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<document id="2ED642CE177A7CFEC643AA2CB066375A" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF40CDA4FF4CEDCFB551EEE0" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Acanthophorella irystoni" docType="treatment" docVersion="8" lastPageNumber="155" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="152" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="F7AD7E552A641AE84194DD60F0696FDB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2BDE8F0EBCB6772F850CEAC6907735F6">
<mods:title id="9D846CC09F6D8CF2A25EBD8049DBF331">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="1CD7078E0192CF9B96BC2B0286E3C2BD" type="personal">
<mods:role id="EB47ACF37566690E56C8DCC971893BF3">
<mods:roleTerm id="A2DB3A662473556918BB89526EE265BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33BE7DAE12CD24666BC992FE69A9C2AA">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="26325A3457F151BCBC1D8ECE788EE29E" type="personal">
<mods:role id="235729D4C220371EBAFCC8229F2DCC73">
<mods:roleTerm id="E38357016F1793A8C2E291E3A2CBDEDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05045D44838985D8B7A618C42E3C7D7A">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FA9B345550BF9E7812FF7582CA99FBC8">text</mods:typeOfResource>
<mods:relatedItem id="3516A96CE63263FCC5FAF14A849BBD8B" type="host">
<mods:titleInfo id="E7AA5180431A37495828C9BE70CD70DB">
<mods:title id="4B268E8D1A380BE535AA4528A2ED9BB0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1007991D6056CEFC6D5B92E2972273E5">
<mods:date id="C3FB9EB2AC48A9D238B5C70A7DE51DE8">2016</mods:date>
<mods:detail id="4B7DCCC1C5698571873A3BF455C0FB97" type="volume">
<mods:number id="34748F609BDB56B96F7EC6FD4050A9F3">4211</mods:number>
</mods:detail>
<mods:detail id="931661E14C04C326D81979B50A15A9BF" type="issue">
<mods:number id="4E774EEFBD199FE6E3201B7C6F0C742B">1</mods:number>
</mods:detail>
<mods:extent id="3E70A4EDE0A5A61EA6E0230B401EEA0F" unit="page">
<mods:start id="40E872A89760786E8710BA802FDBA17A">1</mods:start>
<mods:end id="788F17A1C5CEBF1E11A543EAB70E9277">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3B9E9284316BB17A9AFAE7773421095F">journal article</mods:classification>
<mods:identifier id="D5363F825564DC46F50EEFA8225E9216" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="205F67AF7C24566385E81B3D0883C507" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="2FCA5999112EDDF1B17D97C7883B7BBF" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="A9DFC2B54D1389A103A70AFACA69F87B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3CADA85B1378D2F3C6F4EC395E5DF90D" type="Zenodo-Dep">272750</mods:identifier>
<mods:identifier id="91C5AE681977BF94BD7672FF846482E2" type="ZooBank">6B86C6BA-6AFE-4AAD-870D-04794C138D47</mods:identifier>
</mods:mods>
<treatment id="03B7878CFF40CDA4FF4CEDCFB551EEE0" ID-DOI="http://doi.org/10.5281/zenodo.6074470" ID-GBIF-Taxon="127419187" ID-Zenodo-Dep="6074470" LSID="urn:lsid:plazi:treatment:03B7878CFF40CDA4FF4CEDCFB551EEE0" httpUri="http://treatment.plazi.org/id/03B7878CFF40CDA4FF4CEDCFB551EEE0" lastPageId="154" lastPageNumber="155" pageId="151" pageNumber="152">
<subSubSection id="C3046511FF40CDA9FF4CEDCFB043ECBF" pageId="151" pageNumber="152" type="nomenclature">
<paragraph id="8BA1369AFF40CDA9FF4CEDCFB3F1EC9C" blockId="151.[151,642,1466,1526]" box="[151,642,1466,1493]" pageId="151" pageNumber="152">
<heading id="D0E981F6FF40CDA9FF4CEDCFB3F1EC9C" bold="true" box="[151,642,1466,1493]" fontSize="11" level="1" pageId="151" pageNumber="152" reason="1">
<emphasis id="B96AEA88FF40CDA9FF4CEDCFB3F1EC9C" bold="true" box="[151,642,1466,1493]" pageId="151" pageNumber="152">
<taxonomicName id="4C1E4D19FF40CDA9FF4CEDCFB0B6EC9D" ID-CoL="8466Y" box="[151,453,1466,1492]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="irystoni">
<emphasis id="B96AEA88FF40CDA9FF4CEDCFB0B6EC9D" bold="true" box="[151,453,1466,1492]" italics="true" pageId="151" pageNumber="152">Acanthophorella irystoni</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF40CDA9FF4CEDA8B043ECBF" blockId="151.[151,642,1466,1526]" box="[151,304,1501,1526]" pageId="151" pageNumber="152">
