figureCitation id="134E30B6FFBBFFA4FADFFB24FF5FFB00" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 2</figureCitation>
). Head bears two small, barely visible lateral ocelli, the median ocellus indistinct, difficult to distinguish.
</paragraph>
<paragraph id="8BCA2C33FFBBFFA4FF77FB7CFB24FA6A" blockId="2.[151,1437,981,2031]" pageId="2" pageNumber="548">
Prothoracic sternum of male with diamond-shaped basisternum (b
<subScript id="17F12E76FFBBFFA4FC0CFB4DFC33FB28" attach="left" box="[956,964,1248,1262]" fontSize="6" pageId="2" pageNumber="548">1</subScript>
), furcasternum (f
<subScript id="17F12E76FFBBFFA4FB3EFB4DFB61FB28" attach="left" box="[1166,1174,1248,1262]" fontSize="6" pageId="2" pageNumber="548">1</subScript>
) is a pair of diverging bars, postfurcasternum (p
<subScript id="17F12E76FFBBFFA4FE05FAAAFE4AFAD3" attach="left" box="[437,445,1287,1301]" fontSize="6" pageId="2" pageNumber="548">1</subScript>
)—a large elliptical and medially subdivided sclerite, spinasternum (s
<subScript id="17F12E76FFBBFFA4FB79FAAAFB26FAD3" attach="left" box="[1225,1233,1287,1301]" fontSize="6" pageId="2" pageNumber="548">1</subScript>
) obscure, appears as a small round sclerite surrounded by pleurae, (
<figureCitation id="134E30B6FFBBFFA4FD0DFAB3FCF6FAF1" box="[701,769,1310,1335]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 3</figureCitation>
). Mesothoracic sternum bears basisternum (b
<subScript id="17F12E76FFBBFFA4FB4FFA83FAF0FAFA" attach="left" box="[1279,1287,1326,1340]" fontSize="6" pageId="2" pageNumber="548">2</subScript>
) much wider than that on the prothoracic sternum, with two triangular anterolateral processes; furcasternum (f
<subScript id="17F12E76FFBBFFA4FB6BFAF9FB14FAA4" attach="left" box="[1243,1251,1364,1378]" fontSize="6" pageId="2" pageNumber="548">2</subScript>
) separated from the basisternum by distinct arms; postfurkasternum (p
<subScript id="17F12E76FFBBFFA4FD49FAD6FCF6FA4F" attach="left" box="[761,769,1403,1417]" fontSize="6" pageId="2" pageNumber="548">2</subScript>
) as a pair of sclerotized rounded triangle sclerites laterally to furkasternum arms (f
<subScript id="17F12E76FFBBFFA4FE10FA0FFE5FFA76" attach="left" box="[416,424,1442,1456]" fontSize="6" pageId="2" pageNumber="548">2</subScript>
) not fused to furcasternum; spinasternum (s
<subScript id="17F12E76FFBBFFA4FC26FA0FFC69FA76" attach="left" box="[918,926,1442,1456]" fontSize="6" pageId="2" pageNumber="548">2</subScript>
) narrow elongated (
<figureCitation id="134E30B6FFBBFFA4FB31FA3EFB32FA6A" box="[1153,1221,1427,1452]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BCA2C33FFBBFFA4FF77FA17FC3BF829" blockId="2.[151,1437,981,2031]" pageId="2" pageNumber="548">
<emphasis id="B901F021FFBBFFA4FF77FA17FEFCFA15" bold="true" box="[199,267,1466,1491]" pageId="2" pageNumber="548">Male.</emphasis>
Terga I-VII simple, each tergum with heavily sclerotized anterior margin and semicircular paramedian membranous area, increased in size posteriorly to tergum VIII and covered by dense, long black setae (
<figureCitation id="134E30B6FFBBFFA4FAF7FA73FA79FA31" box="[1351,1422,1502,1527]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Fig. 5</figureCitation>
). Tergum VIII bears indistinct weakly sclerotized anteromedial process looking like a pair of short slightly pigmented fan-shaped spots, closely separated, directed backward and covered by long black setae (
<figureCitation id="134E30B6FFBBFFA4FB5EF98BFAC6F9F9" box="[1262,1329,1574,1599]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Fig. 5</figureCitation>
). Tergum IX mostly membranous with distinct antecosta, divided medially for more than ¼ of segment width; posteromedial sclerite consists of paired small prolonged pigmented spots covered with long, black setae (
<figureCitation id="134E30B6FFBBFFA4FB01F9C3FAD7F941" box="[1201,1312,1646,1671]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Figs. 5, 6</figureCitation>
). Posterior margin of tergum X with wide and deep median notch, each lobe bears pointed process at posterior inner margin, directed inward and oblique backward in dorsal view and risen in lateral view (
