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<document id="EDD7DAD094AA7606C94D57F64EAA1164" ID-DOI="10.11646/zootaxa.3948.1.8" ID-GBIF-Dataset="7e1f0a0c-02db-4e5c-a4eb-02fee3d3c1e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="244386" ID-ZooBank="50DC0370-C8C7-4316-93E4-D7424AB22291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461043375627" checkinUser="plazi" docAuthor="Gasca-Álvarez, Héctor Jaime &amp; Deloya, Cuauhtémoc" docDate="2015" docId="03D9AF4E160D620CFF3CFC8F1176FD94" docLanguage="en" docName="zt03948p136.pdf" docOrigin="Zootaxa 3948 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cotinis patricialvarezae Gasca-Alvarez &amp; Deloya, new species" docType="treatment" docVersion="10" lastPageNumber="130" masterDocId="FFE0D736160F6209FFABFFE81365FF83" masterDocTitle="A new species and first record of Cotinis Burmeister (Coleoptera: Scarabaeidae: Cetoniinae) for Venezuela" masterLastPageNumber="136" masterPageNumber="125" pageNumber="127" updateTime="1698576161018" updateUser="plazi">
<mods:mods id="1E3F312BCF22648D8386D280E18F195B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8C5A21087C522C64933D9BB2A485AF03">
<mods:title id="3F6D0EFEB4D722E2239764CE285B16BB">A new species and first record of Cotinis Burmeister (Coleoptera: Scarabaeidae: Cetoniinae) for Venezuela</mods:title>
</mods:titleInfo>
<mods:name id="3CE5EAD50E9C1D02D55BFB9A157776EE" type="personal">
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<mods:roleTerm id="D2523E18E3D4D2BA7F6892E1680D5970">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="188C426A46CC85A3DB46D9D3584ADD99">Gasca-Álvarez, Héctor Jaime</mods:namePart>
</mods:name>
<mods:name id="6C43255B13443D91A14B3FE9953F816B" type="personal">
<mods:role id="020C816CEE9EEB505E5288242275DF2B">
<mods:roleTerm id="F09DF335D7EE2027FEE8E2A73295BF2B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBE244622D3B92CDF86094E7C17E23D2">Deloya, Cuauhtémoc</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A76600A09D0F8299CBB37A5E61E9A87">text</mods:typeOfResource>
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<mods:title id="073ED1BE831A4EE0158ED58C262A5BEC">Zootaxa</mods:title>
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<mods:part id="4CC2E8A875E57E3BEE4A6C1C9F19E6A4">
<mods:date id="16345027BDA9F59B76EF171CF2BD2913">2015</mods:date>
<mods:detail id="CC1EC3B118A42935DCF9EBCDE46E797B" type="volume">
<mods:number id="4C36E8AAA2E743ACAA750C9E58332ECC">3948</mods:number>
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<mods:classification id="87A07D1635A895B830ADB5052FDFA973">journal article</mods:classification>
<mods:identifier id="4649B1F60A31FD504879B3B52638B554" type="DOI">10.11646/zootaxa.3948.1.8</mods:identifier>
<mods:identifier id="D9A9583F87CD51D6943D8EB7FC026FB3" type="GBIF-Dataset">7e1f0a0c-02db-4e5c-a4eb-02fee3d3c1e1</mods:identifier>
<mods:identifier id="FE12F7DFC0C768F437C2EA1DE8AA0766" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C79DD819AAEE1C52ACA7D6B561E44BFF" type="Zenodo-Dep">244386</mods:identifier>
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</mods:mods>
<treatment id="03D9AF4E160D620CFF3CFC8F1176FD94" ID-DOI="http://doi.org/10.5281/zenodo.5694223" ID-GBIF-Taxon="119610083" ID-Zenodo-Dep="5694223" LSID="urn:lsid:plazi:treatment:03D9AF4E160D620CFF3CFC8F1176FD94" httpUri="http://treatment.plazi.org/id/03D9AF4E160D620CFF3CFC8F1176FD94" lastPageId="5" lastPageNumber="130" pageId="2" pageNumber="127">
<subSubSection id="C36A4DD3160D620BFF3CFC8F1277FC20" pageId="2" pageNumber="127" type="nomenclature">
<paragraph id="8BCF1E58160D620BFF3CFC8F10EAFC01" blockId="2.[151,911,871,931]" box="[151,911,871,898]" pageId="2" pageNumber="127">