<figureCitation id="13252A1FFF40CDA9FF4CEDA8B043ECBF" box="[151,304,1501,1526]" captionStart-0="FIGURE 126" captionStart-1="FIGURE 127" captionStart-2="FIGURE 128" captionStartId-0="152.[151,250,1264,1286]" captionStartId-1="153.[151,250,1965,1987]" captionStartId-2="154.[151,250,1453,1475]" captionTargetBox-0="[180,1404,337,1250]" captionTargetBox-1="[228,1345,193,1938]" captionTargetBox-2="[208,1381,224,1348]" captionTargetId-0="figure@152.[180,1404,337,1250]" captionTargetId-1="figure@153.[228,1345,193,1938]" captionTargetId-2="figure@154.[188,1398,193,1432]" captionTargetPageId-0="152" captionTargetPageId-1="153" captionTargetPageId-2="154" captionText-0="FIGURE 126. Acanthophorella irystoni gen. et sp. nov., paratype male habitus. Scale line = 1 mm." captionText-1="FIGURE 127. Acanthophorella irystoni gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 6. D. Leg 7. E. Leg 10. Scale lines = 0.2 mm." captionText-2="FIGURE 128. Acanthophorella irystoni gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; ll = lateral lamella; mp = medial shield; sp = sternal plate; ss = sternal sac; tp = thorn-like process; us = undefined structure; cvp = chitinised and membranous process; ppr = posterior projection. Scale line = 0.3 mm." httpUri-0="https://zenodo.org/record/272875/files/figure.png" httpUri-1="https://zenodo.org/record/272876/files/figure.png" httpUri-2="https://zenodo.org/record/272877/files/figure.png" pageId="151" pageNumber="152">Figs 126–128</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF40CDA9FF4CEE50B073EFCF" pageId="151" pageNumber="152" type="diagnosis">
<paragraph id="8BA1369AFF40CDA9FF4CEE50B073EFCF" blockId="151.[151,1436,1573,2030]" pageId="151" pageNumber="152">
<emphasis id="B96AEA88FF40CDA9FF4CEE50B062EF77" bold="true" box="[151,273,1573,1598]" pageId="151" pageNumber="152">Diagnosis.</emphasis>
Differs from the other two species of the genus by the presence of denticulate lateral lamellae of the anterior gonopods, by very small coxal processes on the posterior gonopods and by the general shape of the gonopod.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF40CDA9FF1CEEE4B060EF87" pageId="151" pageNumber="152" type="etymology">
<paragraph id="8BA1369AFF40CDA9FF1CEEE4B060EF87" blockId="151.[151,1436,1573,2030]" pageId="151" pageNumber="152">
<emphasis id="B96AEA88FF40CDA9FF1CEEE4B03DEFE3" bold="true" box="[199,334,1681,1706]" pageId="151" pageNumber="152">Etymology.</emphasis>
The new species is named after its
<typeStatus id="54A58838FF40CDA9FD04EEE7B27CEFE3" box="[735,783,1682,1706]" pageId="151" pageNumber="152">type</typeStatus>
locality,
<collectingRegion id="49DAF878FF40CDA9FCACEEE4B2B9EFE3" box="[887,970,1681,1706]" country="Russia" name="North Ossetia" pageId="151" pageNumber="152">Ossetia</collectingRegion>
(Iryston in Ossetian language), a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF40CDA9FF1CEEACB0B6EE83" pageId="151" pageNumber="152" type="materials_examined">
<paragraph id="8BA1369AFF40CDA9FF1CEEACB3E9EE73" blockId="151.[151,1436,1573,2030]" pageId="151" pageNumber="152">
<materialsCitation id="3B763CC7FF40CDA9FF1CEEACB3E9EE73" ID-GBIF-Occurrence="1424566943" collectingDate="1982-06-02" collectionCode="ZMUM" collectorName="S. Golovatch" country="Russia" location="Ordzhonikidze" municipality="Material" pageId="151" pageNumber="152" specimenCount="7" specimenCount-female="3" specimenCount-juvenile="2" specimenCount-male="2" stateProvince="North Ossetia" typeStatus="holotype">
<emphasis id="B96AEA88FF40CDA9FF1CEEACB0E1EFBB" bold="true" box="[199,402,1753,1778]" pageId="151" pageNumber="152">
<collectingMunicipality id="6BC5ACE0FF40CDA9FF1CEEACB05CEFBB" box="[199,303,1753,1778]" pageId="151" pageNumber="152">Material</collectingMunicipality>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF40CDA9FE37EEAFB33AEFBB" box="[492,585,1753,1778]" pageId="151" pageNumber="152" type="male">2 males</specimenCount>
,
<specimenCount id="9D18FD13FF40CDA9FD81EEAFB3BEEFBB" box="[602,717,1753,1778]" pageId="151" pageNumber="152" type="female">3 females</specimenCount>
,
<specimenCount id="9D18FD13FF40CDA9FD06EEAFB212EFBB" box="[733,865,1753,1778]" pageId="151" pageNumber="152" type="juvenile">2 juveniles</specimenCount>
).
