233 lines
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233 lines
18 KiB
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<document ID-DOI="10.11646/palaeoentomology.4.4.13" ID-GBIF-Dataset="6cd3d259-465d-4ab3-8a32-d57a812bb6e0" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507925" ID-ZooBank="D1F5A853-32D1-470B-A609-D54F7B8C60CA" checkinTime="1631645857228" checkinUser="carolina" docAuthor="JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S. & CAI, CHEN-YANG" docDate="2021" docId="03B28787AB45067AFF0D8039FC5AFF37" docLanguage="en" docName="palaeoentomology.4.4.13.pdf" docOrigin="Palaeoentomology 4 (4)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.4.13" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Arceotermes Engel & Jiang 2021, gen. nov." docType="treatment" docVersion="6" lastPageNumber="381" masterDocId="FF8BFFFFAB42067DFF858453FF89FB0A" masterDocTitle="Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)" masterLastPageNumber="389" masterPageNumber="374" pageNumber="381" updateTime="1661815541049" updateUser="admin">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>JIANG, RI-XIN</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ZHANG, HONG-RUI</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ELDREDGE, K. TARO</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>SONG, XIAO-BIN</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>LI, YAN-DA</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>TIHELKA, ERIK</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>HUANG, DI-YING</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>WANG, SHUO</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ENGEL, MICHAEL S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>CAI, CHEN-YANG</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>374</mods:start>
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<mods:end>389</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.4.4.13</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.4.4.13</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6cd3d259-465d-4ab3-8a32-d57a812bb6e0</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5507925</mods:identifier>
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<mods:identifier type="ZooBank">D1F5A853-32D1-470B-A609-D54F7B8C60CA</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5507945" ID-GBIF-Taxon="188191020" ID-Zenodo-Dep="5507945" LSID="urn:lsid:plazi:treatment:03B28787AB45067AFF0D8039FC5AFF37" httpUri="http://treatment.plazi.org/id/03B28787AB45067AFF0D8039FC5AFF37" lastPageNumber="381" pageId="7" pageNumber="381">
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<subSubSection box="[136,578,1130,1156]" pageId="7" pageNumber="381" type="nomenclature">
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<paragraph blockId="7.[136,578,1130,1156]" box="[136,578,1130,1156]" pageId="7" pageNumber="381">
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<heading bold="true" box="[136,578,1130,1156]" fontSize="10" level="2" pageId="7" pageNumber="381" reason="3">
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<emphasis bold="true" box="[136,578,1130,1156]" pageId="7" pageNumber="381">
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<taxonomicName authority="Engel & Jiang" authorityName="Engel & Jiang" authorityYear="2021" box="[136,460,1130,1156]" class="Insecta" family="Staphylinidae" genus="Arceotermes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[136,281,1130,1156]" italics="true" pageId="7" pageNumber="381">Arceotermes</emphasis>
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Engel & Jiang
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</taxonomicName>
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,
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<taxonomicNameLabel box="[473,578,1130,1156]" pageId="7" pageNumber="381" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="381" type="type_taxon">
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<paragraph blockId="7.[136,778,1202,2092]" pageId="7" pageNumber="381">
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<emphasis bold="true" box="[136,296,1202,1228]" pageId="7" pageNumber="381">
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<typeStatus box="[136,195,1202,1228]" pageId="7" pageNumber="381">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName authority="Engel & Jiang" authorityName="Engel & Jiang" authorityYear="2021" box="[307,729,1202,1229]" class="Insecta" family="Staphylinidae" genus="Arceotermes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="hospitis" status="sp. nov.">
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<emphasis box="[307,546,1202,1229]" italics="true" pageId="7" pageNumber="381">Arceotermes hospitis</emphasis>
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Engel & Jiang
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</taxonomicName>
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,
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<emphasis bold="true" pageId="7" pageNumber="381">
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<taxonomicNameLabel pageId="7" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="381" type="etymology">
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<paragraph blockId="7.[136,778,1202,2092]" pageId="7" pageNumber="381">
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<emphasis bold="true" box="[189,324,1274,1300]" pageId="7" pageNumber="381">Etymology.</emphasis>
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The new genus-group name is a combination of the Latin terms
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<emphasis box="[545,608,1311,1336]" italics="true" pageId="7" pageNumber="381">arceo</emphasis>
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(meaning, to “hinder” or “guard”) and
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<emphasis box="[454,529,1347,1372]" italics="true" pageId="7" pageNumber="381">termes</emphasis>
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(“termite”; genitive,
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<emphasis box="[136,222,1383,1408]" italics="true" pageId="7" pageNumber="381">termitis</emphasis>
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). The gender of the name is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="381" type="diagnosis">
