252 lines
33 KiB
XML
252 lines
33 KiB
XML
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<document id="3B6030D1E8ABC8511C71BED085B0A47B" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D378784890DFFD921CA3E0AFCD6F80A" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus pellucidus Sokolov, 2016, sp.n." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="17" updateTime="1698393955637" updateUser="plazi">
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<mods:mods id="3A2EF0C0A12D260EAF6B6785B28284CE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E7F54D8932ACE5F7DF2A4CF0DA778C70">
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<mods:title id="6E1FD4158220E42D30407E6BF5B6539D">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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</mods:titleInfo>
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<mods:name id="6968ACB772BDC5E360BCE06378ECBB01" type="personal">
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<mods:roleTerm id="28097D4056E42BACFE2E7A5438DB0BF4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F8AD63B74C8C615B22D1F4D4DCD8AFB">Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0BCACCFE49EB7D6FC8254741740871E0">text</mods:typeOfResource>
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<mods:title id="A3CF4CE0EE74607044CC3B309025C77C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="05F83C25999BAE3065CB70716AC6E151">
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<mods:date id="1E73B619F7CB3489D7C0432DA3F03C9D">2016</mods:date>
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<mods:detail id="6CBBB59DC17C67E024A892ABB87DD3C1" type="volume">
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<mods:number id="A1D40E06C6D891223EB29C378C2603B6">4196</mods:number>
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</mods:detail>
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<mods:detail id="40C2AA17EAFA0105E9B8B3543278C43A" type="issue">
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<mods:number id="8FBF81516DE392DAA6B441E0473C8E06">1</mods:number>
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<mods:start id="A6038938C4F906CCE236AF56DDCEAC1D">1</mods:start>
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<mods:classification id="234A404CF497D263B18DD2CC5136A2FE">journal article</mods:classification>
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<mods:identifier id="3493A7D5883FCFB3B12A87655ADF38BE" type="DOI">10.11646/zootaxa.4196.1.1</mods:identifier>
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<mods:identifier id="1180AF3C31FC3CD4764DF527A31B192E" type="GBIF-Dataset">b7651a7b-df24-4347-9585-0633b88dba51</mods:identifier>
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<mods:identifier id="2821FFC6577DB139289FC8F6B4BF6AD2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="42DB1AA08238CB3CAE712EFA1AF49CA0" type="Zenodo-Dep">167648</mods:identifier>
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<mods:identifier id="16CD3D2AE0EC42DC6953326925EE5204" type="ZooBank">026D88BD-86C0-4811-814A-CB5CACB3104F</mods:identifier>
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</mods:mods>
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<treatment id="8D378784890DFFD921CA3E0AFCD6F80A" ID-DOI="http://doi.org/10.5281/zenodo.6089678" ID-GBIF-Taxon="124814319" ID-Zenodo-Dep="6089678" LSID="urn:lsid:plazi:treatment:8D378784890DFFD921CA3E0AFCD6F80A" httpUri="http://treatment.plazi.org/id/8D378784890DFFD921CA3E0AFCD6F80A" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
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<subSubSection id="4D846519890DFFDB21CA3E0AFBCBFA60" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="05213692890DFFDB21CA3E0AFE69FA82" blockId="16.[151,1143,1374,1434]" box="[151,469,1374,1400]" pageId="16" pageNumber="17">
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<heading id="5E6981FE890DFFDB21CA3E0AFE69FA82" bold="true" box="[151,469,1374,1400]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="37EAEA80890DFFDB21CA3E0AFE69FA82" bold="true" box="[151,469,1374,1400]" pageId="16" pageNumber="17">
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<emphasis id="37EAEA80890DFFDB21CA3E0AFE2FFA82" bold="true" box="[151,403,1374,1400]" italics="true" pageId="16" pageNumber="17">
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<taxonomicName id="C29E4D11890DFFDB21CA3E0AFE31FA82" ID-CoL="8M9FX" box="[151,397,1374,1400]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pellucidus" status="sp. nov.">Zeanillus pellucidus</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="2CD957FB890DFFDB20C63E0AFE69FA82" box="[411,469,1374,1400]" pageId="16" pageNumber="17" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="05213692890DFFDB21CA3ED5FBCBFA60" blockId="16.[151,1143,1374,1434]" box="[151,1143,1409,1434]" pageId="16" pageNumber="17">
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<figureCitation id="9DA52A17890DFFDB21CA3ED5FF5CFA60" box="[151,224,1409,1434]" captionStart="FIGURE 1" captionStartId="8.[151,250,1917,1939]" captionTargetBox="[172,1423,199,1885]" captionTargetId="figure@8.[164,1430,193,1891]" captionTargetPageId="8" captionText="FIGURE 1. SEM illustrations of head, dorsal aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. pallidus; D— Z. pellucidus; E— Z. lescheni; F— Z. nanus. Legend: cl—clypeus; cs—clypeal seta; fcc—fronto-clypeal carina; fs—frontal seta; lb—labrum; mp 3 — maxillary palpomere 3; mp 4 — maxillary palpomere 4; pos—postorbital seta; ssa—anterior supraorbital seta; ssp—posterior supraorbital seta. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/257540/files/figure.png" pageId="16" pageNumber="17">Figs. 1</figureCitation>