<emphasis id="B96AEA88FF40CDA9FCA3EEACB29FEFBB" bold="true" box="[888,1004,1753,1778]" pageId="151" pageNumber="152">
<typeStatus id="54A58838FF40CDA9FCA3EEACB294EFBB" box="[888,999,1753,1778]" pageId="151" pageNumber="152" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF40CDA9FC2CEEAFB52CEFBB" box="[1015,1119,1754,1778]" name="Russia" pageId="151" pageNumber="152">RUSSIA</collectingCountry>
: male,
<collectingRegion id="49DAF878FF40CDA9FB6CEEACB42AEFBB" box="[1207,1369,1753,1778]" country="Russia" name="North Ossetia" pageId="151" pageNumber="152">North Ossetia</collectingRegion>
, S of
<location id="8EC16041FF40CDA9FF4CEE88B04EEE5F" LSID="urn:lsid:plazi:treatment:03B7878CFF40CDA4FF4CEDCFB551EEE0:8EC16041FF40CDA9FF4CEE88B04EEE5F" box="[151,317,1789,1814]" country="Russia" municipality="Material" name="Ordzhonikidze" pageId="151" pageNumber="152" stateProvince="North Ossetia">Ordzhonikidze</location>
(= Vladikavkaz),
<locationDeviation id="9A132FB8FF40CDA9FDD4EE88B574EE5F" box="[527,1031,1789,1814]" pageId="151" pageNumber="152">
between Chmi and Baltik,
<taxonomicName id="4C1E4D19FF40CDA9FC99EE8BB2D3EE5C" box="[834,928,1790,1813]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="151" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF40CDA9FC99EE8BB2D3EE5C" box="[834,928,1790,1813]" italics="true" pageId="151" pageNumber="152">Quercus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF40CDA9FC1CEE88B574EE5F" box="[967,1031,1789,1814]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="151" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF40CDA9FC1CEE88B574EE5F" box="[967,1031,1789,1814]" italics="true" pageId="151" pageNumber="152">Alnus</emphasis>
</taxonomicName>
</locationDeviation>
on slope, litter and under stones,
<collectingDate id="EFE4E9B2FF40CDA9FA55EE8BB074EE73" pageId="151" pageNumber="152" value="1982-06-02">
<date id="FFA0105AFF40CDA9FA55EE8BB074EE73" pageId="151" pageNumber="152" value="1982-06-02">2 Jun. 1982</date>
</collectingDate>
,
<collectorName id="26EB534CFF40CDA9FEC9EF57B0D7EE73" box="[274,420,1825,1850]" pageId="151" pageNumber="152">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF40CDA9FE38EF57B34FEE73" box="[483,572,1826,1850]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="151" pageNumber="152">ZMUM</collectionCode>
ρ3403).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF40CDA9FF1CEF30B0AFEECB" blockId="151.[151,1436,1573,2030]" pageId="151" pageNumber="152">
<emphasis id="B96AEA88FF40CDA9FF1CEF30B04CEE17" bold="true" box="[199,319,1861,1886]" pageId="151" pageNumber="152">Paratypes</emphasis>
(total: 1 male, 3 females, 2 juveniles).