<figureCitation id="134E30B6FFBBFFA4FBF5F91BFB41F909" box="[1093,1206,1718,1743]" captionStart-0="FIGURES 24" captionStart-1="FIGURES 59" captionStartId-0="3.[151,264,1555,1578]" captionStartId-1="4.[151,264,1975,1998]" captionTargetBox-0="[300,1392,193,1525]" captionTargetBox-1="[170,1400,340,1946]" captionTargetId-0="figure@3.[162,1416,193,1538]" captionTargetId-1="figure@4.[170,1417,337,1955]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." captionText-1="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri-0="https://zenodo.org/record/2640283/files/figure.png" httpUri-1="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Figs. 47</figureCitation>
). Epiproct rounded, mushroom-shaped, membranous with pigmented stalk (
<figureCitation id="134E30B6FFBBFFA4FCA3F977FCADF934" box="[787,858,1754,1779]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Fig. 7</figureCitation>
). Cerci are very specific, each cercus large, wide, scoop-shaped, gently bent inward and upward, hardly sclerotized on outer edge; the inner part membranous, covered with long black setae, denser than on the outer edge; apex wide, with slight depression on edge (
<figureCitation id="134E30B6FFBBFFA4FA9CF88FFF50F899" captionStart-0="FIGURES 24" captionStart-1="FIGURES 59" captionStartId-0="3.[151,264,1555,1578]" captionStartId-1="4.[151,264,1975,1998]" captionTargetBox-0="[300,1392,193,1525]" captionTargetBox-1="[170,1400,340,1946]" captionTargetId-0="figure@3.[162,1416,193,1538]" captionTargetId-1="figure@4.[170,1417,337,1955]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." captionText-1="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri-0="https://zenodo.org/record/2640283/files/figure.png" httpUri-1="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Figs. 46, 8</figureCitation>
). Paraprocts stout (
<figureCitation id="134E30B6FFBBFFA4FE35F8EBFE21F899" box="[389,470,1862,1887]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Figs. 4</figureCitation>
,
<figureCitation id="134E30B6FFBBFFA4FE54F8EAFE03F899" box="[484,500,1863,1887]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">9</figureCitation>
), styles 2.25X shorter than specilla (
<figureCitation id="134E30B6FFBBFFA4FC29F8EBFC17F899" box="[921,992,1862,1887]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Fig. 9</figureCitation>
). Styles with wide base; narrowed and curved sideward near the apex, and ending in rounded tip (
<figureCitation id="134E30B6FFBBFFA4FC93F8C7FC55F844" box="[803,930,1898,1923]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Figs 57, 9</figureCitation>
). Specilla long gently bent inward apically in dorsal view, narrowed to the apex and ending in a sharp point (
<figureCitation id="134E30B6FFBBFFA4FCEDF823FBF5F861" box="[861,1026,1934,1959]" captionStart="FIGURES 59" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[170,1400,340,1946]" captionTargetId="figure@4.[170,1417,337,1955]" captionTargetPageId="4" captionText="FIGURES 59. Male of Leuctra adjariae sp. n. 5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral." httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="2" pageNumber="548">Figs. 5, 6, 8, 9</figureCitation>
). In lateral view the apex of specilla pointed and directed upward (but downward as shown in
<figureCitation id="134E30B6FFBBFFA4FC81F81FFC82F80C" box="[817,885,1970,1995]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 4</figureCitation>
since the abdomen curved, in mating position). Sternum IX with U-shaped unsclerotized area, no ventral vesicle (
<figureCitation id="134E30B6FFBBFFA4FCC9F87BFC48F829" box="[889,959,2006,2031]" captionStart="FIGURES 24" captionStartId="3.[151,264,1555,1578]" captionTargetBox="[300,1392,193,1525]" captionTargetId="figure@3.[162,1416,193,1538]" captionTargetPageId="3" captionText="FIGURES 24. Male of Leuctra adjariae sp. n. 2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b 1 —basisternum, f 1 —furcasternum p 1 —postfurcasternum, s 1 —spinasternum; b 2, f 2, p 2, s 2 —the same on the mesothorax. 4. Abdominal tip, lateral." httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="2" pageNumber="548">Fig. 2</figureCitation>
).