<heading id="D087A934160D620BFF3CFC8F10EAFC01" bold="true" box="[151,911,871,898]" fontSize="11" level="1" pageId="2" pageNumber="127" reason="1">
<emphasis id="B904C24A160D620BFF3CFC8F10EAFC01" bold="true" box="[151,911,871,898]" pageId="2" pageNumber="127">
<taxonomicName id="4C7065DB160D620BFF3CFC8F1197FC01" authority="Gasca-Alvarez &amp; Deloya" authorityName="Gasca-Alvarez &amp; Deloya" box="[151,754,871,898]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="patricialvarezae" status="sp. nov.">
<emphasis id="B904C24A160D620BFF3CFC8F12D0FC02" bold="true" box="[151,437,871,897]" italics="true" pageId="2" pageNumber="127">Cotinis patricialvarezae</emphasis>
Gasca-Álvarez &amp; Deloya
</taxonomicName>
,
<taxonomicNameLabel id="A2377F31160D620BFD54FC8010EAFC01" box="[767,911,872,898]" pageId="2" pageNumber="127" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E58160D620BFF3CFC621277FC20" blockId="2.[151,911,871,931]" box="[151,274,906,931]" pageId="2" pageNumber="127">
(
<figureCitation id="134B02DD160D620BFF34FC62126CFC20" box="[159,265,906,931]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 7" captionStartId-0="3.[151,264,1923,1946]" captionStartId-1="4.[151,264,1953,1976]" captionTargetBox-0="[157,1413,468,1877]" captionTargetBox-1="[303,1282,236,1894]" captionTargetId-0="figure@3.[151,1436,411,1899]" captionTargetId-1="figure@4.[186,1401,193,1932]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 5. Cotinis patricialvarezae new species, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum." captionText-1="FIGURES 6 – 7. Cotinis patricialvarezae new species, 6. Parameres, frontal view; 7. Parameres, lateral view. Scale bar = 0.6 mm." httpUri-0="https://zenodo.org/record/244387/files/figure.png" httpUri-1="https://zenodo.org/record/244388/files/figure.png" pageId="2" pageNumber="127">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A4DD3160D620BFF3CFC3A12B3F95C" pageId="2" pageNumber="127" type="materials_examined">
<paragraph id="8BCF1E58160D620BFF3CFC3A1615FBB0" blockId="2.[151,1437,978,2011]" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF3CFC3A122CFC68" bold="true" box="[151,329,978,1003]" pageId="2" pageNumber="127">
<typeStatus id="54CBA0FA160D620BFF3CFC3A13B7FC68" box="[151,210,978,1003]" pageId="2" pageNumber="127">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54CBA0FA160D620BFEFEFC3A12D8FC68" box="[341,445,978,1003]" pageId="2" pageNumber="127" type="holotype">Holotype</typeStatus>
male labeled “
<collectingCountry id="F3675EC8160D620BFDDAFC3B107BFC68" box="[625,798,979,1003]" name="Venezuela" pageId="2" pageNumber="127">VENEZUELA</collectingCountry>
. State of Lara, Barquisemeto.
<date id="FFCE3898160D620BFB23FC3A17BCFC68" box="[1160,1241,978,1003]" pageId="2" pageNumber="127" value="1968-05-09">9-V-68</date>
. José M. Osorio Coll”. Single male
<typeStatus id="54CBA0FA160D620BFEC5FC1F12ABFB8C" box="[366,462,1015,1039]" pageId="2" pageNumber="127" type="paratype">paratype</typeStatus>
labeled “
<collectingCountry id="F3675EC8160D620BFD9CFC1F1181FB8C" box="[567,740,1015,1039]" name="Venezuela" pageId="2" pageNumber="127">VENEZUELA</collectingCountry>
. Dept. Lara, Aqua Viva (Cabudare).
<date id="FFCE3898160D620BFBD5FC1F17B8FB8C" box="[1150,1245,1015,1039]" pageId="2" pageNumber="127" value="1982-04">IV-1982</date>
, J. S. González”.
<typeStatus id="54CBA0FA160D620BFF3CFBF313BFFBB0" box="[151,218,1051,1075]" pageId="2" pageNumber="127">Types</typeStatus>
deposited at the Florida State Collection of Arthropods (Gainesville, Florida,
<collectingCountry id="F3675EC8160D620BFBEFFBF21600FBB0" box="[1092,1381,1050,1075]" name="United States of America" pageId="2" pageNumber="127">United States of America</collectingCountry>
).
