169 lines
17 KiB
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169 lines
17 KiB
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<document id="FBDE1CBA161B8E7F455082CF61C78F55" ID-DOI="10.11646/zootaxa.3866.1.8" ID-GBIF-Dataset="ce8c4396-e39e-4de0-ab78-0fdee09b6bb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="228760" ID-ZooBank="2BECDD90-3B85-4F44-8C3F-585AC3D7A84B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460916092506" checkinUser="plazi" docAuthor="Oberprieler, Stefanie K. & Yeates, David K." docDate="2014" docId="039B87B7B22B3A69FFC4FA34D88B4CBD" docLanguage="en" docName="zt03866p144.pdf" docOrigin="Zootaxa 3866 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Notoatherix Oberprieler & Yeates, gen. nov." docType="treatment" docVersion="7" lastPageNumber="141" masterDocId="FFA2FFCFB22A3A6AFF53FFF9DA1F4F6D" masterDocTitle="Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae)" masterLastPageNumber="144" masterPageNumber="138" pageNumber="139" updateTime="1698361698483" updateUser="plazi">
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<mods:titleInfo id="206B66FFFD0B8B419C48B99F7D59AF4C">
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<mods:title id="E1993D556505DDDFE505B4FCDA3B4716">Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae)</mods:title>
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<mods:name id="D919FAE2D2561486D1F03BE58CBF91CC" type="personal">
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</mods:role>
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<mods:namePart id="3C8459264703672F164C4484C60A2F9F">Oberprieler, Stefanie K.</mods:namePart>
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</mods:name>
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<mods:name id="CFC5FC3C2031E75F8F7436622EF94F44" type="personal">
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<mods:roleTerm id="DA127137144FFBBE9FA8DD75FE8BB416">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71D9642B5A50C7B89D798A90F663FBAC">Yeates, David K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B8DDBB7B0E19B060E30F7D6C6649B3B1">text</mods:typeOfResource>
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<mods:title id="979566D7652F43962CACCDF314E400F0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0447D96A4CB606485ACE05773B2AFD02">
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<mods:date id="F1373937779388FB73271012939D60A4">2014</mods:date>
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<mods:detail id="C7B3646978DC72A6C4D9F59D6532B430" type="volume">
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<mods:number id="A2B3A5012319F011A4B08B75094ECDF9">3866</mods:number>
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<mods:number id="B2332835FB7E4289F21B2E07CA594814">1</mods:number>
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<mods:identifier id="9EE4BDCEC3C452BA7417B2C2818D33CC" type="DOI">10.11646/zootaxa.3866.1.8</mods:identifier>
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<mods:identifier id="AD9390BD1581828E70CCA59D0039F080" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A9DC5809280E3D4EAFCA6A61E831A9A3" type="Zenodo-Dep">228760</mods:identifier>
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</mods:mods>
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<treatment id="039B87B7B22B3A69FFC4FA34D88B4CBD" ID-DOI="http://doi.org/10.5281/zenodo.6143087" ID-GBIF-Taxon="119588781" ID-Zenodo-Dep="6143087" LSID="urn:lsid:plazi:treatment:039B87B7B22B3A69FFC4FA34D88B4CBD" httpUri="http://treatment.plazi.org/id/039B87B7B22B3A69FFC4FA34D88B4CBD" lastPageId="3" lastPageNumber="141" pageId="1" pageNumber="139">
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<subSubSection id="C328652AB22B3A6BFFC4FA34DB3F4967" pageId="1" pageNumber="139" type="nomenclature">
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<paragraph id="8B8D36A1B22B3A6BFFC4FA34D8F34A8B" blockId="1.[151,748,1485,1546]" box="[151,748,1485,1510]" pageId="1" pageNumber="139">
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<heading id="D0C581CDB22B3A6BFFC4FA34D8F34A8B" bold="true" box="[151,748,1485,1510]" fontSize="10" level="2" pageId="1" pageNumber="139" reason="2">
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<emphasis id="B946EAB3B22B3A6BFFC4FA34D8F34A8B" bold="true" box="[151,748,1485,1510]" pageId="1" pageNumber="139">
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Genus
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<taxonomicName id="4C324D22B22B3A6BFFB9FA34D8634A8B" ID-CoL="89QV2" authority="Oberprieler & Yeates" authorityName="Oberprieler & Yeates" box="[234,636,1485,1510]" class="Insecta" family="Athericidae" genus="Notoatherix" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B946EAB3B22B3A6BFFB9FA34DB6A4A8B" bold="true" box="[234,373,1485,1510]" italics="true" pageId="1" pageNumber="139">Notoatherix</emphasis>