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<paragraph blockId="7.[136,778,1202,2092]" lastBlockId="7.[810,1452,158,1085]" pageId="7" pageNumber="381">
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<emphasis bold="true" box="[189,312,1418,1444]" pageId="7" pageNumber="381">Diagnosis.</emphasis>
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Total body length (as preserved) 5.00 mm, head length (excluding mandibles)
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<quantity box="[657,773,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="7" pageNumber="381" unit="mm" value="2.08">2.08 mm</quantity>
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, maximum width
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<quantity box="[326,429,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="7" pageNumber="381" unit="mm" value="1.83">1.83 mm</quantity>
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. Like most soldier termites, the head is large and longer than broad, although the posterior temples are not tumid (tumid in
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<taxonomicName authority="Desneux, 1904 b" authorityName="Desneux" authorityYear="1904" class="Insecta" family="Archotermopsidae" genus="Archotermopsis" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[491,667,1562,1588]" italics="true" pageId="7" pageNumber="381">Archotermopsis</emphasis>
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<bibRefCitation author="Desneux, J." pageId="7" pageNumber="381" pagination="445 - 446" refId="ref9680" refString="Desneux, J. (1904 b) A new termite from India. Journal of the Bombay Natural History Society, 15, 445 - 446." type="journal article" year="1904">Desneux, 1904b</bibRefCitation>
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</taxonomicName>
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) and the head is not dorsoventrally compressed (compressed in
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<taxonomicName authorityName="Holmgren" authorityYear="1910" box="[323,501,1634,1661]" class="Insecta" family="Stolotermitidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="family">Stolotermitidae</taxonomicName>
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). The lateral surface of the head lacks striations (present in
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<taxonomicName authorityName="Desneux" authorityYear="1904" box="[585,761,1670,1696]" class="Insecta" family="Archotermopsidae" genus="Archotermopsis" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[585,761,1670,1696]" italics="true" pageId="7" pageNumber="381">Archotermopsis</emphasis>
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</taxonomicName>
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), and the lateral margins are largely parallel. The dorsal surface of the head lacks a clearly visible Y-shaped ecdysial cleavage scar (it is difficult, though, to determine whether it is absent or merely exceedingly faint), lacks a fontanelle, and lacks ocelli. The compound eyes are absent and the antenna is composed of 17, largely moniliform articles. Mandibles are large, elongate, and gently bent ventrad toward their apices, with prominent apical and marginal teeth. Like
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<taxonomicName box="[518,725,1994,2020]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
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and
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<taxonomicName authorityName="Desneux" authorityYear="1904" box="[136,315,2030,2056]" class="Insecta" family="Hodotermitidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="family">Hodotermitidae</taxonomicName>
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, the soldier mandibles have two right marginal teeth and three left marginal teeth (in addition to the apical tooth), the right marginal teeth are broad, much like that of
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<taxonomicName authorityName="Desneux" authorityYear="1904" box="[1005,1181,194,220]" class="Insecta" family="Archotermopsidae" genus="Archotermopsis" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[1005,1181,194,220]" italics="true" pageId="7" pageNumber="381">Archotermopsis</emphasis>
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</taxonomicName>
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, and simple (sometimes with two points, such as in
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<taxonomicName authority="Holmgren, 1911" authorityName="Holmgren" authorityYear="1911" class="Insecta" family="Archotermopsidae" genus="Hodotermopsis" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[1147,1318,230,256]" italics="true" pageId="7" pageNumber="381">Hodotermopsis</emphasis>
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<bibRefCitation author="Holmgren, N." pageId="7" pageNumber="381" pagination="1 - 86" refId="ref10494" refString="Holmgren, N. (1911) Termitenstudien: 2. Systematik der Termiten. Die Familien Protermitidae und Mesotermitidae. Kungliga Svenska Vetenskapsakademiens Handlingar, 46 (6), 1 - 86." type="journal article" year="1911">Holmgren, 1911</bibRefCitation>
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</taxonomicName>
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(
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<taxonomicName authority="Engel" authorityName="Engel" authorityYear="2021" box="[878,1153,266,292]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="379" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Hodotermopsinae">Hodotermopsinae Engel</taxonomicName>
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,
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<emphasis bold="true" box="[1164,1315,266,292]" pageId="7" pageNumber="381">
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<taxonomicNameLabel box="[1164,1315,266,292]" pageId="7" pageNumber="381" rank="subFamily">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<emphasis box="[1328,1435,266,292]" italics="true" pageId="7" pageNumber="381">vide infra</emphasis>
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),