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D, 2C, 3E–F, 4D, 5D, 6D, 7E, 8C, 9C, 10CHN, 11E, 12Q–T, 14E, 15E, 16D, 18A
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</paragraph>
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</subSubSection>
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<subSubSection id="4D846519890DFFDB21CA3E9DFE88F96C" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="05213692890DFFDB21CA3E9DFC17F9FC" blockId="16.[151,1437,1481,2010]" pageId="16" pageNumber="17">
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<materialsCitation id="B5F63CCF890DFFDB21CA3E9DFC17F9FC" ID-GBIF-Occurrence="1341350620" collectingDate="2 Aug" collectionCode="NZAC" collectorName="DN Roy Cooper Res Oamaru" country="New Zealand" location="New Zealand" pageId="16" pageNumber="17" specimenCount="2" typeStatus="holotype">
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<emphasis id="37EAEA80890DFFDB21CA3E9DFEF5FA18" bold="true" box="[151,329,1481,1506]" pageId="16" pageNumber="17">
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<typeStatus id="DA258830890DFFDB21CA3E9DFF6EFA18" box="[151,210,1481,1506]" pageId="16" pageNumber="17">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="DA258830890DFFDB20083E9EFE50FA18" box="[341,492,1482,1506]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
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, male, in
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<collectionCode id="638FAE57890DFFDB233A3E9EFD0AFA18" box="[615,694,1482,1506]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="16" pageNumber="17">NZAC</collectionCode>
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, board-mounted, undissected, labeled: /
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<collectingCountry id="7D897602890DFFDB25CF3E9EFA91FA18" box="[1170,1325,1481,1506]" name="New Zealand" pageId="16" pageNumber="17">New Zealand</collectingCountry>
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<collectorName id="A86B5344890DFFDB24643E9EFEC7F9FC" pageId="16" pageNumber="17">DN Roy Cooper Res Oamaru</collectorName>
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<collectingDate id="6164E9BA890DFFDB20DF3EBAFE7BF9FC" box="[386,455,1518,1542]" pageId="16" pageNumber="17">
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<date id="71201052890DFFDB20DF3EBAFE7BF9FC" box="[386,455,1518,1542]" pageId="16" pageNumber="17">2 Aug</date>
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</collectingDate>
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0 7 / Washed soil sample broadleaf forest /.
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</materialsCitation>
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</paragraph>
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<paragraph id="05213692890DFFDB219A3D46FE88F96C" blockId="16.[151,1437,1481,2010]" pageId="16" pageNumber="17">
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PARATYPES (21 specimens, 5 dissected), 8 males and 11 females labeled same as holotype;
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<materialsCitation id="B5F63CCF890DFFDB25813D46FB9BF9B4" ID-GBIF-Occurrence="1341350612" collectingDate="1919-01" collectorName="DN Glen Warren Res Oamaru" country="New Zealand" location="New Zealand" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
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<specimenCount id="1398FD1B890DFFDB25813D46FA81F9D0" box="[1244,1341,1553,1578]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
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labeled: /
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<collectingCountry id="7D897602890DFFDB21FA3D62FE83F9B4" box="[167,319,1589,1614]" name="New Zealand" pageId="16" pageNumber="17">New Zealand</collectingCountry>
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<collectorName id="A86B5344890DFFDB20153D62FD1CF9B4" box="[328,672,1589,1614]" pageId="16" pageNumber="17">DN Glen Warren Res Oamaru</collectorName>
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<collectingDate id="6164E9BA890DFFDB23F43D62FD4FF9B7" box="[681,755,1590,1614]" pageId="16" pageNumber="17" value="1919-01">
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<date id="71201052890DFFDB23F43D62FD4FF9B7" box="[681,755,1590,1614]" pageId="16" pageNumber="17" value="1919-01">19 Jan</date>
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</collectingDate>
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0 7 / Washed soil sample /
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</materialsCitation>
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;
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<materialsCitation id="B5F63CCF890DFFDB25693D62FE88F96C" ID-GBIF-Occurrence="1341350618" collectingDate="2008-01-12" collectorName="J. Nunn" country="New Zealand" elevation="150" location="Glen Warren Reserve" municipality="Oamaru" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
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<specimenCount id="1398FD1B890DFFDB25693D62FB3DF9B4" box="[1076,1153,1589,1614]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
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labeled: /