</paragraph>
<caption id="DF0A7CBBFFBAFFA5FF27F9BEFE8AF9AC" httpUri="https://zenodo.org/record/2640283/files/figure.png" pageId="3" pageNumber="549" startId="3.[151,264,1555,1578]" targetBox="[300,1392,193,1525]" targetPageId="3">
<paragraph id="8BCA2C33FFBAFFA5FF27F9BEFE8AF9AC" blockId="3.[151,1437,1555,1642]" pageId="3" pageNumber="549">
<emphasis id="B901F021FFBAFFA5FF27F9BEFEB5F9EF" bold="true" box="[151,322,1555,1578]" pageId="3" pageNumber="549">FIGURES 24.</emphasis>
Male of
<taxonomicName id="4C7557B0FFBAFFA5FE03F9B8FD91F9EC" authority="Teslenko &amp; Palatov &amp; Semenchenko, 2019" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[435,614,1556,1578]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="549" phylum="Arthropoda" rank="species" species="adjariae" status="sp. nov.">
<emphasis id="B901F021FFBAFFA5FE03F9B8FD91F9EC" box="[435,614,1556,1578]" italics="true" pageId="3" pageNumber="549">Leuctra adjariae</emphasis>
</taxonomicName>
<emphasis id="B901F021FFBAFFA5FDC3F9B9FD44F9EF" bold="true" box="[627,691,1556,1577]" pageId="3" pageNumber="549">
<taxonomicNameLabel id="A2324D5AFFBAFFA5FDC3F9B9FD44F9EF" box="[627,691,1556,1577]" pageId="3" pageNumber="549" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
2. Habitus, dorsal view. 3. Pro- meso- and methatorax. Prothorax: b
<subScript id="17F12E76FFBAFFA5FF14F9EDFF5CF98A" attach="left" box="[164,171,1600,1612]" fontSize="5" pageId="3" pageNumber="549">1</subScript>
—basisternum, f
<subScript id="17F12E76FFBAFFA5FEE8F9EDFEA8F98A" attach="left" box="[344,351,1600,1612]" fontSize="5" pageId="3" pageNumber="549">1</subScript>
—furcasternum p
<subScript id="17F12E76FFBAFFA5FDA5F9EDFDEBF98A" attach="left" box="[533,540,1600,1612]" fontSize="5" pageId="3" pageNumber="549">1</subScript>
—postfurcasternum, s
<subScript id="17F12E76FFBAFFA5FCB0F9EDFCF0F98A" attach="left" box="[768,775,1600,1612]" fontSize="5" pageId="3" pageNumber="549">1</subScript>
—spinasternum; b
<subScript id="17F12E76FFBAFFA5FC76F9EDFC3AF98A" attach="left" box="[966,973,1600,1612]" fontSize="5" pageId="3" pageNumber="549">2</subScript>
, f
<subScript id="17F12E76FFBAFFA5FC56F9EDFC05F98A" attach="left" box="[998,1010,1600,1612]" fontSize="5" pageId="3" pageNumber="549">2,</subScript>
p
<subScript id="17F12E76FFBAFFA5FBB8F9EDFBF8F98A" attach="left" box="[1032,1039,1600,1612]" fontSize="5" pageId="3" pageNumber="549">2</subScript>
, s
<subScript id="17F12E76FFBAFFA5FB9BF9EDFBC5F98A" attach="left" box="[1067,1074,1600,1612]" fontSize="5" pageId="3" pageNumber="549">2</subScript>
—the same on the mesothorax. 4. Abdominal tip, lateral.
</paragraph>
</caption>
<paragraph id="8BCA2C33FFBAFFA2FF77F93BFDEDFEDA" blockId="3.[151,1436,1686,2035]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="550" pageId="3" pageNumber="549">
<emphasis id="B901F021FFBAFFA5FF77F93BFED2F969" bold="true" box="[199,293,1686,1711]" pageId="3" pageNumber="549">Female.</emphasis>
Terga II-VI with dorsal sclerite and pair of unsclerotized paramedian bands, the size of dorsal sclerite increasing to tergum VI; terga VII-X completely sclerotized. Sternum VII convex, very large and strongly sclerotized, almost trapezoidal with truncated anterolateral margins, anterior margin slightly concave, posterior margin broadly rounded, overlapping the anterior margin of sternum VIII and partly hiding the base of subgenital plate (
<figureCitation id="134E30B6FFBAFFA5FF6EF88BFEC3F8F9" box="[222,308,1830,1855]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1435,1458]" captionTargetBox="[150,1421,193,1407]" captionTargetId="figure@5.[150,1436,193,1415]" captionTargetPageId="5" captionText="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="3" pageNumber="549">Fig. 10</figureCitation>
). Subgenital plate triangular, prolonged, reaches anterior margin of sternum IX: a wide, triangular, unpigmented base delimited by a pair of small sclerotized sports anterolaterally, the second pair of small pigmented spots supports the plate laterally in the first ¼ of its length; further to the subgenital plate narrowing, ending in a pair of sclerotized finger-shaped prolonged posterior lobes, shortly bifurcated at the apex and directed slightly sideward; apex with membranous area mesoposteriorly (
<figureCitation id="134E30B6FFBAFFA5FC8CF81BFC60F809" box="[828,919,1974,1999]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1435,1458]" captionTargetBox="[150,1421,193,1407]" captionTargetId="figure@5.[150,1436,193,1415]" captionTargetPageId="5" captionText="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="3" pageNumber="549">Fig. 10</figureCitation>