</paragraph>
<paragraph id="8BCF1E58160D620BFF6CFBD612B3F95C" blockId="2.[151,1437,978,2011]" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF6CFBD6125EFBD4" bold="true" box="[199,315,1086,1111]" pageId="2" pageNumber="127">
<typeStatus id="54CBA0FA160D620BFF6CFBD61253FBD4" box="[199,310,1086,1111]" pageId="2" pageNumber="127" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="134B02DD160D620BFE26FBD6129FFBD4" box="[397,506,1086,1111]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[157,1413,468,1877]" captionTargetId="figure@3.[151,1436,411,1899]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cotinis patricialvarezae new species, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum." httpUri="https://zenodo.org/record/244387/files/figure.png" pageId="2" pageNumber="127">Figs. 1–2</figureCitation>
). Length
<quantity id="4C88B3BD160D620BFDC3FBD611B5FBD5" box="[616,720,1086,1111]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="2" pageNumber="127" unit="mm" value="23.6">23.6 mm</quantity>
; width across humeri
<quantity id="4C88B3BD160D620BFC60FBD71756FBD5" box="[971,1075,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="127" unit="mm" value="13.1">13.1 mm</quantity>
.
<emphasis id="B904C24A160D620BFB94FBD617E8FBD4" bold="true" box="[1087,1165,1086,1111]" pageId="2" pageNumber="127">Color:</emphasis>
Head bright green with some red-brown reflections. Pronotum bright green. Elytra opaque red-brown with green reflections. Ventral region and legs glossy greenish brown. Pygidium bright red-brown.
<emphasis id="B904C24A160D620BFCF2FB6E10C7FB1C" bold="true" box="[857,930,1158,1183]" pageId="2" pageNumber="127">Head:</emphasis>
Surface rugopunctate, punctures strong and sparse; sparsely setose with sparse, small, pale yellow setae. Clypeal projection developed, triangular, rounded at apex (
<figureCitation id="134B02DD160D620BFF70FB261245FB64" box="[219,288,1230,1255]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[157,1413,468,1877]" captionTargetId="figure@3.[151,1436,411,1899]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cotinis patricialvarezae new species, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum." httpUri="https://zenodo.org/record/244387/files/figure.png" pageId="2" pageNumber="127">Fig. 3</figureCitation>
). Frontoclypeal depression with conspicuous medial ridge extending to the base of clypeal projection. Frontal projection straight, apex expanded apically and truncate, partially free, fused 80% to head on basal third (
<figureCitation id="134B02DD160D620BFF34FAFE1380FAAC" box="[159,229,1302,1327]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[157,1413,468,1877]" captionTargetId="figure@3.[151,1436,411,1899]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cotinis patricialvarezae new species, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum." httpUri="https://zenodo.org/record/244387/files/figure.png" pageId="2" pageNumber="127">Fig. 4</figureCitation>
). Antenna with 10 antennomeres.
<emphasis id="B904C24A160D620BFDC2FAFE118EFAAC" bold="true" box="[617,747,1302,1327]" pageId="2" pageNumber="127">Pronotum:</emphasis>
Surface finely punctate with a shallow, median, longitudinal impression; disc smooth, punctures becoming much larger and strong impressed laterally. Marginal bead complete at sides. Apical margin with rounded protuberance at middle. Dorsal surface of mesepimera punctate, punctures moderate in size.
<emphasis id="B904C24A160D620BFEF0FA6A12D5FA18" bold="true" box="[347,432,1410,1435]" pageId="2" pageNumber="127">Elytra:</emphasis>
Surface smooth, finely punctate with 2 slightly marked, longitudinal costae converging at apical umbone. Humerus prominent.
<emphasis id="B904C24A160D620BFD9EFA4E11D7FA3C" bold="true" box="[565,690,1446,1471]" pageId="2" pageNumber="127">Pygidium:</emphasis>
Base and angles rugose, with transverse strigulations, unicolorous, without setae; apex rounded, surface slightly convex in lateral view.
<emphasis id="B904C24A160D620BFC0CFA22108DFA60" bold="true" box="[935,1000,1482,1507]" pageId="2" pageNumber="127">Legs:</emphasis>
Protibiae tridentate, basal tooth short, shorter than second tooth; apical and median teeth closer to each other than to basal tooth. Metatibial spurs slender, with acute apices.