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Oberprieler & Yeates
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</taxonomicName>
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<taxonomicNameLabel id="A27557C8B22B3A6BFDD0FA34D8F34A8B" box="[643,748,1485,1510]" pageId="1" pageNumber="139" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D36A1B22B3A6BFFC4FA08DB3F4967" blockId="1.[151,748,1485,1546]" box="[151,288,1521,1546]" pageId="1" pageNumber="139">
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(
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<figureCitation id="13092A24B22B3A6BFFCCFA08DAFA4967" box="[159,229,1521,1546]" captionStart="FIGURE 1" captionStartId="2.[151,250,2003,2025]" captionTargetBox="[163,1424,334,1950]" captionTargetId="figure@2.[151,1436,331,1978]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of Notoatherix antiqua gen. et sp. nov., holotype; A — Part, B — Counterpart." httpUri="https://zenodo.org/record/228761/files/figure.png" pageId="1" pageNumber="139">Figs 1</figureCitation>
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& 2)
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</paragraph>
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</subSubSection>
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<subSubSection id="C328652AB22B3A6BFFC4F9C0D967493F" box="[151,888,1593,1618]" pageId="1" pageNumber="139" type="materials_examined">
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<paragraph id="8B8D36A1B22B3A6BFFC4F9C0D967493F" blockId="1.[151,1437,1593,2038]" box="[151,888,1593,1618]" pageId="1" pageNumber="139">
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<emphasis id="B946EAB3B22B3A6BFFC4F9C0DB34493F" bold="true" box="[151,299,1593,1618]" pageId="1" pageNumber="139">
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<typeStatus id="54898803B22B3A6BFFC4F9C0DACD493F" box="[151,210,1593,1618]" pageId="1" pageNumber="139">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C324D22B22B3A6BFE6AF9C0D90C493C" authority="Oberprieler & Yeates" authorityName="Oberprieler & Yeates" box="[313,787,1593,1618]" class="Insecta" family="Athericidae" genus="Notoatherix" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="species" species="antiqua" status="sp. nov.">
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<emphasis id="B946EAB3B22B3A6BFE6AF9C0D804493C" box="[313,539,1593,1617]" italics="true" pageId="1" pageNumber="139">Notoatherix antiqua</emphasis>
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Oberprieler & Yeates
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</taxonomicName>
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,
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<emphasis id="B946EAB3B22B3A6BFC4DF9C0D967493F" bold="true" box="[798,888,1593,1618]" pageId="1" pageNumber="139">
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<taxonomicNameLabel id="A27557C8B22B3A6BFC4DF9C0D967493F" box="[798,888,1593,1618]" pageId="1" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328652AB22B3A6BFF94F9A4D8774855" pageId="1" pageNumber="139" type="diagnosis">
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<paragraph id="8B8D36A1B22B3A6BFF94F9A4D9F5487D" blockId="1.[151,1437,1593,2038]" pageId="1" pageNumber="139">
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<emphasis id="B946EAB3B22B3A6BFF94F9A4DB5E491B" bold="true" box="[199,321,1629,1654]" pageId="1" pageNumber="139">Diagnosis.</emphasis>
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Only a single wing blade is preserved, with the anterior base obscured. R1 and R2+3 relatively straight, R2+3 meeting R1 at costa. R4+5 proximally straight, then weakly curved upwards posteriorly, meeting costa of wing just before apex. Crossvein r-m almost in middle of cell
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<emphasis id="B946EAB3B22B3A6BFC1EF952D94449AE" box="[845,859,1707,1731]" italics="true" pageId="1" pageNumber="139">d</emphasis>
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, but tending slightly towards its basal end. Vein M1 sinuous at base, remainder almost straight, meeting wing margin just behind wing apex. M2 relatively straight, meeting wing margin. Crossvein m-m closing cell
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<emphasis id="B946EAB3B22B3A6BFD9DF900D8C3487C" box="[718,732,1785,1809]" italics="true" pageId="1" pageNumber="139">d</emphasis>
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at level of origin of M1.