). Subgenital plate covered with long setae especially in less pigmented part between sclerotized finger-shaped lobes, along inner edges of the lateral sclerites and on posterior margin (
<figureCitation id="134E30B6FFBDFFA2FE02FF3AFDF3FF76" box="[434,516,151,176]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1435,1458]" captionTargetBox="[150,1421,193,1407]" captionTargetId="figure@5.[150,1436,193,1415]" captionTargetPageId="5" captionText="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="4" pageNumber="550">Fig. 11</figureCitation>
). Sternum IX has two pairs of small triangular, membranous spots and one pair of sclerotized rounded spots between distal tops of each pair of membranous spots, below the subgenital plate (
<figureCitation id="134E30B6FFBDFFA2FADFFF11FF41FF3E" captionStart="FIGURES 1013" captionStartId="5.[151,264,1435,1458]" captionTargetBox="[150,1421,193,1407]" captionTargetId="figure@5.[150,1436,193,1415]" captionTargetPageId="5" captionText="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="4" pageNumber="550">Fig. 10</figureCitation>
). Seminal receptacle spheroid enclosing spermathecal ring-shaped slightly sclerotized sclerite with pair of lateral pointed arms (
<figureCitation id="134E30B6FFBDFFA2FE35FEA9FDFBFEDA" box="[389,524,260,285]" captionStart="FIGURES 1013" captionStartId="5.[151,264,1435,1458]" captionTargetBox="[150,1421,193,1407]" captionTargetId="figure@5.[150,1436,193,1415]" captionTargetPageId="5" captionText="FIGURES 1013. Female of Leuctra adjariae sp. n. 10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral." httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="4" pageNumber="550">Figs. 12, 13</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0A7CBBFFBDFFA2FF27F81AFB9BF82B" httpUri="https://zenodo.org/record/2640285/files/figure.png" pageId="4" pageNumber="550" startId="4.[151,264,1975,1998]" targetBox="[170,1400,340,1946]" targetPageId="4">
<paragraph id="8BCA2C33FFBDFFA2FF27F81AFB9BF82B" blockId="4.[151,1436,1975,2029]" pageId="4" pageNumber="550">
<emphasis id="B901F021FFBDFFA2FF27F81AFECDF80B" bold="true" box="[151,314,1975,1998]" pageId="4" pageNumber="550">FIGURES 59.</emphasis>
Male of
<taxonomicName id="4C7557B0FFBDFFA2FE25F814FDB7F808" authority="Teslenko &amp; Palatov &amp; Semenchenko, 2019" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[405,576,1976,1998]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="550" phylum="Arthropoda" rank="species" species="adjariae" status="sp. nov.">
<emphasis id="B901F021FFBDFFA2FE25F814FDB7F808" box="[405,576,1976,1998]" italics="true" pageId="4" pageNumber="550">Leuctra adjariae</emphasis>
</taxonomicName>
<emphasis id="B901F021FFBDFFA2FDF6F815FD88F80B" bold="true" box="[582,639,1976,1997]" pageId="4" pageNumber="550">
<taxonomicNameLabel id="A2324D5AFFBDFFA2FDF6F815FD88F80B" box="[582,639,1976,1997]" pageId="4" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
5. Abdominal tip, VII-X terga, dorsal. 6. Abdominal tip, IX-X terga, dorsal. 7. Epiproct, and styles, dorsal. 8. Abdominal tip, IX-X terga, ventral. 9. Styles and specilla, ventral.
</paragraph>
</caption>
<caption id="DF0A7CBBFFBCFFA3FF27FA36FAF4FA17" httpUri="https://zenodo.org/record/2640287/files/figure.png" pageId="5" pageNumber="551" startId="5.[151,264,1435,1458]" targetBox="[150,1421,193,1407]" targetPageId="5">
<paragraph id="8BCA2C33FFBCFFA3FF27FA36FAF4FA17" blockId="5.[151,1436,1435,1489]" pageId="5" pageNumber="551">
<emphasis id="B901F021FFBCFFA3FF27FA36FEA4FA77" bold="true" box="[151,339,1435,1458]" pageId="5" pageNumber="551">FIGURES 1013.</emphasis>
Female of
<taxonomicName id="4C7557B0FFBCFFA3FE75FA30FD87FA74" authority="Teslenko &amp; Palatov &amp; Semenchenko, 2019" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[453,624,1436,1458]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="551" phylum="Arthropoda" rank="species" species="adjariae" status="sp. nov.">
<emphasis id="B901F021FFBCFFA3FE75FA30FD87FA74" box="[453,624,1436,1458]" italics="true" pageId="5" pageNumber="551">Leuctra adjariae</emphasis>
</taxonomicName>
<emphasis id="B901F021FFBCFFA3FDC5FA31FD59FA77" bold="true" box="[629,686,1436,1457]" pageId="5" pageNumber="551">
<taxonomicNameLabel id="A2324D5AFFBCFFA3FDC5FA31FD59FA77" box="[629,686,1436,1457]" pageId="5" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
10. Abdominal tip, subgenital plate, ventral. 11. Subgenital plate, cleared, mounted, ventral. 12. Spermatheca, cleared, mounted ventral. 13. Spermatheca, cleared, mounted, ventrolateral.