<emphasis id="B904C24A160D620BFEDFF9FA12A8F9A8" bold="true" box="[372,461,1554,1579]" pageId="2" pageNumber="127">Venter:</emphasis>
Mesometasternal projection broad, expanded apically; sides weakly concave; apex rounded (
<figureCitation id="134B02DD160D620BFEA9F9DE1223F9CC" box="[258,326,1590,1615]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[157,1413,468,1877]" captionTargetId="figure@3.[151,1436,411,1899]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cotinis patricialvarezae new species, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum." httpUri="https://zenodo.org/record/244387/files/figure.png" pageId="2" pageNumber="127">Fig. 5</figureCitation>
). Metasternum with central longitudinal line, with sparse, small punctures at center, sides with fine impressed punctures; pale yellow setae short.
<emphasis id="B904C24A160D620BFD3DF9B21045F9F0" bold="true" box="[662,800,1626,1651]" pageId="2" pageNumber="127">Parameres:</emphasis>
Elongate; narrow at middle, apex rounded; anterocentral margins slightly curved, anterolateral projections developed, former into lobes directed at right angle. Sides with a conspicuous, rounded tooth under the anterolateral projections (
<figureCitation id="134B02DD160D620BFCCDF94A10B1F938" box="[870,980,1698,1723]" captionStart="FIGURES 6 – 7" captionStartId="4.[151,264,1953,1976]" captionTargetBox="[303,1282,236,1894]" captionTargetId="figure@4.[186,1401,193,1932]" captionTargetPageId="4" captionText="FIGURES 6 – 7. Cotinis patricialvarezae new species, 6. Parameres, frontal view; 7. Parameres, lateral view. Scale bar = 0.6 mm." httpUri="https://zenodo.org/record/244388/files/figure.png" pageId="2" pageNumber="127">Figs. 6–7</figureCitation>
). Internal sac with sclerotized area dark, bearing a single, stout spine.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DD3160D620BFF6CF902176EF8C9" pageId="2" pageNumber="127" type="description">
<paragraph id="8BCF1E58160D620BFF6CF90212C7F880" blockId="2.[151,1437,978,2011]" box="[199,418,1770,1795]" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF6CF9021240F880" bold="true" box="[199,293,1770,1795]" pageId="2" pageNumber="127">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCF1E58160D620BFF6CF8E6176EF8C9" blockId="2.[151,1437,978,2011]" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF6CF8E6125EF8A4" bold="true" box="[199,315,1806,1831]" pageId="2" pageNumber="127">Variation.</emphasis>
Male (1
<typeStatus id="54CBA0FA160D620BFE01F8E71175F8A4" box="[426,528,1807,1831]" pageId="2" pageNumber="127" type="paratype">paratype</typeStatus>
). Length
<quantity id="4C88B3BD160D620BFDD7F8E71180F8A5" box="[636,741,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="2" pageNumber="127" unit="mm" value="21.1">21.1 mm</quantity>
; width across humeri
<quantity id="4C88B3BD160D620BFC48F8E7172EF8A5" box="[995,1099,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="2" pageNumber="127" unit="mm" value="11.9">11.9 mm</quantity>
. The
<typeStatus id="54CBA0FA160D620BFB27F8E71789F8A4" box="[1164,1260,1807,1831]" pageId="2" pageNumber="127" type="paratype">paratype</typeStatus>
does not differ significantly from the
<typeStatus id="54CBA0FA160D620BFE3BF8DA1297F8C8" box="[400,498,1842,1867]" pageId="2" pageNumber="127" type="holotype">holotype</typeStatus>
except the pygidium is slightly smooth at apex.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DD3160D620BFF6CF8BE16F7F811" pageId="2" pageNumber="127" type="diagnosis">
<paragraph id="8BCF1E58160D620BFF6CF8BE16F7F811" blockId="2.[151,1437,978,2011]" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF6CF8BE1224F8EC" bold="true" box="[199,321,1878,1903]" pageId="2" pageNumber="127">Diagnosis.</emphasis>
<taxonomicName id="4C7065DB160D620BFEE1F8BF113FF8ED" box="[330,602,1878,1902]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160D620BFEE1F8BF113FF8ED" box="[330,602,1878,1902]" italics="true" pageId="2" pageNumber="127">Cotinis patricialvarezae</emphasis>
</taxonomicName>
can be distinguished by the violet color of elytra with bright reflections, triangular shape of the clypeal projection, parameres elongate (
<figureCitation id="134B02DD160D620BFCF2F89210A0F810" box="[857,965,1914,1939]" captionStart="FIGURES 6 – 7" captionStartId="4.[151,264,1953,1976]" captionTargetBox="[303,1282,236,1894]" captionTargetId="figure@4.[186,1401,193,1932]" captionTargetPageId="4" captionText="FIGURES 6 – 7. Cotinis patricialvarezae new species, 6. Parameres, frontal view; 7. Parameres, lateral view. Scale bar = 0.6 mm." httpUri="https://zenodo.org/record/244388/files/figure.png" pageId="2" pageNumber="127">Figs. 6–7</figureCitation>
), and the single spine on the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DD3160D620AFF6CF87617F6FEBC" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127" type="distribution">
<paragraph id="8BCF1E58160D620AFF6CF87617F6FEBC" blockId="2.[151,1437,978,2011]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127">
<emphasis id="B904C24A160D620BFF6CF876123CF834" bold="true" box="[199,345,1950,1975]" pageId="2" pageNumber="127">Distribution</emphasis>
.