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</paragraph>
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<paragraph id="8B8D36A1B22B3A6BFF94F8E6D8774855" blockId="1.[151,1437,1593,2038]" box="[199,616,1823,1848]" pageId="1" pageNumber="139">
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<emphasis id="B946EAB3B22B3A6BFF94F8E6DB8C4855" bold="true" box="[199,403,1823,1848]" pageId="1" pageNumber="139">Species included.</emphasis>
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<typeStatus id="54898803B22B3A6BFEC9F8D9DBCD4855" box="[410,466,1824,1848]" pageId="1" pageNumber="139">Type</typeStatus>
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species only.
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</paragraph>
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</subSubSection>
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<subSubSection id="C328652AB22B3A69FF94F8BAD88B4CBD" lastPageId="3" lastPageNumber="141" pageId="1" pageNumber="139" type="discussion">
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<paragraph id="8B8D36A1B22B3A68FF94F8BADE5A4E4F" blockId="1.[151,1437,1593,2038]" lastBlockId="2.[151,1437,151,290]" lastPageId="2" lastPageNumber="140" pageId="1" pageNumber="139">
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<emphasis id="B946EAB3B22B3A6BFF94F8BADB244831" bold="true" box="[199,315,1859,1884]" pageId="1" pageNumber="139">Remarks.</emphasis>
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Venational features characteristic of the
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<taxonomicName id="4C324D22B22B3A6BFC71F8BAD9BA4831" box="[802,933,1859,1884]" class="Insecta" family="Athericidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="family">Athericidae</taxonomicName>
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were defined by
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<bibRefCitation id="EFA34B50B22B3A6BFB2FF8BADF7E4831" author="Stuckenberg" box="[1148,1377,1859,1884]" pageId="1" pageNumber="139" refString="Stuckenberg, B. R. (1973) The Athericidae, a new family in the lower Brachycera (Diptera). Annals of the Natal Museum, 21, 649 - 673." type="journal article" year="1973">Stuckenberg (1973)</bibRefCitation>
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. The diagnostic feature of the family, the close terminal convergence of veins R1 and R2+3, occurs in
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<emphasis id="B946EAB3B22B3A6BFBBEF89EDF644812" box="[1261,1403,1895,1919]" italics="true" pageId="1" pageNumber="139">Notoatherix.</emphasis>
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In addition, crossvein r-m meets the discal cell further from its basal end, another feature to distinguish rhagionids and athericids used by
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<bibRefCitation id="EFA34B50B22B3A6BFE3EF84BD85048A6" author="Stuckenberg" box="[365,591,1970,1995]" pageId="1" pageNumber="139" refString="Stuckenberg, B. R. (1973) The Athericidae, a new family in the lower Brachycera (Diptera). Annals of the Natal Museum, 21, 649 - 673." type="journal article" year="1973">Stuckenberg (1973)</bibRefCitation>
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. Two other features of many athericids, veins R4 and R5 diverging at a far smaller angle than in rhagionids and tabanids, and R4 ending before the tip of the wing (
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<bibRefCitation id="EFA34B50B22B3A6BFB22F820DF5E489C" author="Stuckenberg" box="[1137,1345,2009,2034]" pageId="1" pageNumber="139" refString="Stuckenberg, B. R. (1973) The Athericidae, a new family in the lower Brachycera (Diptera). Annals of the Natal Museum, 21, 649 - 673." type="journal article" year="1973">Stuckenberg 1973</bibRefCitation>
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), do not occur in
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<emphasis id="B946EAB3B2283A68FFAAFF61DB604FDD" box="[249,383,152,176]" italics="true" pageId="2" pageNumber="140">Notoatherix</emphasis>
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because R4 and R5 are fused throughout their length.