</paragraph>
</caption>
<subSubSection id="C36F7FB8FFBCFFA0FF77FA50FF08FDDE" lastPageId="6" lastPageNumber="552" pageId="5" pageNumber="551" type="diagnosis">
<paragraph id="8BCA2C33FFBCFFA3FF77FA50FF0BF867" blockId="5.[151,1437,1533,2026]" pageId="5" pageNumber="551">
<emphasis id="B901F021FFBCFFA3FF77FA50FEB6F9D0" bold="true" box="[199,321,1533,1558]" pageId="5" pageNumber="551">Diagnosis.</emphasis>
Apterous in both sexes. Terga of male I-VII simple, with heavily sclerotized anterior margin and semicircular paramedian membranous area, increased in size posteriorly to tergum VIII. Tergum VIII bears a pair of anteromedial, short, slightly pigmented fan-shaped spots, covered by long black setae. Tergum IX with distinct antecosta, posteromedial sclerite consists of paired small prolonged pigmented spots covered with long, black setae. Tergum X with wide and deep median notch, each lobe bears very specific pointed process at posterior inner margin. Epiproct large, rounded, mushroom-shaped, membranous with pigmented stalk. Cerci large, wide, scoopshaped, bent inward and upward, apex wide, with slight depression. Paraprocts strong, styles shorter than specilla, narrowed and curved sideward near the apex. Specilla long, narrowed to the apex, ending in a sharp point. Ventral vesicle is absent. Female subgenital plate also conspicuous: a triangular, prolonged, reaches anterior margin of sternum IX, with a wide, triangular, unpigmented base delimited by two pairs of small sclerotized sports anterolaterally and laterally, with a pair of sclerotized finger-shaped prolonged lateral lobes shortly bifurcated at the apex.
</paragraph>
<paragraph id="8BCA2C33FFBCFFA0FF77F800FF08FDDE" blockId="5.[151,1437,1533,2026]" lastBlockId="6.[151,1436,151,824]" lastPageId="6" lastPageNumber="552" pageId="5" pageNumber="551">
<emphasis id="B901F021FFBCFFA3FF77F800FEC8F800" bold="true" box="[199,319,1965,1990]" pageId="5" pageNumber="551">Affinities:</emphasis>
According to the shape of processes and pigmented spots on the male abdominal terga, the structure of male epiproct and paraprocts,
<taxonomicName id="4C7557B0FFBCFFA3FDBEF87FFD7BF82C" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[526,652,2002,2026]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="551" phylum="Arthropoda" rank="species" species="adjariae">
<emphasis id="B901F021FFBCFFA3FDBEF87FFD7BF82C" box="[526,652,2002,2026]" italics="true" pageId="5" pageNumber="551">L. adjariae</emphasis>
</taxonomicName>
appears to belong to the
<emphasis id="B901F021FFBCFFA3FC1FF87FFBE1F82C" box="[943,1046,2002,2026]" italics="true" pageId="5" pageNumber="551">hippopus</emphasis>
species group (
<bibRefCitation id="EFE451C2FFBCFFA0FB79F87CFF25FF76" author="Ravizza, C. &amp; Vincon, G." lastPageId="6" lastPageNumber="552" pageId="5" pageNumber="551" pagination="285 - 342" refId="ref7187" refString="Ravizza, C. &amp; Vincon, G. (1998) Les Leuctrides (Plecoptera, Leuctridae) des Alpes. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71, 285 - 342." type="journal article" year="1998">Ravizza &amp; Vinçon 1998</bibRefCitation>
,
<bibRefCitation id="EFE451C2FFBFFFA0FF6EFF3AFE8AFF76" author="Ravizza, C." box="[222,381,151,176]" pageId="6" pageNumber="552" pagination="1 - 42" refId="ref7065" refString="Ravizza, C. (2002) Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species. Lauterbornia, 44, 1 - 42." type="journal article" year="2002">Ravizza 2002</bibRefCitation>
). However, the indistinct, short, fan-shaped processes on tergum VIII, paired pointed process at posterior inner margin of tergum X and other features mentioned in diagnosis, readily distinguish
<taxonomicName id="4C7557B0FFBFFFA0FAADFF10FA6CFF12" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[1309,1435,188,212]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="adjariae">
<emphasis id="B901F021FFBFFFA0FAADFF10FA6CFF12" box="[1309,1435,188,212]" italics="true" pageId="6" pageNumber="552">L. adjariae</emphasis>
</taxonomicName>
males from all congeners. No closely related species are indicated based on external morphology. In the male, a pair of pointed processes of tergum X are unique and they only resemble those of male of Alpine-Carpathian