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<taxonomicName authority="(Banks, 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[810,1248,302,329]" class="Insecta" family="Archotermopsidae" genus="Zootermopsis" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="species" species="laticeps">
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<emphasis box="[810,1064,302,328]" italics="true" pageId="7" pageNumber="381">Zootermopsis laticeps</emphasis>
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(
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<bibRefCitation author="Banks, N." box="[1087,1241,302,329]" pageId="7" pageNumber="381" pagination="336 - 337" refId="ref9042" refString="Banks, N. (1906) Two new termites. Entomological News, 17 (9), 336 - 337." type="journal article" year="1906">Banks, 1906</bibRefCitation>
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)
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</taxonomicName>
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, and to a lesser degree like
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<taxonomicName authority="Sjostedt, 1926" authorityName="Sjostedt" authorityYear="1926" box="[942,1313,338,365]" class="Insecta" family="Hodotermitidae" genus="Microhodotermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[942,1140,338,364]" italics="true" pageId="7" pageNumber="381">Microhodotermes</emphasis>
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<bibRefCitation author="Sjostedt, Y." box="[1148,1313,338,365]" pageId="7" pageNumber="381" pagination="1 - 419" refId="ref11579" refString="Sjostedt, Y. (1926) Revision der Termiten Afrikas: 3. Monographie. Kungliga Svenska Vetenskapsakademiens Handlingar, 3 (3), 1 - 419." type="journal article" year="1926">Sjöstedt, 1926</bibRefCitation>
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</taxonomicName>
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). The apical teeth are long, pointed, and curved. The postmentum is greatly broadened anteriorly, with concave lateral margins tapering posteriorly. The pronotum is broad (not massive and elongate as in
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<taxonomicName authority="Engel et al., 2016" authorityName="Engel" authorityYear="2016" box="[1013,1393,482,509]" class="Insecta" genus="Ginormotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
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<emphasis box="[1013,1189,483,509]" italics="true" pageId="7" pageNumber="381">Ginormotermes</emphasis>
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<bibRefCitation author="Engel, M. S. & Barden, P. & Riccio, M. L. & Grimaldi, D. A." box="[1194,1393,482,509]" pageId="7" pageNumber="381" pagination="522 - 530" refId="ref10116" refString="Engel, M. S., Barden, P., Riccio, M. L. & Grimaldi, D. A. (2016) Morphologically specialized termite castes and advanced sociality in the Early Cretaceous. Current Biology, 26 (4), 522 - 530. https: // doi. org / 10.1016 / j. cub. 2015.12.061" type="journal article" year="2016">
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Engel
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<emphasis box="[1265,1313,482,508]" italics="true" pageId="7" pageNumber="381">et al</emphasis>
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., 2016
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</bibRefCitation>
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</taxonomicName>
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), and about as wide as the head, with the anterior margin broadly concave and the lateral margins faintly convex, roughly parallel, and broadly rounded posteriorly to a broadly and weakly convex posterior margin. The surface of the pronotum is flat.The tibial spur formula is 3-3-? (metatarsi not preserved in
|
||
|
|
<typeStatus box="[1000,1102,698,725]" pageId="7" pageNumber="381">holotype</typeStatus>
|
||
|
|
), and the tarsi are tetramerous, while the pretarsus lacks an arolium. Unfortunately, the abdominal apex is damaged in the
|
||
|
|
<typeStatus box="[1217,1315,770,796]" pageId="7" pageNumber="381">holotype</typeStatus>
|
||
|
|
of the type species and the cerci are not preserved.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="7.[810,1452,158,1085]" pageId="7" pageNumber="381">
|
||
|
|
The genus differs from soldiers of
|
||
|
|
<taxonomicName box="[1241,1406,842,868]" class="Insecta" genus="Krishnatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis box="[1241,1406,842,868]" italics="true" pageId="7" pageNumber="381">Krishnatermes</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName box="[810,986,879,905]" class="Insecta" genus="Ginormotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis box="[810,986,879,905]" italics="true" pageId="7" pageNumber="381">Ginormotermes</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
by the tetramerous tarsi (pentamerous in those genera), and from
|
||
|
|
<taxonomicName authority="Zhao et al., 2020" authorityName="Zhao" authorityYear="2020" box="[1092,1451,914,941]" class="Insecta" family="Stolotermitidae" genus="Cosmotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis box="[1092,1244,915,941]" italics="true" pageId="7" pageNumber="381">Cosmotermes</emphasis>
|
||
|
|
<bibRefCitation author="Zhao, Z. P. & Yin, X. C. & Shih, C. K. & Gao, T. P. & Ren, D." box="[1254,1451,914,941]" pageId="7" pageNumber="381" pagination="381 - 390" refId="ref12049" refString="Zhao, Z. P., Yin, X. C., Shih, C. K., Gao, T. P. & Ren, D. (2020) Termite colonies from mid-Cretaceous Myanmar demonstrate their early eusocial lifestyle in damp wood. National Science Review, 7 (2), 381 - 390. https: // doi. org / 10.1093 / nsr / nwz 141" type="journal article" year="2020">
|
||
|
|
Zhao
|
||
|
|
<emphasis box="[1321,1372,914,940]" italics="true" pageId="7" pageNumber="381">et al</emphasis>
|
||
|
|
., 2020
|
||
|
|
</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
by the absence of compound eyes, the long left first marginal tooth relative to the second marginal tooth, and the broader pronotum (distinctly narrower than head in
|
||
|
|
<taxonomicName box="[810,962,1059,1085]" class="Insecta" family="Stolotermitidae" genus="Cosmotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="381" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis box="[810,962,1059,1085]" italics="true" pageId="7" pageNumber="381">Cosmotermes</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|