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<document id="32130F085FB7BC6433B8DDA96193B9B8" ID-DOI="10.11646/zootaxa.3686.2.1" ID-GBIF-Dataset="4d62d5da-3890-4af3-9758-674a13b43a79" ID-ISSN="1175-5326" ID-Zenodo-Dep="221213" ID-ZooBank="1AF48240-6A27-452E-9D32-9C6D45C715E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460646465142" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Kurina, Olavi" docDate="2013" docId="039B87AB0058FFD337E775E2FB4818DE" docLanguage="en" docName="zt03686p139.pdf" docOrigin="Zootaxa 3686 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manota coxata Enderlein" docType="treatment" docVersion="8" lastPageNumber="105" masterDocId="FFA2FFD3005BFFD73770753FFF811A31" masterDocTitle="New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region" masterLastPageNumber="139" masterPageNumber="101" pageNumber="104" updateTime="1698337046318" updateUser="plazi">
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<mods:title id="04C74564FA0B54438172AA5F8B52B216">New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region</mods:title>
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<mods:namePart id="08C0A4D60B6EEE3C874D45B1D5EAD230">Hippa, Heikki</mods:namePart>
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<mods:namePart id="E7DDC586FB3253FE19449FBC15919700">Kurina, Olavi</mods:namePart>
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<mods:date id="E0C6D3D3C662576744A9B2AA99C63B68">2013</mods:date>
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<treatment id="039B87AB0058FFD337E775E2FB4818DE" ID-DOI="http://doi.org/10.5281/zenodo.6155058" ID-GBIF-Taxon="119572312" ID-Zenodo-Dep="6155058" LSID="urn:lsid:plazi:treatment:039B87AB0058FFD337E775E2FB4818DE" httpUri="http://treatment.plazi.org/id/039B87AB0058FFD337E775E2FB4818DE" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
<subSubSection id="C32865360058FFD437E775E2FCE31AC6" box="[151,866,221,247]" pageId="3" pageNumber="104" type="nomenclature">
<paragraph id="8B8D36BD0058FFD437E775E2FCE31AC6" blockId="3.[151,866,221,247]" box="[151,866,221,247]" pageId="3" pageNumber="104">
<heading id="D0C581D10058FFD437E775E2FCE31AC6" bold="true" box="[151,866,221,247]" fontSize="11" level="1" pageId="3" pageNumber="104" reason="1">
<emphasis id="B946EAAF0058FFD437E775E2FCE31AC6" bold="true" box="[151,866,221,247]" pageId="3" pageNumber="104">
Notes on
<taxonomicName id="4C324D3E0058FFD4367A75E2FDD41AC6" ID-CoL="3XVVH" authority="Enderlein" authorityName="Enderlein" box="[266,597,221,247]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF0058FFD4367A75E2FE3E1AC6" bold="true" box="[266,447,221,247]" italics="true" pageId="3" pageNumber="104">Manota coxata</emphasis>
(Enderlein)
</taxonomicName>
and
<taxonomicName id="4C324D3E0058FFD435E475E2FCE31AC6" authority="Lane" authorityName="Lane" box="[660,866,221,247]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="B946EAAF0058FFD435E475E2FC9B1AC6" bold="true" box="[660,794,221,247]" italics="true" pageId="3" pageNumber="104">M. palpalis</emphasis>
Lane
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<caption id="DF4D66350058FFD437E7724BFE961DF9" httpUri="https://zenodo.org/record/221215/files/figure.png" pageId="3" pageNumber="104" targetBox="[151,1436,299,1886]" targetPageId="3">
<paragraph id="8B8D36BD0058FFD437E7724BFE961DF9" blockId="3.[151,1436,1908,1992]" pageId="3" pageNumber="104">
<emphasis id="B946EAAF0058FFD437E7724BFE951DBB" bold="true" box="[151,276,1908,1931]" pageId="3" pageNumber="104">FIGURE 2.</emphasis>
<taxonomicName id="4C324D3E0058FFD4366B7249FDB21DBA" authority="Enderlein" authorityName="Enderlein" box="[283,563,1909,1931]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF0058FFD4366B7249FE341DBA" box="[283,437,1910,1931]" italics="true" pageId="3" pageNumber="104">Manota coxata</emphasis>
(Enderlein)
</taxonomicName>
.
<emphasis id="B946EAAF0058FFD4354D724AFDD61DBB" bold="true" box="[573,599,1909,1930]" pageId="3" pageNumber="104">A.</emphasis>
Double pinned holotype with labels.
<emphasis id="B946EAAF0058FFD434AA724AFC701DBB" bold="true" box="[986,1009,1909,1930]" pageId="3" pageNumber="104">B.</emphasis>
Remains of the holotype, closer view.