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<collectingCountry id="7D897602890DFFDB25A03D62FA25F9B4" box="[1277,1433,1589,1614]" name="New Zealand" pageId="16" pageNumber="17">New Zealand</collectingCountry>
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: DN,
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<location id="00416049890DFFDB21933D0DFE78F98B" LSID="urn:lsid:plazi:treatment:8D378784890DFFD921CA3E0AFCD6F80A:00416049890DFFDB21933D0DFE78F98B" box="[206,452,1625,1650]" country="New Zealand" municipality="Oamaru" name="Glen Warren Reserve" pageId="16" pageNumber="17">Glen Warren Reserve</location>
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,
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<collectingMunicipality id="E545ACE8890DFFDB208D3D0EFD91F988" box="[464,557,1626,1650]" pageId="16" pageNumber="17">Oamaru</collectingMunicipality>
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,
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<elevation id="8EB3D1A1890DFFDB23643D0DFDC0F988" box="[569,636,1625,1650]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="150.0">
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<quantity id="C2669B77890DFFDB23643D0DFDC0F988" box="[569,636,1625,1650]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="150.0">150m</quantity>
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</elevation>
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, washed soil sample, dry coastal broadleaf forest, H45/864483
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<collectorName id="A86B5344890DFFDB24113D0EFA20F98B" box="[1356,1436,1626,1649]" pageId="16" pageNumber="17">J.Nunn</collectorName>
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<collectingDate id="6164E9BA890DFFDB21CA3D2AFEA2F96C" box="[151,286,1662,1686]" pageId="16" pageNumber="17" value="2008-01-12">
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<date id="71201052890DFFDB21CA3D2AFEA2F96C" box="[151,286,1662,1686]" pageId="16" pageNumber="17" value="2008-01-12">12 Jan 2008</date>
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</collectingDate>
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/.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D846519890DFFDB219A3DF5FAC5F924" pageId="16" pageNumber="17" type="description">
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<paragraph id="05213692890DFFDB219A3DF5FAC5F924" blockId="16.[151,1437,1481,2010]" pageId="16" pageNumber="17">
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<emphasis id="37EAEA80890DFFDB219A3DF5FE3BF940" bold="true" box="[199,391,1697,1722]" pageId="16" pageNumber="17">Specific epithet.</emphasis>
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The specific epithet is a Latinized adjective in the masculine form and is derived from the Latin adjective
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<taxonomicName id="C29E4D11890DFFDB20183D91FE04F927" box="[325,440,1733,1757]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pellucidus">
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<emphasis id="37EAEA80890DFFDB20183D91FE04F927" box="[325,440,1733,1757]" italics="true" pageId="16" pageNumber="17">pellūcidus</emphasis>
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</taxonomicName>
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meaning "transparent". The epithet refers to the translucent elytra of the new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D846519890DFFDB219A3DBDFE72F8B0" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="05213692890DFFDB219A3DBDFBE2F8FB" blockId="16.[151,1437,1481,2010]" box="[199,1118,1769,1794]" pageId="16" pageNumber="17">
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<materialsCitation id="B5F63CCF890DFFDB219A3DBDFBE2F8FB" ID-GBIF-Occurrence="1341350613" box="[199,1118,1769,1794]" country="New Zealand" location="Oamaru" municipality="South Island" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
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<emphasis id="37EAEA80890DFFDB219A3DBDFED9F8F8" bold="true" box="[199,357,1769,1794]" pageId="16" pageNumber="17">
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<typeStatus id="DA258830890DFFDB219A3DBDFEBEF8F8" box="[199,258,1769,1794]" pageId="16" pageNumber="17">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="7D897602890DFFDB20313DBEFDB9F8F8" box="[364,517,1769,1794]" name="New Zealand" pageId="16" pageNumber="17">New Zealand</collectingCountry>
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,
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<collectingMunicipality id="E545ACE8890DFFDB234D3DBDFD23F8F8" box="[528,671,1769,1794]" pageId="16" pageNumber="17">South Island</collectingMunicipality>
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,
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<collectingRegion id="C75AF870890DFFDB23F63DBEFD4EF8F8" box="[683,754,1770,1794]" country="New Zealand" name="Otago" pageId="16" pageNumber="17">Otago</collectingRegion>
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,
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<location id="00416049890DFFDB23A03DBEFCE6F8F8" LSID="urn:lsid:plazi:treatment:8D378784890DFFD921CA3E0AFCD6F80A:00416049890DFFDB23A03DBEFCE6F8F8" box="[765,858,1770,1794]" country="New Zealand" municipality="South Island" name="Oamaru" pageId="16" pageNumber="17" stateProvince="Otago">Oamaru</location>
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, Glen Warren Reserve.