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<bibRefCitation id="EFA34B50B2283A68FC85FF6EDEC24FDD" box="[982,1245,151,176]" pageId="2" pageNumber="140" refString="Mostovski, M. B., Jarzembowski, E. A. & Coram, R. A. (2003) Horseflies and athericids (Diptera: Tabanidae, Athericidae) from the lower Cretaceous of England and Transbaikalia. Paleontological Journal C / C of Paleontologicheskii Zhurnal, 37 (2), 162 - 169." type="journal article">
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Mostovski
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<emphasis id="B946EAB3B2283A68FB00FF60DE934FDD" box="[1107,1164,152,176]" italics="true" pageId="2" pageNumber="140">et al.</emphasis>
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(2003)
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</bibRefCitation>
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reported that the conformation of R4 and R5 is more variable in the fossil record of the three families than in extant forms.
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<emphasis id="B946EAB3B2283A68FFC4FF1CDB024F90" box="[151,285,229,253]" italics="true" pageId="2" pageNumber="140">Notoatherix</emphasis>
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dates to an age when these three families are believed to have diverged (
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<bibRefCitation id="EFA34B50B2283A68FB37FF1CDF7C4F93" author="Wiegmann" box="[1124,1379,229,254]" pageId="2" pageNumber="140" refString="Wiegmann, B. M., Trautwein, M. D., Winkler, I. S., Barr, N. B., Kim, J., Lambkin, C., Bertone, M. A., Cassel, B. K., Bayless, K. M., Heimberg, A. M., Wheeler, B. M., Peterson, K. J., Pape, T., Sinclair, B. J., Skevington, J. H., Blagoderov, V., Caravas, J., Kutty, S. N., Schmidt-Ott, U., Kampmeier, E., Thompson, F. C., Grimaldi, D. A., Beckenbach, A. T., Courtney, G. W., Friedrich, M., Meier, R. & Yeates, D. K. (2011) Episodic radiations in the fly tree of life. Proceedings of the National Academy of Science of the U. S. A., 108, 5690 - 5695. http: // dx. doi. org / 10.1073 / pnas. 1012675108" type="journal article" year="2011">
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Wiegmann
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<emphasis id="B946EAB3B2283A68FBBBFF1FDF3D4F90" box="[1256,1314,229,253]" italics="true" pageId="2" pageNumber="140">et al.</emphasis>
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2011
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</bibRefCitation>
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) and could possess a venation different from that of the current
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<taxonomicName id="4C324D22B2283A68FC76FEF0D9B74E4F" box="[805,936,265,290]" class="Insecta" family="Athericidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="family">Athericidae</taxonomicName>
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crown group.
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</paragraph>
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<caption id="DF4D6629B2283A68FFC4F82ADEB74884" box="[151,1192,2003,2025]" httpUri="https://zenodo.org/record/228761/files/figure.png" pageId="2" pageNumber="140" targetBox="[163,1424,334,1950]" targetPageId="2">
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<paragraph id="8B8D36A1B2283A68FFC4F82ADEB74884" blockId="2.[151,1192,2003,2025]" box="[151,1192,2003,2025]" pageId="2" pageNumber="140">
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<emphasis id="B946EAB3B2283A68FFC4F82ADB0C4885" bold="true" box="[151,275,2003,2025]" pageId="2" pageNumber="140">FIGURE 1.</emphasis>
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Photographs of
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<taxonomicName id="4C324D22B2283A68FEEAF82AD89A4884" box="[441,645,2003,2025]" class="Insecta" family="Athericidae" genus="Notoatherix" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="species" species="antiqua">
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<emphasis id="B946EAB3B2283A68FEEAF82AD89A4884" box="[441,645,2003,2025]" italics="true" pageId="2" pageNumber="140">Notoatherix antiqua</emphasis>
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</taxonomicName>
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<emphasis id="B946EAB3B2283A68FDDFF82AD9364885" bold="true" box="[652,809,2003,2024]" pageId="2" pageNumber="140">gen. et sp. nov.</emphasis>
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, holotype;
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<emphasis id="B946EAB3B2283A68FCCFF82AD9B14885" bold="true" box="[924,942,2003,2024]" pageId="2" pageNumber="140">A</emphasis>
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—Part,
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<emphasis id="B946EAB3B2283A68FCAEF82ADE114885" bold="true" box="[1021,1038,2003,2024]" pageId="2" pageNumber="140">B</emphasis>
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—Counterpart.