<materialsCitation id="3B763CC7FF40CDA9FD2DEF30B0AFEECB" ID-GBIF-Occurrence="1424566926" collectingDate="1982-06-02" collectionCode="ZMUM" collectorName="S. Golovatch" country="Russia" location="All" municipality="Material" pageId="151" pageNumber="152" specimenCount="6" specimenCount-female="3" specimenCount-juvenile="2" specimenCount-male="1" stateProvince="North Ossetia">
<location id="8EC16041FF40CDA9FD2DEF30B269EE17" LSID="urn:lsid:plazi:treatment:03B7878CFF40CDA4FF4CEDCFB551EEE0:8EC16041FF40CDA9FD2DEF30B269EE17" box="[758,794,1861,1886]" country="Russia" name="All" pageId="151" pageNumber="152">All</location>
from
<collectingCountry id="F309760AFF40CDA9FC85EF33B2B5EE17" box="[862,966,1862,1886]" name="Russia" pageId="151" pageNumber="152">RUSSIA</collectingCountry>
:
<specimenCount id="9D18FD13FF40CDA9FC0BEF33B56CEE17" box="[976,1055,1861,1886]" pageId="151" pageNumber="152" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF40CDA9FBF1EF33B5EAEE17" box="[1066,1177,1861,1886]" pageId="151" pageNumber="152" type="female">3 females</specimenCount>
,
<specimenCount id="9D18FD13FF40CDA9FB7FEF33B452EE17" box="[1188,1313,1861,1886]" pageId="151" pageNumber="152" type="juvenile">2 juveniles</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFF40CDA9FEFDEF1FB00CEECB" box="[294,383,1898,1922]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="151" pageNumber="152">ZMUM</collectionCode>
ρ3404).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF40CDA9FF1CEFF8B0B6EE83" blockId="151.[151,1436,1573,2030]" pageId="151" pageNumber="152">
<materialsCitation id="3B763CC7FF40CDA9FF1CEFF8B0B6EE83" ID-GBIF-Occurrence="1424566944" country="Russia" location="Ordzhonikidze" pageId="151" pageNumber="152" specimenCount="1" stateProvince="North Ossetia" typeStatus="holotype">
<emphasis id="B96AEA88FF40CDA9FF1CEFF8B018EEEF" bold="true" box="[199,363,1933,1958]" pageId="151" pageNumber="152">
<typeStatus id="54A58838FF40CDA9FF1CEFF8B071EEEF" box="[199,258,1933,1958]" pageId="151" pageNumber="152">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF40CDA9FEACEFFBB0ACEEEF" box="[375,479,1934,1958]" name="Russia" pageId="151" pageNumber="152">RUSSIA</collectingCountry>
:
<collectingRegion id="49DAF878FF40CDA9FE34EFF8B3E1EEEF" box="[495,658,1933,1958]" country="Russia" name="North Ossetia" pageId="151" pageNumber="152">North Ossetia</collectingRegion>
, S of
<location id="8EC16041FF40CDA9FD38EFF8B2FAEEEF" LSID="urn:lsid:plazi:treatment:03B7878CFF40CDA4FF4CEDCFB551EEE0:8EC16041FF40CDA9FD38EFF8B2FAEEEF" box="[739,905,1933,1958]" country="Russia" name="Ordzhonikidze" pageId="151" pageNumber="152" stateProvince="North Ossetia">Ordzhonikidze</location>
(= Vladikavkaz),
<locationDeviation id="9A132FB8FF40CDA9FBBCEFF8B02BEE83" pageId="151" pageNumber="152">
between Chmi and Baltik,
<taxonomicName id="4C1E4D19FF40CDA9FF4CEFC7B186EE80" box="[151,245,1970,1993]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="151" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF40CDA9FF4CEFC7B186EE80" box="[151,245,1970,1993]" italics="true" pageId="151" pageNumber="152">Quercus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF40CDA9FEC3EFC4B02BEE83" box="[280,344,1969,1994]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="151" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF40CDA9FEC3EFC4B02BEE83" box="[280,344,1969,1994]" italics="true" pageId="151" pageNumber="152">Alnus</emphasis>
</taxonomicName>
</locationDeviation>
on slope.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF40CDA4FF1CEFA0B0BFEECD" lastPageId="154" lastPageNumber="155" pageId="151" pageNumber="152" type="description">
<paragraph id="8BA1369AFF40CDA9FF1CEFA0B2D1EEA7" blockId="151.[151,1436,1573,2030]" box="[199,930,2005,2030]" pageId="151" pageNumber="152">
<emphasis id="B96AEA88FF40CDA9FF1CEFA0B02AEEA7" bold="true" box="[199,345,2005,2030]" pageId="151" pageNumber="152">Description.</emphasis>
Body with 31 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CE8EDB2EBE99D" blockId="152.[151,1436,151,285]" pageId="152" pageNumber="153">
MEASUREMENTS. Males
<quantity id="4CE69B7FFF4FCDA6FE00E8E2B33EE9E6" box="[475,589,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="152" pageNumber="153" unit="mm" value="9.25" valueMax="9.5" valueMin="9.0">9–9.5 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF4FCDA6FB5AE8EDB5A9E9E6" box="[1153,1242,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="152" pageNumber="153" unit="mm" value="0.8">0.8 mm</quantity>
. Females
<quantity id="4CE69B7FFF4FCDA6FA93E8E2B1B0E99D" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="152" pageNumber="153" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF4FCDA6FD20E8C9B2E0E99D" box="[763,915,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="152" pageNumber="153" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.8–0.85 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CE895B57EE855" blockId="152.[151,1436,151,285]" pageId="152" pageNumber="153">
COLORATION (
<figureCitation id="13252A1FFF4FCDA6FEB4E8AAB0A6E9B1" box="[367,469,223,248]" captionStart="FIGURE 126" captionStartId="152.[151,250,1264,1286]" captionTargetBox="[180,1404,337,1250]" captionTargetId="figure@152.[180,1404,337,1250]" captionTargetPageId="152" captionText="FIGURE 126. Acanthophorella irystoni gen. et sp. nov., paratype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272875/files/figure.png" pageId="152" pageNumber="153">Fig. 126</figureCitation>
). Dorsal and lateral sides of prozonites greyish, dorsolateral and ventrolateral sides with yellowish spots. Metazonites brownish, lateral keels with yellowish spots.