<taxonomicName id="4C7557B0FFBFFFA0FA35FEA8FE45FE86" authority="Aubert, 1948" authorityName="Aubert" authorityYear="1948" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="autumnalis">
<emphasis id="B901F021FFBFFFA0FA35FEA8FEE3FE86" italics="true" pageId="6" pageNumber="552">L. autumnalis</emphasis>
<bibRefCitation id="EFE451C2FFBFFFA0FEABFE8AFE45FE86" author="Aubert, J." box="[283,434,295,320]" pageId="6" pageNumber="552" pagination="469 - 470" refId="ref6043" refString="Aubert, J. (1948) Un Plecoptere nouveau des Prealpes vaudoises: Leuctra autumnalis n. sp. Mitteilungen des Schweizerischen Entomologischen Gesellschaft, 21, 469 - 470." type="journal article" year="1948">Aubert, 1948</bibRefCitation>
</taxonomicName>
. The short styles of the paraprocts of
<taxonomicName id="4C7557B0FFBFFFA0FCECFE84FC2EFE86" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[860,985,296,320]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="adjariae">
<emphasis id="B901F021FFBFFFA0FCECFE84FC2EFE86" box="[860,985,296,320]" italics="true" pageId="6" pageNumber="552">L. adjariae</emphasis>
</taxonomicName>
are also unusual in the
<emphasis id="B901F021FFBFFFA0FB55FE85FABBFE86" box="[1253,1356,296,320]" italics="true" pageId="6" pageNumber="552">hippopus</emphasis>
group, that are only shared with
<taxonomicName id="4C7557B0FFBFFFA0FE0FFEE0FD87FEA2" box="[447,624,332,356]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="tarnogradski">
<emphasis id="B901F021FFBFFFA0FE0FFEE0FD87FEA2" box="[447,624,332,356]" italics="true" pageId="6" pageNumber="552">L. tarnogradski</emphasis>
</taxonomicName>
and
<emphasis id="B901F021FFBFFFA0FD1CFEE0FCCCFEA2" box="[684,827,332,356]" italics="true" pageId="6" pageNumber="552">
<taxonomicName id="4C7557B0FFBFFFA0FD1CFEE0FCC0FEA2" authorityName="Balinsky" authorityYear="1950" box="[684,823,332,356]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="dispinata">L. dispinata</taxonomicName>
,
</emphasis>
both species with short styles (
<bibRefCitation id="EFE451C2FFBFFFA0FB18FEE1FAACFEA2" author="Zhiltzova, L. A." box="[1192,1371,332,357]" pageId="6" pageNumber="552" pagination="1 - 538" refId="ref7537" refString="Zhiltzova, L. A. (2003) Stoneflies (Plecoptera). Gruppe Euholognata. Fauna of Russia and Neighbouring Countries, New Series, No. 145, 1 (1), 1 - 538. [in Russian]" type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
). The triangular, prolonged subgenital plate of female is not similar to other
<taxonomicName id="4C7557B0FFBFFFA0FC14FEDCFC0BFE4E" authorityName="Stephens" authorityYear="1836" box="[932,1020,369,392]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="B901F021FFBFFFA0FC14FEDCFC0BFE4E" box="[932,1020,369,392]" italics="true" pageId="6" pageNumber="552">Leuctra</emphasis>
</taxonomicName>
species and only remotely resembles members of the
<emphasis id="B901F021FFBFFFA0FEFCFE39FE39FE6A" box="[332,462,404,428]" italics="true" pageId="6" pageNumber="552">brevipennis</emphasis>
sub-group,
<emphasis id="B901F021FFBFFFA0FDE4FE39FD4CFE6A" box="[596,699,404,428]" italics="true" pageId="6" pageNumber="552">hippopus</emphasis>
group, e.g.
<taxonomicName id="4C7557B0FFBFFFA0FCF1FE38FF38FE16" authority="Ravizza &amp; Ravizza Dematteis, 1994" authorityName="Ravizza &amp; Ravizza Dematteis" authorityYear="1994" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="subSpecies" species="vinconi" subSpecies="aubertorum">
<emphasis id="B901F021FFBFFFA0FCF1FE38FBCAFE6A" box="[833,1085,404,428]" italics="true" pageId="6" pageNumber="552">L. vinconi aubertorum</emphasis>
<bibRefCitation id="EFE451C2FFBFFFA0FBF4FE39FF38FE16" author="Ravizza, C. &amp; Ravizza Dematteis, E." pageId="6" pageNumber="552" pagination="37 - 41" refId="ref7137" refString="Ravizza, C. &amp; Ravizza Dematteis, E. (1994): Leuctra vinconi aubertorum, a new subspecies of Leuctra from the Ticino canton, Switzerland (Plecoptera Leuctridae). Mitteilungen desSchweizerischen Entomologischen Gesellschaft, 67 (1 - 2), 37 - 41." type="journal article" year="1994">Ravizza &amp; Ravizza Dematteis, 1994</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="4C7557B0FFBFFFA0FF44FE14FCAAFE16" authority="vinconi Ravizza &amp; Ravizza Dematteis, 1993" authorityName="Ravizza &amp; Ravizza Dematteis" authorityYear="1993" box="[244,861,439,464]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="subSpecies" species="vinconi" subSpecies="vinconi">