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</paragraph>
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</caption>
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<paragraph id="8B8D36A1B2293A69FFC4FCEED88B4CBD" blockId="3.[151,1436,791,976]" pageId="3" pageNumber="141">
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<emphasis id="B946EAB3B2293A69FFC4FCEEDB104C41" bold="true" box="[151,271,791,813]" pageId="3" pageNumber="141">FIGURE 2</emphasis>
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. Line drawing of wing of venation of
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<taxonomicName id="4C324D22B2293A69FDF7FCEED96D4C40" box="[676,882,791,813]" class="Insecta" family="Athericidae" genus="Notoatherix" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="antiqua">
|
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<emphasis id="B946EAB3B2293A69FDF7FCEED96D4C40" box="[676,882,791,813]" italics="true" pageId="3" pageNumber="141">Notoatherix antiqua</emphasis>
|
||
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</taxonomicName>
|
||
|
|
<emphasis id="B946EAB3B2293A69FC28FCEEDE004C41" bold="true" box="[891,1055,791,812]" pageId="3" pageNumber="141">gen. et sp. nov.</emphasis>
|
||
|
|
,
|
||
|
|
<typeStatus id="54898803B2293A69FB7DFCEEDE954C40" box="[1070,1162,791,813]" pageId="3" pageNumber="141" type="holotype">holotype</typeStatus>
|
||
|
|
; composite, from part and counterpart. Abbreviations: C = costa; Sc = subcosta; R = radius; R2+3 = anterior branch of the radial sector; R4+5 = posterior branch of the radial sector; M1, M2 = anterior (sectoral) branches of media; M3 = posterior (sectoral) branch of media; CuA1, CuA2 = anterior branches of cubitus; A = anal vein; r-m = radial to medial cross-vein; m-m = medial to medial cross-vein; m-cu = medial to cubital cross-vein;
|
||
|
|
<emphasis id="B946EAB3B2293A69FE9DFC60DBC44CC2" box="[462,475,921,943]" italics="true" pageId="3" pageNumber="141">d</emphasis>
|
||
|
|
= discal cell;
|
||
|
|
<emphasis id="B946EAB3B2293A69FD34FC60D8614CC2" box="[615,638,921,943]" italics="true" pageId="3" pageNumber="141">br</emphasis>
|
||
|
|
= first basal cell;
|
||
|
|
<emphasis id="B946EAB3B2293A69FC62FC60D94F4CC2" box="[817,848,921,943]" italics="true" pageId="3" pageNumber="141">bm</emphasis>
|
||
|
|
= second basal cell;
|
||
|
|
<emphasis id="B946EAB3B2293A69FB73FC63DE5A4CDE" box="[1056,1093,922,947]" italics="true" pageId="3" pageNumber="141">m1-3</emphasis>
|
||
|
|
= first to fourth medial cells;
|
||
|
|
<emphasis id="B946EAB3B2293A69FA23FC63DF834CDE" box="[1392,1436,922,947]" italics="true" pageId="3" pageNumber="141">cua1</emphasis>
|
||
|
|
= anterior cubital cell;
|
||
|
|
<emphasis id="B946EAB3B2293A69FE2EFC42DBBD4CBD" box="[381,418,955,976]" italics="true" pageId="3" pageNumber="141">cup</emphasis>
|
||
|
|
= posterior cubital cell.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
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</document>
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