<taxonomicName id="4C7065DB160D620BFEC2F877111FF835" box="[361,634,1950,1974]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160D620BFEC2F877111FF835" box="[361,634,1950,1974]" italics="true" pageId="2" pageNumber="127">Cotinis patricialvarezae</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54CBA0FA160D620BFC30F87710AEF834" box="[923,971,1951,1975]" pageId="2" pageNumber="127">type</typeStatus>
locality in Lara,
<collectingCountry id="F3675EC8160D620BFB3EF876166DF834" box="[1173,1288,1950,1975]" name="Venezuela" pageId="2" pageNumber="127">Venezuela</collectingCountry>
(
<figureCitation id="134B02DD160D620BFAB2F876163CF834" box="[1305,1369,1950,1975]" captionStart="FIGURE 8" captionStartId="5.[151,250,2005,2027]" captionText="FIGURE 8. Distribution records of Cotinis patricalvarezae in Venezuela." pageId="2" pageNumber="127">Fig 8</figureCitation>
). The habitat is lowland in which the vegetation is composed of thorny shrubs (
<bibRefCitation id="EFE163A9160D620BFC75F82A1632F858" author="Huber" box="[990,1367,1986,2011]" pageId="2" pageNumber="127" refString="Huber, O. &amp; Oliveira-Miranda, M. A. (2010) Ambientes terrestres de Venezuela. In: Rodriguez, J. P., Rojas-Suarez, F. &amp; Giraldo-Hernandez, D. (Eds.), Libro Rojo de los Ecosistemas Terrestres de Venezuela. Provita, Shell Venezuela, Lenovo (Venezuela), Caracas, Venezuela, pp. 29 - 89." type="book chapter" year="2010">Huber &amp; Oliveira-Miranda 2010</bibRefCitation>
). This species represents the first record of
<taxonomicName id="4C7065DB160C620AFD9BFF7011E4FF2C" box="[560,641,152,175]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B904C24A160C620AFD9BFF7011E4FF2C" box="[560,641,152,175]" italics="true" pageId="3" pageNumber="128">Cotinis</emphasis>
</taxonomicName>
for
<collectingCountry id="F3675EC8160C620AFD1BFF7F1046FF33" box="[688,803,151,176]" name="Venezuela" pageId="3" pageNumber="128">Venezuela</collectingCountry>
and is the furthest southeastern occurrence of the genus, expanding its distributional range by approximately
<quantity id="4C88B3BD160C620AFD52FF541030FF56" box="[761,853,188,213]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="128" unit="km" value="600.0">600 km</quantity>
. The geographical distribution of the new species corresponds to the continuity of the Colombian Andes mountain range, which extends into
<collectingCountry id="F3675EC8160C620AFB39FF371660FF7B" box="[1170,1285,223,248]" name="Venezuela" pageId="3" pageNumber="128">Venezuela</collectingCountry>
ending at the Merida Cordillera.