<emphasis id="B946EAAF0058FFD432F3724BFA1D1DBA" bold="true" box="[1411,1436,1908,1931]" pageId="3" pageNumber="104">C.</emphasis>
Abdomen mounted on a piece of celluloid.
<emphasis id="B946EAAF0058FFD4353E72ABFDE61D98" bold="true" box="[590,615,1940,1961]" pageId="3" pageNumber="104">D.</emphasis>
Enderleins handwritten determination label.
<emphasis id="B946EAAF0058FFD4334772ABFBCF1D98" bold="true" box="[1079,1102,1940,1961]" pageId="3" pageNumber="104">E.</emphasis>
Small board box with the mount of one wing.
</paragraph>
</caption>
<subSubSection id="C3286536005FFFD337E775A8FA06180D" pageId="4" pageNumber="105" type="description">
<paragraph id="8B8D36BD005FFFD337E775A8FB291AC6" blockId="4.[151,1437,151,752]" pageId="4" pageNumber="105">
Prior to the study of
<taxonomicName id="4C324D3E005FFFD336E475A6FE6A1A81" box="[404,491,153,176]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAAF005FFFD336E475A6FE6A1A81" box="[404,491,153,176]" italics="true" pageId="4" pageNumber="105">Manota</emphasis>
</taxonomicName>
in
<collectingCountry id="F325762D005FFFD3356875A8FD191A9E" box="[536,664,151,175]" name="Costa Rica" pageId="4" pageNumber="105">Costa Rica</collectingCountry>
(
<bibRefCitation id="EFA34B4C005FFFD335DA75A8FC461A9E" author="Jaschhof" box="[682,967,151,175]" pageId="4" pageNumber="105" refString="Jaschhof, M. &amp; Hippa, H. (2005) The genus Manota in Costa Rica (Diptera: Mycetophilidae). Zootaxa, 1011, 1 - 54." type="journal article" year="2005">Jaschhof &amp; Hippa 2005</bibRefCitation>
), three species were described from the Neotropical region.
<bibRefCitation id="EFA34B4C005FFFD336077583FD121AE5" author="Jaschhof" box="[375,659,188,212]" pageId="4" pageNumber="105" refString="Jaschhof, M. &amp; Hippa, H. (2005) The genus Manota in Costa Rica (Diptera: Mycetophilidae). Zootaxa, 1011, 1 - 54." type="journal article" year="2005">Jaschhof &amp; Hippa (2005)</bibRefCitation>
studied the
<typeStatus id="5489881F005FFFD3346B7582FCCA1AE4" box="[795,843,189,213]" pageId="4" pageNumber="105">type</typeStatus>
material of
<taxonomicName id="4C324D3E005FFFD334A37582FBC81AE5" box="[979,1097,188,212]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="defecta">
<emphasis id="B946EAAF005FFFD334A37582FBC81AE5" box="[979,1097,188,212]" italics="true" pageId="4" pageNumber="105">M. defecta</emphasis>
</taxonomicName>
described from the Caribbean Islands (
<bibRefCitation id="EFA34B4C005FFFD3378575E0FE1C1AC6" author="Williston" box="[245,413,223,247]" pageId="4" pageNumber="105" refString="Williston, S. W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Royal Entomological Society of London, 1896, 253 - 446." type="journal article" year="1896">Williston 1896</bibRefCitation>
) but not that of the two other species, viz.
<taxonomicName id="4C324D3E005FFFD3340C75DEFC6A1AC9" box="[892,1003,225,248]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF005FFFD3340C75DEFC6A1AC9" box="[892,1003,225,248]" italics="true" pageId="4" pageNumber="105">M. coxata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D3E005FFFD3335275DEFB201AC9" box="[1058,1185,224,248]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="B946EAAF005FFFD3335275DEFB201AC9" box="[1058,1185,224,248]" italics="true" pageId="4" pageNumber="105">M. palpalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8D36BD005FFFD337B7743AFA06180D" blockId="4.[151,1437,151,752]" pageId="4" pageNumber="105">
<taxonomicName id="4C324D3E005FFFD337B7743AFEF41B2D" box="[199,373,261,284]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF005FFFD337B7743AFEF41B2D" box="[199,373,261,284]" italics="true" pageId="4" pageNumber="105">Manota coxata</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C324D3E005FFFD336DD743BFD371B2D" box="[429,694,260,284]" class="Insecta" family="Mycetophilidae" genus="Aphanizophleps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF005FFFD336DD743BFD371B2D" box="[429,694,260,284]" italics="true" pageId="4" pageNumber="105">Aphanizophleps coxata</emphasis>
</taxonomicName>
) was described from the
<typeStatus id="5489881F005FFFD3349C743BFBCF1B2D" box="[1004,1102,260,284]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
male from Santa Catharina,
<collectingCountry id="F325762D005FFFD337E77418FF5A1B0E" box="[151,219,295,319]" name="Brazil" pageId="4" pageNumber="105">Brazil</collectingCountry>
(