</paragraph>
<caption id="DF616612FF4FCDA6FF4CEC85B5F9EC4F" box="[151,1162,1264,1286]" httpUri="https://zenodo.org/record/272875/files/figure.png" pageId="152" pageNumber="153" targetBox="[180,1404,337,1250]" targetPageId="152">
<paragraph id="8BA1369AFF4FCDA6FF4CEC85B5F9EC4F" blockId="152.[151,1162,1264,1286]" box="[151,1162,1264,1286]" pageId="152" pageNumber="153">
<emphasis id="B96AEA88FF4FCDA6FF4CEC85B05FEC4C" bold="true" box="[151,300,1264,1286]" pageId="152" pageNumber="153">FIGURE 126.</emphasis>
<taxonomicName id="4C1E4D19FF4FCDA6FEE9EC85B35CEC4F" box="[306,559,1264,1286]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="irystoni">
<emphasis id="B96AEA88FF4FCDA6FEE9EC85B35CEC4F" box="[306,559,1264,1286]" italics="true" pageId="152" pageNumber="153">Acanthophorella irystoni</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4FCDA6FDEEEC84B3A1EC4F" bold="true" box="[565,722,1265,1286]" pageId="152" pageNumber="153">gen. et sp. nov.</emphasis>
, paratype male habitus. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF4FCDA6FF1CED5AB185EF57" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">
HEAD. Flattened in males. Labrum with three medial teeth and 5+5 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 6+6 setae, on each plate arranged in two rows. Stipites with 25+25 setae. Antennae
<quantity id="4CE69B7FFF4FCDA6FE38ED03B34FECC7" box="[483,572,1398,1422]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="152" pageNumber="153" unit="mm" value="1.4">1.4 mm</quantity>
long in
<typeStatus id="54A58838FF4FCDA6FD79ED00B27BECC7" box="[674,776,1397,1422]" pageId="152" pageNumber="153" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.09), II (0.15), III (0.34), IV (0.19), V (0.35), VI (0.17), VII (0.1) and VIII (0.01). Length/breadth ratios of antennomeres I–VII: I (1), II (1.5), III (4.2), IV (2.4), V (3.5), VI (1.7) and VII (1.3). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 13–16, arranged in 4–5 rows in males;
<quantity id="4CE69B7FFF4FCDA6FB74ED97B461ECB2" box="[1199,1298,1506,1531]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.318" metricValueMin="3.302" pageId="152" pageNumber="153" unit="in" value="15.0" valueMax="17.0" valueMin="13.0">13–17 in</quantity>
4–5 rows in females.
</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CEE5FB189EF2F" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CEE06B40EEFE7" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">
BODY SEGMENTS (
<figureCitation id="13252A1FFF4FCDA6FE7EEE07B37CEFC2" box="[421,527,1650,1675]" captionStart="FIGURE 126" captionStartId="152.[151,250,1264,1286]" captionTargetBox="[180,1404,337,1250]" captionTargetId="figure@152.[180,1404,337,1250]" captionTargetPageId="152" captionText="FIGURE 126. Acanthophorella irystoni gen. et sp. nov., paratype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272875/files/figure.png" pageId="152" pageNumber="153">Fig. 126</figureCitation>
). Lateral keels like lateral swellings. Macrochaetae long and trichoid. CIX (pleurotergite 15) ~ 0.9; MIX (pleurotergite 15) ~ 2; PIX (pleurotergite 15) ~ 0.5; MA (pleurotergite 15) ~ 106˚.