<emphasis id="B901F021FFBFFFA0FF44FE14FE4AFE09" box="[244,445,440,464]" italics="true" pageId="6" pageNumber="552">L. vinconi vinconi</emphasis>
<bibRefCitation id="EFE451C2FFBFFFA0FE74FE1AFCAAFE16" author="Ravizza, C. &amp; Ravizza Dematteis, E." box="[452,861,439,464]" pageId="6" pageNumber="552" pagination="41 - 44" refId="ref7098" refString="Ravizza, C. &amp; Ravizza Dematteis, E. (1993) Leuctra vinconi new species of Plecoptera from the Pennine Alps (Italy). Aquatic Insects, 15 (1), 41 - 44." type="journal article" year="1993">Ravizza &amp; Ravizza Dematteis, 1993</bibRefCitation>
</taxonomicName>
, found in the western Alps of
<collectingCountry id="F3626CA3FFBFFFA0FB1DFE1AFAC4FE16" box="[1197,1331,439,464]" name="Switzerland" pageId="6" pageNumber="552">Switzerland</collectingCountry>
and
<collectingCountry id="F3626CA3FFBFFFA0FAD9FE1AFA6BFE16" box="[1385,1436,439,464]" name="Italy" pageId="6" pageNumber="552">Italy</collectingCountry>
(
<bibRefCitation id="EFE451C2FFBFFFA0FF2FFE71FEB1FE32" author="Ravizza, C." box="[159,326,476,501]" pageId="6" pageNumber="552" pagination="1 - 42" refId="ref7065" refString="Ravizza, C. (2002) Atlas of the Italian Leuctridae (Insecta, Plecoptera) with an appendix including Central European species. Lauterbornia, 44, 1 - 42." type="journal article" year="2002">Ravizza, 2002</bibRefCitation>
) and another apterous species
<taxonomicName id="4C7557B0FFBFFFA0FD1DFE70FBBCFE32" authority="Braasch &amp; Joost 1977" authorityName="Braasch &amp; Joost" authorityYear="1977" box="[685,1099,476,501]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="kumanskii">
<emphasis id="B901F021FFBFFFA0FD1DFE70FCB7FE32" box="[685,832,476,500]" italics="true" pageId="6" pageNumber="552">L. kumanskii</emphasis>
Braasch &amp; Joost 1977
</taxonomicName>
from the Balkan Mountains,
<collectingCountry id="F3626CA3FFBFFFA0FF27FE52FF0CFDDE" box="[151,251,511,536]" name="Bulgaria" pageId="6" pageNumber="552">Bulgaria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F7FB8FFBFFFA0FF77FD89FCD7FCD3" pageId="6" pageNumber="552" type="distribution">
<paragraph id="8BCA2C33FFBFFFA0FF77FD89FCD7FCD3" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="552">
<emphasis id="B901F021FFBFFFA0FF77FD89FE97FDFB" bold="true" box="[199,352,548,573]" pageId="6" pageNumber="552">Distribution.</emphasis>
<taxonomicName id="4C7557B0FFBFFFA0FEDAFD88FDDEFDFA" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[362,553,548,572]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="adjariae">
<emphasis id="B901F021FFBFFFA0FEDAFD88FDDEFDFA" box="[362,553,548,572]" italics="true" pageId="6" pageNumber="552">Leuctra adjariae</emphasis>
</taxonomicName>
was collected at a single locality along the Kintrishi River near the town of Kobulety in
<collectingRegion id="49B1E2D1FFBFFFA0FE99FDEAFE77FDA6" box="[297,384,583,608]" country="Georgia" name="Ajaria" pageId="6" pageNumber="552">Adjaria</collectingRegion>
, in southwestern
<collectingCountry id="F3626CA3FFBFFFA0FDFDFDEAFD5DFDA6" box="[589,682,583,608]" name="Georgia" pageId="6" pageNumber="552">Georgia</collectingCountry>
. The Autonomous Republic
<collectingRegion id="49B1E2D1FFBFFFA0FC48FDEAFBB8FDA6" box="[1016,1103,583,608]" country="Georgia" name="Ajaria" pageId="6" pageNumber="552">Adjaria</collectingRegion>
, a historical, geographic and political-administrative region of
<collectingCountry id="F3626CA3FFBFFFA0FDA2FDC1FD98FD43" box="[530,623,620,645]" name="Georgia" pageId="6" pageNumber="552">Georgia</collectingCountry>
, borders
<collectingCountry id="F3626CA3FFBFFFA0FD65FDC1FCD2FD43" box="[725,805,620,645]" name="Turkey" pageId="6" pageNumber="552">Turkey</collectingCountry>
to the south and the eastern edge of the Black Sea (
<figureCitation id="134E30B6FFBFFFA0FADFFDC1FF40FD6E" captionStart="FIGURES 2528" captionStartId="10.[151,264,1178,1201]" captionTargetBox="[150,1436,197,1151]" captionTargetId="figure@10.[150,1436,193,1157]" captionTargetPageId="10" captionText="FIGURES 2528. 25. Collecting place of Plecoptera in Adjaria, Georgia (black dot). Square shows the nearest collecting site of Capnioneura gouanerae in Artvin, Turkey. 26. Female of Leuctra adjariae sp. n. on the snow. 27 &amp; 28. The Kintrishi River, Meskheti Range, the type locality of Leuctra adjariae sp. n. and Leuctra georgiae sp. n. Photo by D. Palatov." httpUri="https://zenodo.org/record/2640295/files/figure.png" pageId="6" pageNumber="552">Fig. 25</figureCitation>