<bibRefCitation id="EFA34B4C005FFFD3379C7418FE1D1B0E" author="Enderlein" box="[236,412,295,319]" pageId="4" pageNumber="105" refString="Enderlein, G. (1911) Neue Gattungen und Arten aussereuropaischer Fliegen. Stettin entomologische Zeitung, 72, 135 - 209." type="journal article" year="1911">Enderlein 1911</bibRefCitation>
) and has not been reported since. The detailed description, including a drawing of wing, does not give any characters of the hypopygium which would be necessary for identification of the species. We have studied the
<typeStatus id="5489881F005FFFD3362A7450FE3D1BB6" box="[346,444,367,391]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
(in MIZ) that is in a relatively poor condition (see
<figureCitation id="13092A38005FFFD3337D7450FBD21BB6" box="[1037,1107,367,391]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1931]" captionTargetBox="[151,1436,299,1886]" captionTargetId="figure@3.[151,1436,299,1887]" captionTargetPageId="3" captionText="FIGURE 2. Manota coxata (Enderlein). A. Double pinned holotype with labels. B. Remains of the holotype, closer view. C. Abdomen mounted on a piece of celluloid. D. Enderleins handwritten determination label. E. Small board box with the mount of one wing." httpUri="https://zenodo.org/record/221215/files/figure.png" pageId="4" pageNumber="105">Fig. 2</figureCitation>
). It is pinned through thorax which bears the head with incomplete antennae and palpi, all coxae and the complete left foreleg. One wing is mounted separately between two cover slips and is stored in a small board box next to the specimen. The abdomen is mounted, probably in
<collectingCountry id="F325762D005FFFD336DD74E3FD801BC5" box="[429,513,476,500]" name="Canada" pageId="4" pageNumber="105">Canada</collectingCountry>
balsam, on a piece of celluloid attached to the pin with the other dry parts of the specimen but the hypopygium is lacking. The hypopygium has obviously been slide-mounted separately but has lost the contact with the other parts of the specimen. Therefore, the identity of
<taxonomicName id="4C324D3E005FFFD33377771AFBF6180D" box="[1031,1143,549,572]" class="Insecta" family="Mycetophilidae" genus="Aphanizophleps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="coxata">
<emphasis id="B946EAAF005FFFD33377771AFBF6180D" box="[1031,1143,549,572]" italics="true" pageId="4" pageNumber="105">M. coxata</emphasis>
</taxonomicName>
remains currently open.
</paragraph>
</subSubSection>
<subSubSection id="C3286536005FFFD337B77776FB4818DE" pageId="4" pageNumber="105" type="reference_group">
<paragraph id="8B8D36BD005FFFD337B77776FB4818DE" blockId="4.[151,1437,151,752]" pageId="4" pageNumber="105">
<taxonomicName id="4C324D3E005FFFD337B77776FE011851" ID-CoL="3XVXM" box="[199,384,584,608]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="B946EAAF005FFFD337B77776FE011851" box="[199,384,584,608]" italics="true" pageId="4" pageNumber="105">Manota palpalis</emphasis>
</taxonomicName>
was described from the
<typeStatus id="5489881F005FFFD335EB7778FD7C186E" box="[667,765,583,607]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
male from Estado de Goiás, Corumbá,
<collectingCountry id="F325762D005FFFD333B47778FA89186E" box="[1220,1288,583,607]" name="Brazil" pageId="4" pageNumber="105">Brazil</collectingCountry>
(
<bibRefCitation id="EFA34B4C005FFFD332697778FA15186E" author="Lane" box="[1305,1428,583,607]" pageId="4" pageNumber="105" refString="Lane, J. (1948) &quot; Mycetophilidae &quot; do Brasil (Diptera, Nemocera). Revista Brasileira de Biologia, 8 (2), 247 - 254." type="journal article" year="1948">Lane 1948</bibRefCitation>
) and has not reported since. The description includes a sufficiently detailed drawing of male hypopygium (cf.
<bibRefCitation id="EFA34B4C005FFFD332147752FF521896" author="Lane" pageId="4" pageNumber="105" refString="Lane, J. (1948) &quot; Mycetophilidae &quot; do Brasil (Diptera, Nemocera). Revista Brasileira de Biologia, 8 (2), 247 - 254." type="journal article" year="1948">Lane 1948</bibRefCitation>
: fig. 4) to allow the identification of the species when at hand. The row of six hook-like structures at the posterodorsal margin of the gonocoxa, probably megasetae, are quite unlike what we have seen in any of the species we studied and we have regarded it unnecessary to study the
<typeStatus id="5489881F005FFFD334EA77E7FC4B18C1" box="[922,970,728,752]" pageId="4" pageNumber="105">type</typeStatus>
in the present context.
</paragraph>
</subSubSection>
</treatment>
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