<bibRefCitation id="EFE163A9160C620AFED9FEEC1298FE9F" author="Solis" box="[370,509,260,285]" pageId="3" pageNumber="128" refString="Solis A. (2004) Escarabajos Fruteros de Costa Rica. Instituto Nacional de Biodiversidad INBIO, Editorial INBIO, Costa Rica, 237 pp." type="book" year="2004">Solís (2004)</bibRefCitation>
reported
<taxonomicName id="4C7065DB160C620AFDC7FEEC11B4FE9F" box="[620,721,260,284]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="lebasi">
<emphasis id="B904C24A160C620AFDC7FEEC11B4FE9F" box="[620,721,260,284]" italics="true" pageId="3" pageNumber="128">C. lebasi</emphasis>
</taxonomicName>
from
<collectingCountry id="F3675EC8160C620AFCB2FEEC1015FE9F" box="[793,880,260,284]" name="Guyana" pageId="3" pageNumber="128">Guyana</collectingCountry>
but without any evidence to support this record. Due this species is so far out of their normal range, we consider the record to be erroneous.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DD3160C620CFF6CFEA41176FD94" lastPageId="5" lastPageNumber="130" pageId="3" pageNumber="128" type="etymology">
<paragraph id="8BCF1E58160C620AFF6CFEA417B8FEE6" blockId="3.[151,1437,151,357]" box="[199,1245,332,357]" pageId="3" pageNumber="128">
<emphasis id="B904C24A160C620AFF6CFEA4122BFEE6" bold="true" box="[199,334,332,357]" pageId="3" pageNumber="128">Etymology.</emphasis>
This species is dedicated to Patricia Álvarez de Gasca, mother of the first author.
</paragraph>
<caption id="DF0F4ED0160C620AFF3CF86B11FCF83B" httpUri="https://zenodo.org/record/244387/files/figure.png" pageId="3" pageNumber="128" targetBox="[157,1413,468,1877]" targetPageId="3">
<paragraph id="8BCF1E58160C620AFF3CF86B11FCF83B" blockId="3.[151,1436,1923,1976]" pageId="3" pageNumber="128">
<emphasis id="B904C24A160C620AFF3CF86B1259F81A" bold="true" box="[151,316,1923,1946]" pageId="3" pageNumber="128">FIGURES 1–5.</emphasis>
<taxonomicName id="4C7065DB160C620AFEEEF86C1159F819" box="[325,572,1924,1946]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="patricialvarezae" status="sp. nov.">
<emphasis id="B904C24A160C620AFEEEF86C1159F819" box="[325,572,1924,1946]" italics="true" pageId="3" pageNumber="128">Cotinis patricialvarezae</emphasis>
</taxonomicName>
<emphasis id="B904C24A160C620AFDEDF86C11A0F819" bold="true" box="[582,709,1924,1946]" pageId="3" pageNumber="128">
<taxonomicNameLabel id="A2377F31160C620AFDEDF86C11A0F819" box="[582,709,1924,1946]" pageId="3" pageNumber="128" rank="species">new species</taxonomicNameLabel>
</emphasis>
, 1. Habitus, dorsal view; 2. Habitus, lateral view; 3. Clypeus, frontal view; 4. Head, lateral view; 5. Meso-metasternum.
</paragraph>
</caption>
<caption id="DF0F4ED0160B620DFF3CF84913A3F855" httpUri="https://zenodo.org/record/244388/files/figure.png" pageId="4" pageNumber="129" targetBox="[303,1282,236,1894]" targetPageId="4">
<paragraph id="8BCF1E58160B620DFF3CF84913A3F855" blockId="4.[151,1436,1953,2006]" pageId="4" pageNumber="129">
<emphasis id="B904C24A160B620DFF3CF849125EF835" bold="true" box="[151,315,1953,1976]" pageId="4" pageNumber="129">FIGURES 6–7.</emphasis>
<taxonomicName id="4C7065DB160B620DFEE8F84A115CF834" box="[323,569,1953,1975]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="patricialvarezae" status="sp. nov.">
<emphasis id="B904C24A160B620DFEE8F84A115CF834" box="[323,569,1953,1975]" italics="true" pageId="4" pageNumber="129">Cotinis patricialvarezae</emphasis>
</taxonomicName>
<emphasis id="B904C24A160B620DFDEBF84A11DBF834" bold="true" box="[576,702,1953,1975]" pageId="4" pageNumber="129">
<taxonomicNameLabel id="A2377F31160B620DFDEBF84A11DBF834" box="[576,702,1953,1975]" pageId="4" pageNumber="129" rank="species">new species</taxonomicNameLabel>
</emphasis>
, 6. Parameres, frontal view; 7. Parameres, lateral view. Scale bar = 0.6 mm.