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</materialsCitation>
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</paragraph>
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<paragraph id="05213692890DFFDB219A3C59FE72F8B0" blockId="16.[151,1437,1481,2010]" pageId="16" pageNumber="17">
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<emphasis id="37EAEA80890DFFDB219A3C59FEE1F8DC" bold="true" box="[199,349,1805,1830]" pageId="16" pageNumber="17">Recognition.</emphasis>
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Adults of this species (
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<figureCitation id="9DA52A17890DFFDB233A3C59FD0AF8DC" box="[615,694,1805,1830]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
|
|||
|
|
E) are distinguished from other congeners by the characters listed in the subgeneric diagnosis.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="4D846519890DFFD9219A3C01FCD6F80A" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
|
|||
|
|
<paragraph id="05213692890DFFDB219A3C01FB61F894" blockId="16.[151,1437,1481,2010]" box="[199,1245,1877,1902]" pageId="16" pageNumber="17">
|
|||
|
|
<emphasis id="37EAEA80890DFFDB219A3C01FEE5F894" bold="true" box="[199,345,1877,1902]" pageId="16" pageNumber="17">Description.</emphasis>
|
|||
|
|
Size. Small for genus (SBL range
|
|||
|
|
<quantity id="C2669B77890DFFDB23833C01FC3FF897" box="[734,899,1877,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.64" metricValueMin="1.41" pageId="16" pageNumber="17" unit="mm" value="1.525" valueMax="1.64" valueMin="1.41">1.41–1.64 mm</quantity>
|
|||
|
|
, mean 1.54±
|
|||
|
|
<quantity id="C2669B77890DFFDB254F3C01FB3BF897" box="[1042,1159,1877,1902]" metricMagnitude="-5" metricUnit="m" metricValue="5.8" pageId="16" pageNumber="17" unit="mm" value="0.058">0.058 mm</quantity>
|
|||
|
|
, n=20).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="05213692890DFFDB219A3C2DFE17F820" blockId="16.[151,1437,1481,2010]" pageId="16" pageNumber="17">Habitus. Body form subdepressed, ovoid, moderately elongate (WE/SBL 0.35±0.007), head of moderate width for genus compared to pronotum (WH/WPm 0.75±0.019), pronotum of moderate width in comparison to elytra (WPm/WE 0.74±0.022).</paragraph>
|
|||
|
|
<caption id="51E1661A890CFFDA21CA3C21FD81F810" httpUri="https://zenodo.org/record/257545/files/figure.png" pageId="17" pageNumber="18" targetBox="[252,1326,204,1867]" targetPageId="17">
|
|||
|
|
<paragraph id="05213692890CFFDA21CA3C21FD81F810" blockId="17.[151,1437,1909,2026]" pageId="17" pageNumber="18">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA21CA3C21FEA9F870" bold="true" box="[151,277,1909,1931]" pageId="17" pageNumber="18">FIGURE 8.</emphasis>
|
|||
|
|
SEM illustrations of chaetotaxy of elytra, dorso-lateral aspect, of
|
|||
|
|
<taxonomicName id="C29E4D11890CFFDA22903C21FB97F871" box="[973,1067,1909,1931]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA22903C21FB97F871" box="[973,1067,1909,1931]" italics="true" pageId="17" pageNumber="18">Zeanillus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species. A—
|
|||
|
|
<taxonomicName id="C29E4D11890CFFDA25E53C22FAF1F871" box="[1208,1357,1910,1931]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="punctigerus">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA25E53C22FAF1F871" box="[1208,1357,1910,1931]" italics="true" pageId="17" pageNumber="18">Z. punctigerus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; B—
|
|||
|
|
<taxonomicName id="C29E4D11890CFFDA24DA3C22FF55F856" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA24DA3C22FF55F856" italics="true" pageId="17" pageNumber="18">Z. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; C—
|
|||
|
|
<taxonomicName id="C29E4D11890CFFDA207D3CCCFE1FF856" box="[288,419,1942,1965]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pellucidus">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA207D3CCCFE1FF856" box="[288,419,1942,1965]" italics="true" pageId="17" pageNumber="18">Z. pellucidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; D—
|
|||
|
|
<taxonomicName id="C29E4D11890CFFDA20803CCCFDF0F856" box="[477,588,1942,1965]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lescheni">
|
|||
|
|