</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CEECEB39FEFBF" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CEF76B295EE77" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">WALKING LEGS. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.</paragraph>
<paragraph id="8BA1369AFF4FCDA6FF1CEF3EB054EEBA" blockId="152.[151,1437,1325,2035]" pageId="152" pageNumber="153">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF4FCDA6FDC2EF3FB30EEE2B" box="[537,637,1866,1891]" captionStart="FIGURE 127" captionStartId="153.[151,250,1965,1987]" captionTargetBox="[228,1345,193,1938]" captionTargetId="figure@153.[228,1345,193,1938]" captionTargetPageId="153" captionText="FIGURE 127. Acanthophorella irystoni gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 6. D. Leg 7. E. Leg 10. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272876/files/figure.png" pageId="152" pageNumber="153">Fig. 127</figureCitation>
). Leg-pairs 3–7 enlarged. Leg-pairs 3 and 4 each with a basal exterior protrusion on prefemur. Leg-pair 5 with a basal oral protrusion on prefemur. Leg-pair 6 with a small, basal, exterior protrusion on prefemur. Leg-pair 7 with a posterior, wide, squarish, thin, lamellar coxal process. Leg-pair 10 with coxal glands and with a short coxal horn orientated posteriorly. Leg-pair 11 with coxal glands, without other peculiarities.
</paragraph>
<caption id="DF616612FF4ECDA7FF4CEFD8B000EEAB" httpUri="https://zenodo.org/record/272876/files/figure.png" pageId="153" pageNumber="154" targetBox="[228,1345,193,1938]" targetPageId="153">
<paragraph id="8BA1369AFF4ECDA7FF4CEFD8B000EEAB" blockId="153.[151,1436,1965,2018]" pageId="153" pageNumber="154">
<emphasis id="B96AEA88FF4ECDA7FF4CEFD8B05EEE8B" bold="true" box="[151,301,1965,1987]" pageId="153" pageNumber="154">FIGURE 127.</emphasis>
<taxonomicName id="4C1E4D19FF4ECDA7FEEEEFD8B341EE8A" box="[309,562,1965,1987]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="irystoni">
<emphasis id="B96AEA88FF4ECDA7FEEEEFD8B341EE8A" box="[309,562,1965,1987]" italics="true" pageId="153" pageNumber="154">Acanthophorella irystoni</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4ECDA7FDE1EFD8B3A8EE8B" bold="true" box="[570,731,1965,1986]" pageId="153" pageNumber="154">gen. et sp. nov.</emphasis>
, holotype male.
<emphasis id="B96AEA88FF4ECDA7FC52EFD8B2D1EE8B" bold="true" box="[905,930,1965,1986]" pageId="153" pageNumber="154">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF4ECDA7FC28EFD8B579EE8B" bold="true" box="[1011,1034,1965,1986]" pageId="153" pageNumber="154">B.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF4ECDA7FB80EFD8B507EE8A" bold="true" box="[1115,1140,1965,1987]" pageId="153" pageNumber="154">C.</emphasis>
Leg 6.
<emphasis id="B96AEA88FF4ECDA7FB1EEFD8B5ADEE8B" bold="true" box="[1221,1246,1965,1986]" pageId="153" pageNumber="154">D.</emphasis>
Leg 7.
<emphasis id="B96AEA88FF4ECDA7FAF5EFD8B435EE8B" bold="true" box="[1326,1350,1965,1986]" pageId="153" pageNumber="154">E.</emphasis>
Leg 10. Scale lines = 0.2 mm.
</paragraph>
</caption>
<caption id="DF616612FF4DCDA4FF4CEDD8B4F8EF56" httpUri="https://zenodo.org/record/272877/files/figure.png" pageId="154" pageNumber="155" targetBox="[208,1381,224,1348]" targetPageId="154">
<paragraph id="8BA1369AFF4DCDA4FF4CEDD8B4F8EF56" blockId="154.[151,1436,1453,1567]" pageId="154" pageNumber="155">
<emphasis id="B96AEA88FF4DCDA4FF4CEDD8B05DEC8B" bold="true" box="[151,302,1453,1475]" pageId="154" pageNumber="155">FIGURE 128.</emphasis>
<taxonomicName id="4C1E4D19FF4DCDA4FEECEDD8B345EC8A" box="[311,566,1453,1475]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="irystoni">
<emphasis id="B96AEA88FF4DCDA4FEECEDD8B345EC8A" box="[311,566,1453,1475]" italics="true" pageId="154" pageNumber="155">Acanthophorella irystoni</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4DCDA4FDE5EDDBB391EC8A" bold="true" box="[574,738,1454,1475]" pageId="154" pageNumber="155">gen. et sp. nov.</emphasis>
, holotype male gonopods.