</paragraph>
</caption>
<paragraph id="8BCF1E58160A620CFF6CFF7F1061FE2F" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="130">
<emphasis id="B904C24A160A620CFF6CFF7F12D8FF33" bold="true" box="[199,445,151,176]" pageId="5" pageNumber="130">Taxonomic remarks.</emphasis>
<bibRefCitation id="EFE163A9160A620CFE6FFF7F111AFF33" author="Goodrich" box="[452,639,151,176]" pageId="5" pageNumber="130" refString="Goodrich, M. A. (1966) A revision of the genus Cotinis (Coleoptera: Scarabaeidae). Annals of the Entomological Society of America, 59, 550 - 568. http: // dx. doi. org / 10.1093 / aesa / 59.3.550" type="journal article" year="1966">Goodrich (1966)</bibRefCitation>
suggested that
<taxonomicName id="4C7065DB160A620CFC87FF7017D9FF33" authority="Gory &amp; Percheron" authorityName="Gory &amp; Percheron" box="[812,1212,151,176]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="barthelemyi">
<emphasis id="B904C24A160A620CFC87FF7010B7FF33" box="[812,978,152,176]" italics="true" pageId="5" pageNumber="130">C. barthelemyi</emphasis>
(Gory &amp; Percheron)
</taxonomicName>
is closely related to
<taxonomicName id="4C7065DB160A620CFF3CFF54120EFF57" authority="Bates" authorityName="Bates" box="[151,363,188,212]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="laticornis">
<emphasis id="B904C24A160A620CFF3CFF541246FF57" box="[151,291,188,212]" italics="true" pageId="5" pageNumber="130">C. laticornis</emphasis>
Bates
</taxonomicName>
, but he did not provide information to explain this relationship. Our character analysis indicates that
<taxonomicName id="4C7065DB160A620CFF6CFF0812FBFF7B" box="[199,414,224,248]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFF6CFF0812FBFF7B" box="[199,414,224,248]" italics="true" pageId="5" pageNumber="130">C. patricialvarezae</emphasis>
</taxonomicName>
is related to
<taxonomicName id="4C7065DB160A620CFD86FF0811B7FF7B" box="[557,722,224,248]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="barthelemyi">
<emphasis id="B904C24A160A620CFD86FF0811B7FF7B" box="[557,722,224,248]" italics="true" pageId="5" pageNumber="130">C. barthelemyi</emphasis>
</taxonomicName>
(
<figureCitation id="134B02DD160A620CFD49FF37103EFF7B" box="[738,859,223,248]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1656,1679]" captionTargetBox="[159,1431,260,1555]" captionTargetId="figure@6.[151,1436,243,1586]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Cotinis barthelemyi. 9. Habitus, dorsal view; 10. Habitus, lateral view; 11. Clypeus, frontal view; 12. Head, lateral view; 13. Meso-metasternum." httpUri="https://zenodo.org/record/244389/files/figure.png" pageId="5" pageNumber="130">Figs. 9–10</figureCitation>
) base on the shape and color of the body, and shape of the frontal projection. They differ by the form of the clypeal projection, which is subtrapezoidal in
<taxonomicName id="4C7065DB160A620CFA29FEEC122BFEC3" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFA29FEEC122BFEC3" italics="true" pageId="5" pageNumber="130">C. patricialvarezae</emphasis>
</taxonomicName>
and parallel-sided in
<taxonomicName id="4C7065DB160A620CFDE2FEC01197FEC3" box="[585,754,296,320]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="barthelemyi">
<emphasis id="B904C24A160A620CFDE2FEC01197FEC3" box="[585,754,296,320]" italics="true" pageId="5" pageNumber="130">C. barthelemyi</emphasis>
</taxonomicName>
(
<figureCitation id="134B02DD160A620CFCA8FECF1033FEC3" box="[771,854,295,320]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1656,1679]" captionTargetBox="[159,1431,260,1555]" captionTargetId="figure@6.[151,1436,243,1586]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Cotinis barthelemyi. 9. Habitus, dorsal view; 10. Habitus, lateral view; 11. Clypeus, frontal view; 12. Head, lateral view; 13. Meso-metasternum." httpUri="https://zenodo.org/record/244389/files/figure.png" pageId="5" pageNumber="130">Fig. 11</figureCitation>
); the greater fusion of the frontal projection to the head in
<taxonomicName id="4C7065DB160A620CFF51FEA412B2FEE7" box="[250,471,332,356]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFF51FEA412B2FEE7" box="[250,471,332,356]" italics="true" pageId="5" pageNumber="130">C. patricialvarezae</emphasis>
</taxonomicName>
(