<emphasis id="37EAEA80890CFFDA20803CCCFDF0F856" box="[477,588,1942,1965]" italics="true" pageId="17" pageNumber="18">Z. lescheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Legend: ed2—scutellar seta; ed3—1st discal seta; ed4–5—2nd discal seta; ed6–7— 3d discal seta; ed8—apical seta; eo1–9—setae 1–9 from the umbilical series; smes—subapical marginal elytral seta; sssubapical sinuation. Scale bars = 0.2 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="51E1661A890FFFD921CA3F20FDB0FB3D" httpUri="https://zenodo.org/record/257546/files/figure.png" pageId="18" pageNumber="19" targetBox="[189,1406,189,1111]" targetPageId="18">
|
|||
|
|
<paragraph id="05213692890FFFD921CA3F20FDB0FB3D" blockId="18.[151,1437,1140,1223]" pageId="18" pageNumber="19">
|
|||
|
|
<emphasis id="37EAEA80890FFFD921CA3F20FEA9FB73" bold="true" box="[151,277,1140,1162]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
|
|||
|
|
SEM illustrations of pygidium and apex of elytra, dorsal aspect, of
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD922803F20FB87FB70" box="[989,1083,1140,1162]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="37EAEA80890FFFD922803F20FB87FB70" box="[989,1083,1140,1162]" italics="true" pageId="18" pageNumber="19">Zeanillus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species. A—
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD9259A3F21FAF1FB70" box="[1223,1357,1140,1162]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="phyllobius">
|
|||
|
|
<emphasis id="37EAEA80890FFFD9259A3F21FAF1FB70" box="[1223,1357,1140,1162]" italics="true" pageId="18" pageNumber="19">Z. phyllobius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; B—
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD924DA3F21FEB3FB53" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="punctigerus">
|
|||
|
|
<emphasis id="37EAEA80890FFFD924DA3F21FEB3FB53" italics="true" pageId="18" pageNumber="19">Z. punctigerus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; C—
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD920143FC0FE73FB53" box="[329,463,1171,1193]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pellucidus">
|
|||
|
|
<emphasis id="37EAEA80890FFFD920143FC0FE73FB53" box="[329,463,1171,1193]" italics="true" pageId="18" pageNumber="19">Z. pellucidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; D—
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD923563FC0FDC7FB53" box="[523,635,1171,1193]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lescheni">
|
|||
|
|
<emphasis id="37EAEA80890FFFD923563FC0FDC7FB53" box="[523,635,1171,1193]" italics="true" pageId="18" pageNumber="19">Z. lescheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Legend: ea—apex of right elytron; ed8—apical seta; pgd—pygidium; suelytral suture. Scale bars = 0.05 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3FBBFCA1FAFD" blockId="18.[151,1437,1263,2032]" box="[199,797,1263,1288]" pageId="18" pageNumber="19">Color. Body color testaceous, appendages testaceous.</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3E46FBF5FA88" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">Prothorax. Pronotum slightly long (LP/LE 0.41±0.017) and slightly transverse (WPm/LP 1.34±0.039), with slightly sinuated and markedly constricted posteriorly lateral margins (WPm/WPp 1.84±0.090). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.44±0.057).</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3E29FEA1FA27" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">Elytra. Moderately depressed, comparatively short (LE/SBL 0.47±0.023) and moderately wide (WE/LE 0.75±0.031). Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third.</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3EB3FDDCF990" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">
|
|||
|
|
Male genitalia. Median lobe (
|
|||
|
|
<figureCitation id="9DA52A17890FFFD9237F3EB3FDCAFA05" box="[546,630,1511,1536]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
|
|||
|
|
Q) with slightly concave ventral margin and semicircular short apex. Apical orifice area with a pair of short ostial fields. Sclerites of internal sac very short, united basally, with unequally sclerotized branches. Left paramere without apical constriction (
|
|||
|
|
<figureCitation id="9DA52A17890FFFD922B53D7AFB80F9BD" box="[1000,1084,1582,1607]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
|
|||
|
|
S). Ring sclerite with short and moderately transverse handle (
|
|||
|
|
<figureCitation id="9DA52A17890FFFD920B33D05FDFCF990" box="[494,576,1617,1642]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="18" pageNumber="19">Fig. 14</figureCitation>