<emphasis id="B96AEA88FF4DCDA4FC25EDD8B564EC8B" bold="true" box="[1022,1047,1453,1474]" pageId="154" pageNumber="155">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF4DCDA4FA5FEDDBB4E8EC8A" bold="true" box="[1412,1435,1454,1475]" pageId="154" pageNumber="155">B.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF4DCDA4FE32EDB9B372ECAB" bold="true" box="[489,513,1484,1506]" pageId="154" pageNumber="155">C.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF4DCDA4FCADEDB9B2FCECA8" bold="true" box="[886,911,1484,1505]" pageId="154" pageNumber="155">D.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FF4DCDA4FAD2EDB8B467ECAB" box="[1289,1300,1485,1506]" italics="true" pageId="154" pageNumber="155">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF4DCDA4FAA4EDB8B4F5ECAB" box="[1407,1414,1485,1506]" italics="true" pageId="154" pageNumber="155">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF4DCDA4FED7ED9EB057EF49" box="[268,292,1515,1536]" italics="true" pageId="154" pageNumber="155">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FF4DCDA4FE3AED9FB09CEF49" box="[481,495,1514,1536]" italics="true" pageId="154" pageNumber="155">ll</emphasis>
= lateral lamella;
<emphasis id="B96AEA88FF4DCDA4FD68ED9EB3A1EF49" box="[691,722,1515,1536]" italics="true" pageId="154" pageNumber="155">mp</emphasis>
= medial shield;
<emphasis id="B96AEA88FF4DCDA4FC56ED9EB2D0EF49" box="[909,931,1515,1536]" italics="true" pageId="154" pageNumber="155">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF4DCDA4FB89ED9EB515EF49" box="[1106,1126,1515,1536]" italics="true" pageId="154" pageNumber="155">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF4DCDA4FADFED9EB46BEF49" box="[1284,1304,1515,1536]" italics="true" pageId="154" pageNumber="155">tp</emphasis>
= thorn-like process;
<emphasis id="B96AEA88FF4DCDA4FF2BEE7FB074EF56" box="[240,263,1546,1567]" italics="true" pageId="154" pageNumber="155">us</emphasis>
= undefined structure;
<emphasis id="B96AEA88FF4DCDA4FE28EE7FB365EF56" box="[499,534,1546,1567]" italics="true" pageId="154" pageNumber="155">cvp</emphasis>
= chitinised and membranous process;
<emphasis id="B96AEA88FF4DCDA4FC7CEE7FB2B9EF56" box="[935,970,1546,1567]" italics="true" pageId="154" pageNumber="155">ppr</emphasis>
= posterior projection. Scale line = 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF4DCDA4FF1CEE39B06CEE51" blockId="154.[151,1436,1612,1961]" pageId="154" pageNumber="155">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF4DCDA4FE01EE39B34CEF2D" box="[474,575,1612,1637]" captionStart="FIGURE 128" captionStartId="154.[151,250,1453,1475]" captionTargetBox="[208,1381,224,1348]" captionTargetId="figure@154.[188,1398,193,1432]" captionTargetPageId="154" captionText="FIGURE 128. Acanthophorella irystoni gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; ll = lateral lamella; mp = medial shield; sp = sternal plate; ss = sternal sac; tp = thorn-like process; us = undefined structure; cvp = chitinised and membranous process; ppr = posterior projection. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272877/files/figure.png" pageId="154" pageNumber="155">Fig. 128</figureCitation>
A–C). Sternal plate (
<emphasis id="B96AEA88FF4DCDA4FCE7EE38B226EF2D" box="[828,853,1613,1636]" italics="true" pageId="154" pageNumber="155">sp</emphasis>
) with a small, medial, hairless, sternal sac (
<emphasis id="B96AEA88FF4DCDA4FAADEE38B4FFEF2D" box="[1398,1420,1613,1636]" italics="true" pageId="154" pageNumber="155">ss</emphasis>
). Gonopods consisting of two medial shields (
<emphasis id="B96AEA88FF4DCDA4FD51EE04B3DFEFC1" box="[650,684,1649,1672]" italics="true" pageId="154" pageNumber="155">mp</emphasis>
), divided distally; apically, mesal edges folded inside and forming a posterior projection (
<emphasis id="B96AEA88FF4DCDA4FE47EEE0B0B1EFE5" box="[412,450,1685,1708]" italics="true" pageId="154" pageNumber="155">ppr</emphasis>
); lateral sides partly covering well-developed thorn-like processes (
<emphasis id="B96AEA88FF4DCDA4FB16EEE0B590EFE5" box="[1229,1251,1685,1708]" italics="true" pageId="154" pageNumber="155">tp</emphasis>
) that arise from the fused base of the gonopods. The lateralmost parts of anterior gonopods are lateral denticulate lamellae (
<emphasis id="B96AEA88FF4DCDA4FA58EECDB4E0EF98" box="[1411,1427,1720,1745]" italics="true" pageId="154" pageNumber="155">ll</emphasis>
) curved posteriorly and mesally. Lateral lamellae lower than thorn-like processes. Posteriorly a pair of lobes (
<emphasis id="B96AEA88FF4DCDA4FAA1EEA8B4E0EFBD" box="[1402,1427,1757,1780]" italics="true" pageId="154" pageNumber="155">us</emphasis>
) can be seen.