). The Kintrishi River drains the western slopes of Meskheti Range (Lesser Caucasus) into the Black Sea. At the
<typeStatus id="54CE9291FFBFFFA0FF27FD18FF30FD0B" box="[151,199,693,717]" pageId="6" pageNumber="552">type</typeStatus>
locality, where the altitude does not exceed
<quantity id="4C8D81D6FFBFFFA0FD0BFD19FCC1FD0A" box="[699,822,692,716]" metricMagnitude="2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="6" pageNumber="552" unit="m" value="185.0" valueMax="190.0" valueMin="180.0">180-190 m</quantity>
a.s.l, the Kintrishi River flows through a canyon, with a width
<quantity id="4C8D81D6FFBFFFA0FF40FD7AFEC2FD29" box="[240,309,727,752]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="552" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">
5-
<specimenCount id="9D73E7BAFFBFFFA0FEB6FD7AFEC2FD29" box="[262,309,727,752]" pageId="6" pageNumber="552" type="generic">6 m</specimenCount>
</quantity>
. The water level increased
<quantity id="4C8D81D6FFBFFFA0FDD2FD75FD55FD29" box="[610,674,728,752]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="552" unit="m" value="0.3">0.3 m</quantity>
during flooding, but was clear, flowing at
<quantity id="4C8D81D6FFBFFFA0FBCBFD75FB1FFD36" box="[1147,1256,728,752]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="6" pageNumber="552" unit="m" value="0.6" valueMax="0.9" valueMin="0.3">0.3-0.9 m</quantity>
/s (
<figureCitation id="134E30B6FFBFFFA0FAB8FD7AFA67FD36" box="[1288,1424,727,752]" captionStart="FIGURES 2528" captionStartId="10.[151,264,1178,1201]" captionTargetBox="[150,1436,197,1151]" captionTargetId="figure@10.[150,1436,193,1157]" captionTargetPageId="10" captionText="FIGURES 2528. 25. Collecting place of Plecoptera in Adjaria, Georgia (black dot). Square shows the nearest collecting site of Capnioneura gouanerae in Artvin, Turkey. 26. Female of Leuctra adjariae sp. n. on the snow. 27 &amp; 28. The Kintrishi River, Meskheti Range, the type locality of Leuctra adjariae sp. n. and Leuctra georgiae sp. n. Photo by D. Palatov." httpUri="https://zenodo.org/record/2640295/files/figure.png" pageId="6" pageNumber="552">Figs. 27, 28</figureCitation>
). Adults of
<taxonomicName id="4C7557B0FFBFFFA0FEB7FD50FE74FCD2" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2019" box="[263,387,764,788]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="552" phylum="Arthropoda" rank="species" species="adjariae">
<emphasis id="B901F021FFBFFFA0FEB7FD50FE74FCD2" box="[263,387,764,788]" italics="true" pageId="6" pageNumber="552">L. adjariae</emphasis>
</taxonomicName>
were walking on the snow (
<figureCitation id="134E30B6FFBFFFA0FD0FFD51FCE3FCD3" box="[703,788,764,789]" captionStart="FIGURES 2528" captionStartId="10.[151,264,1178,1201]" captionTargetBox="[150,1436,197,1151]" captionTargetId="figure@10.[150,1436,193,1157]" captionTargetPageId="10" captionText="FIGURES 2528. 25. Collecting place of Plecoptera in Adjaria, Georgia (black dot). Square shows the nearest collecting site of Capnioneura gouanerae in Artvin, Turkey. 26. Female of Leuctra adjariae sp. n. on the snow. 27 &amp; 28. The Kintrishi River, Meskheti Range, the type locality of Leuctra adjariae sp. n. and Leuctra georgiae sp. n. Photo by D. Palatov." httpUri="https://zenodo.org/record/2640295/files/figure.png" pageId="6" pageNumber="552">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F7FB8FFBFFFA0FF77FCB2FB83FCF1" box="[199,1140,799,824]" pageId="6" pageNumber="552" type="etymology">
<paragraph id="8BCA2C33FFBFFFA0FF77FCB2FB83FCF1" blockId="6.[151,1436,151,824]" box="[199,1140,799,824]" pageId="6" pageNumber="552">
<emphasis id="B901F021FFBFFFA0FF77FCB2FEB9FCFE" bold="true" box="[199,334,799,824]" pageId="6" pageNumber="552">Etymology.</emphasis>
This species is named after
<collectingRegion id="49B1E2D1FFBFFFA0FD3BFCB2FD15FCFE" box="[651,738,799,824]" country="Georgia" name="Ajaria" pageId="6" pageNumber="552">Adjaria</collectingRegion>
, region of
<collectingCountry id="F3626CA3FFBFFFA0FCE9FCB2FC41FCFE" box="[857,950,799,824]" name="Georgia" pageId="6" pageNumber="552">Georgia</collectingCountry>
, where it occurs.
</paragraph>
</subSubSection>
</treatment>
</document>