|
|||
|
|
E).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3D21FD29F92F" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">
|
|||
|
|
Female internal genitalia. Goncoxite 2 with long ensiform setae (
|
|||
|
|
<figureCitation id="9DA52A17890FFFD922F63D21FC42F974" box="[939,1022,1653,1678]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="18" pageNumber="19">Fig. 15</figureCitation>
|
|||
|
|
E). Spermatheca sclerotized, slightly elongate, with two compartments (
|
|||
|
|
<figureCitation id="9DA52A17890FFFD923463DCCFDD0F94B" box="[539,620,1688,1713]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="18" pageNumber="19">Fig. 16</figureCitation>
|
|||
|
|
E). Spermathecal duct short with few coils. Attachments of spermathecal duct and gland to spermatheca close together.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3D8BFDA0F8E6" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">
|
|||
|
|
<emphasis id="37EAEA80890FFFD9219A3D8BFDBAF902" bold="true" box="[199,518,1759,1784]" pageId="18" pageNumber="19">Geographical distribution.</emphasis>
|
|||
|
|
This species inhabits coastal plain of North
|
|||
|
|
<collectingRegion id="C75AF870890FFFD925553DB4FBF3F902" box="[1032,1103,1760,1784]" country="New Zealand" name="Otago" pageId="18" pageNumber="19">Otago</collectingRegion>
|
|||
|
|
, where it is known from one locality only (18A, blue diamond).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3C72FBEAF8C5" blockId="18.[151,1437,1263,2032]" box="[199,1110,1830,1855]" pageId="18" pageNumber="19">
|
|||
|
|
<emphasis id="37EAEA80890FFFD9219A3C72FEF3F8C5" bold="true" box="[199,335,1830,1855]" pageId="18" pageNumber="19">Way of life.</emphasis>
|
|||
|
|
Specimens were collected from soil in a dry broadleaf coastal forest.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="05213692890FFFD9219A3C1EFCD6F80A" blockId="18.[151,1437,1263,2032]" pageId="18" pageNumber="19">
|
|||
|
|
<emphasis id="37EAEA80890FFFD9219A3C1EFECCF899" bold="true" box="[199,368,1866,1891]" pageId="18" pageNumber="19">Relationships.</emphasis>
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD920243C1EFDE2F898" box="[377,606,1866,1890]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pellucidus">
|
|||
|
|
<emphasis id="37EAEA80890FFFD920243C1EFDE2F898" box="[377,606,1866,1890]" italics="true" pageId="18" pageNumber="19">Zeanillus pellucidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a morphologically isolated species within the genus. The presence of postorbital seta, shared with the members of
|
|||
|
|
<emphasis id="37EAEA80890FFFD923C93C39FC8EF87F" box="[660,818,1901,1925]" italics="true" pageId="18" pageNumber="19">
|
|||
|
|
<taxonomicName id="C29E4D11890FFFD923C93C39FD40F87F" box="[660,764,1901,1925]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" sensu="stricto">Zeanillus</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="2CD957FB890FFFD922593C3AFC8EF87F" box="[772,818,1902,1925]" pageId="18" pageNumber="19" sensu="stricto">s.str</taxonomicNameLabel>
|
|||
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</emphasis>
|
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|
., putatively, suggests the latter as the closest relatives. However, striking differences between two taxa in the structure of male genitalia and spermatheca suggests that they are not very close. It is likely that
|
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|
<taxonomicName id="C29E4D11890FFFD923123CE1FD5CF836" box="[591,736,1972,1996]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pellucidus">
|
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|
<emphasis id="37EAEA80890FFFD923123CE1FD5CF836" box="[591,736,1972,1996]" italics="true" pageId="18" pageNumber="19">Z. pellucidus</emphasis>
|
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|
</taxonomicName>
|
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|
|
is a highly derived taxon of ancient origin and its relationship to the representatives of other subgenera is equidistantly remote.
|
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|
|
</paragraph>
|
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|
|
</subSubSection>
|
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|
|
</treatment>
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</document>
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