<figureCitation id="134B02DD160A620CFE40FEA41126FEE7" box="[491,579,332,357]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1656,1679]" captionTargetBox="[159,1431,260,1555]" captionTargetId="figure@6.[151,1436,243,1586]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Cotinis barthelemyi. 9. Habitus, dorsal view; 10. Habitus, lateral view; 11. Clypeus, frontal view; 12. Head, lateral view; 13. Meso-metasternum." httpUri="https://zenodo.org/record/244389/files/figure.png" pageId="5" pageNumber="130">Fig. 12</figureCitation>
); the curved mesometasternal sutures in
<taxonomicName id="4C7065DB160A620CFB81FEA41663FEE7" box="[1066,1286,332,356]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFB81FEA41663FEE7" box="[1066,1286,332,356]" italics="true" pageId="5" pageNumber="130">C. patricialvarezae</emphasis>
</taxonomicName>
right-angled sutures in
<taxonomicName id="4C7065DB160A620CFEA7FE9812D1FE0B" box="[268,436,368,392]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="barthelemyi">
<emphasis id="B904C24A160A620CFEA7FE9812D1FE0B" box="[268,436,368,392]" italics="true" pageId="5" pageNumber="130">C. barthelemyi</emphasis>
</taxonomicName>
(
<figureCitation id="134B02DD160A620CFE6FFE87117DFE0B" box="[452,536,367,392]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1656,1679]" captionTargetBox="[159,1431,260,1555]" captionTargetId="figure@6.[151,1436,243,1586]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Cotinis barthelemyi. 9. Habitus, dorsal view; 10. Habitus, lateral view; 11. Clypeus, frontal view; 12. Head, lateral view; 13. Meso-metasternum." httpUri="https://zenodo.org/record/244389/files/figure.png" pageId="5" pageNumber="130">Fig. 13</figureCitation>
); and the shape of the parameres (
<figureCitation id="134B02DD160A620CFC34FE87174FFE0B" box="[927,1066,367,392]" captionStart="FIGURES 14 – 15" captionStartId="7.[151,264,1954,1977]" captionTargetBox="[273,1241,202,1898]" captionTargetId="figure@7.[214,1372,193,1932]" captionTargetPageId="7" captionText="FIGURES 14 – 15. Cotinis barthelemyi. 14. Parameres, frontal view; 15. Parameres, lateral view. Scale bar = 0.6 mm." httpUri="https://zenodo.org/record/244390/files/figure.png" pageId="5" pageNumber="130">Figs. 14–15</figureCitation>
). Other diagnostic characters for distinguishing these species are summarized in
<tableCitation id="C6F22BE3160A620CFD00FE7C1064FE2F" box="[683,769,404,429]" captionStart="TABLE 2" captionStartId="5.[151,239,581,604]" captionTargetBox="[159,1279,632,1026]" captionTargetPageId="5" captionText="TABLE 2. Differential characters between Cotinis patricialvarezae and C. barthelemyi." httpUri="http://table.plazi.org/id/DF0F4ED0160A620CFF3CFDAD1725FDDF" pageId="5" pageNumber="130" tableUuid="DF0F4ED0160A620CFF3CFDAD1725FDDF">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BCF1E58160A620CFF6CFE501176FD94" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="130">
<taxonomicName id="4C7065DB160A620CFF6CFE5012BDFE53" box="[199,472,440,464]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFF6CFE5012BDFE53" box="[199,472,440,464]" italics="true" pageId="5" pageNumber="130">Cotinis patricialvarezae</emphasis>
</taxonomicName>
is also morphologically similar to
<taxonomicName id="4C7065DB160A620CFCC1FE501722FE53" authority="Woodruff" authorityName="Woodruff" box="[874,1095,439,464]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="aliena">
<emphasis id="B904C24A160A620CFCC1FE5010B6FE53" box="[874,979,440,464]" italics="true" pageId="5" pageNumber="130">C. aliena</emphasis>
Woodruff
</taxonomicName>
in body size, body form, and form of the clypeal projection, but the frontal projection, elytra punctation, and parameres of
<taxonomicName id="4C7065DB160A620CFB0DFE341669FE77" box="[1190,1292,476,500]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="aliena">
<emphasis id="B904C24A160A620CFB0DFE341669FE77" box="[1190,1292,476,500]" italics="true" pageId="5" pageNumber="130">C. aliena</emphasis>
</taxonomicName>
are different from those of
<taxonomicName id="4C7065DB160A620CFE9FFDE8116EFD9B" box="[308,523,512,536]" class="Insecta" family="Cetoniidae" genus="Cotinis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="patricialvarezae">
<emphasis id="B904C24A160A620CFE9FFDE8116EFD9B" box="[308,523,512,536]" italics="true" pageId="5" pageNumber="130">C. patricialvarezae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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