</paragraph>
<paragraph id="8BA1369AFF4DCDA4FF1CEF50B0BFEECD" blockId="154.[151,1436,1612,1961]" pageId="154" pageNumber="155">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF4DCDA4FE0DEF51B344EE75" box="[470,567,1828,1853]" captionStart="FIGURE 128" captionStartId="154.[151,250,1453,1475]" captionTargetBox="[208,1381,224,1348]" captionTargetId="figure@154.[188,1398,193,1432]" captionTargetPageId="154" captionText="FIGURE 128. Acanthophorella irystoni gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; ll = lateral lamella; mp = medial shield; sp = sternal plate; ss = sternal sac; tp = thorn-like process; us = undefined structure; cvp = chitinised and membranous process; ppr = posterior projection. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272877/files/figure.png" pageId="154" pageNumber="155">Fig. 128</figureCitation>
D). Coxites (
<emphasis id="B96AEA88FF4DCDA4FD13EF50B3A7EE75" box="[712,724,1829,1852]" italics="true" pageId="154" pageNumber="155">c</emphasis>
) divided. Telopodites (
<emphasis id="B96AEA88FF4DCDA4FC07EF50B297EE75" box="[988,996,1829,1852]" italics="true" pageId="154" pageNumber="155">t</emphasis>
) present on posterolateral sides. Coxal processes (
<emphasis id="B96AEA88FF4DCDA4FECFEF3CB05CEE29" box="[276,303,1865,1888]" italics="true" pageId="154" pageNumber="155">cp</emphasis>
) on posterior side, small. Anteriorly, high, partly chitinized and partly membranous processes (
<emphasis id="B96AEA88FF4DCDA4FAB7EF3CB4E1EE29" box="[1388,1426,1865,1888]" italics="true" pageId="154" pageNumber="155">cvp</emphasis>
) with coxal vesicles present.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF4DCDA4FF1CEFE5B551EEE0" box="[199,1058,1936,1961]" pageId="154" pageNumber="155" type="distribution">
<paragraph id="8BA1369AFF4DCDA4FF1CEFE5B551EEE0" blockId="154.[151,1436,1612,1961]" box="[199,1058,1936,1961]" pageId="154" pageNumber="155">
<emphasis id="B96AEA88FF4DCDA4FF1CEFE5B013EEE0" bold="true" box="[199,352,1936,1961]" pageId="154" pageNumber="155">Distribution.</emphasis>
<collectingCountry id="F309760AFF4DCDA4FEBCEFE5B0C1EEE0" box="[359,434,1936,1961]" name="Russia" pageId="154" pageNumber="155">Russia</collectingCountry>
(known only from
<typeStatus id="54A58838FF4DCDA4FD57EFE4B3CFEEE0" box="[652,700,1937,1961]" pageId="154" pageNumber="155">type</typeStatus>
locality) (
<figureCitation id="13252A1FFF4DCDA4FCF4EFE5B2E3EEE0" box="[815,912,1936,1961]" captionStart="FIGURE 169" captionStartId="197.[151,250,1981,2003]" captionTargetBox="[207,1379,1136,1959]" captionTargetId="figure@197.[207,1379,1136,1959]" captionTargetPageId="197" captionText="FIGURE 169. Distribution of the Caucasian Anthroleucosomatidae: the Flagellophorella complex." httpUri="https://zenodo.org/record/272918/files/figure.png" pageId="154" pageNumber="155">Fig. 169</figureCitation>
, blue circle).
</paragraph>
</subSubSection>
</treatment>
</document>
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