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<document id="7939C2CBF4688ADF99898925D33920C0" ID-DOI="10.11646/phytotaxa.298.3.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13694274" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725551073606" checkinUser="felipe" docAuthor="Jiménez-Mejías, Pedro &amp; Roalson, Eric H." docDate="2017" docId="AC4B18474876FB0008AC4BAD16FDD5AE" docLanguage="en" docName="phytotaxa.298.3.8.pdf" docOrigin="Phytotaxa 298 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.298.3.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Carex herbacoeli Jim. Mejias &amp; Roalson 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="287" masterDocId="5072603F4874FB0408244E7E147DD349" masterDocTitle="Two new Asian species of Carex (Cyperaceae)" masterLastPageNumber="288" masterPageNumber="283" pageNumber="285" updateTime="1725551450493" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="86F938F6FF9744265093FF5358CB0230">Two new Asian species of Carex (Cyperaceae)</mods:title>
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<mods:namePart id="739B0B9907E80656A04F0A3AA0517EA5">Jiménez-Mejías, Pedro</mods:namePart>
<mods:affiliation id="0911ACF4BC6CA9CEC46ABCAE1001B921">School of Biological Sciences, Washington State University, Pullman, WA 99164 - 4236, USA. &amp; Current address: New York Botanical Garden, 2900 Southern Blvd, Bronx, NY 10458, USA.</mods:affiliation>
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<mods:namePart id="07AEAE1F73F488489E8C8AE247AB604E">Roalson, Eric H.</mods:namePart>
<mods:affiliation id="6F6FF35FD9172E8A8415E0690D30CF32">School of Biological Sciences, Washington State University, Pullman, WA 99164 - 4236, USA.</mods:affiliation>
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<mods:title id="0A2EFC855BC68C36A8D5A1CBF8FF36EA">Phytotaxa</mods:title>
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<mods:date id="090C611512238F104CE6628FAE2E65E7">2017</mods:date>
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<mods:classification id="04253E386614B59EFC371E5009B6E5BD">journal article</mods:classification>
<mods:identifier id="5FC32A544B70BB30E2957DC520FED459" type="DOI">10.11646/phytotaxa.298.3.8</mods:identifier>
<mods:identifier id="318944C2C2FE67A3D66461E7B26BF678" type="ISSN">1179-3163</mods:identifier>
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<treatment id="AC4B18474876FB0008AC4BAD16FDD5AE" ID-DOI="http://doi.org/10.5281/zenodo.13694284" ID-Zenodo-Dep="13694284" LSID="urn:lsid:plazi:treatment:AC4B18474876FB0008AC4BAD16FDD5AE" httpUri="http://treatment.plazi.org/id/AC4B18474876FB0008AC4BAD16FDD5AE" lastPageId="4" lastPageNumber="287" pageId="2" pageNumber="285">
<subSubSection id="6CF8FADA4876FB0608AC4BAD16B5D6A7" box="[136,712,1491,1518]" pageId="2" pageNumber="285" type="nomenclature">
<paragraph id="245DA9514876FB0608AC4BAD16B5D6A7" blockId="2.[136,1455,1491,1769]" box="[136,712,1491,1518]" pageId="2" pageNumber="285">
<heading id="7F151E3D4876FB0608AC4BAD16B5D6A7" bold="true" box="[136,712,1491,1518]" fontSize="11" level="1" pageId="2" pageNumber="285" reason="1">
<emphasis id="169675434876FB0608AC4BAD16B5D6A7" bold="true" box="[136,712,1491,1518]" pageId="2" pageNumber="285">
<taxonomicName id="E3E2D2D24876FB0608AC4BAD161FD6A7" ID-CoL="8WS6X" authority="Jim.Mejias &amp; Roalson" authorityName="Jim. Mejias &amp; Roalson" authorityYear="2017" box="[136,610,1491,1518]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="herbacoeli" status="sp. nov.">
<emphasis id="169675434876FB0608AC4BAD1533D6A7" bold="true" box="[136,334,1491,1518]" italics="true" pageId="2" pageNumber="285">Carex herbacoeli</emphasis>
Jim.Mejías &amp; Roalson
</taxonomicName>
,
<taxonomicNameLabel id="0DA5C8384876FB060A4A4BAB16B5D6A7" box="[622,712,1493,1518]" pageId="2" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6CF8FADA4876FB0008AC4B8215B0D57A" lastPageId="4" lastPageNumber="287" pageId="2" pageNumber="285" type="diagnosis">
<paragraph id="245DA9514876FB0608AC4B821075D5E8" blockId="2.[136,1455,1491,1769]" pageId="2" pageNumber="285">
<emphasis id="169675434876FB0608AC4B821496D558" bold="true" box="[136,235,1532,1553]" pageId="2" pageNumber="285">Diagnosis</emphasis>
:—This distinctive species superficially resembles the co-occurring
<taxonomicName id="E3E2D2D24876FB060BA64B8210B7D558" authority="Koyama (1954: 112)" authorityName="Koyama" authorityPageNumber="112" authorityYear="1954" box="[898,1226,1532,1553]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="nakaoana">
<emphasis id="169675434876FB060BA64B821783D558" box="[898,1022,1532,1553]" italics="true" pageId="2" pageNumber="285">C. nakaoana</emphasis>
<bibRefCitation id="4073D4A04876FB060C214B8210B7D558" author="Koyama, T." box="[1029,1226,1532,1553]" pageId="2" pageNumber="285" pagination="111 - 114" refId="ref2584" refString="Koyama, T. (1954) New carices from Nepal collected by Sasuke Nakao. Acta Phytotaxonica et Geobotanica 15: 111 - 114." type="journal article" year="1954">Koyama (1954:112)</bibRefCitation>
</taxonomicName>
, and also
<taxonomicName id="E3E2D2D24876FB060D0A4B821508D57C" authority="Schkuhr (1801: 91)" authorityName="Schkuhr" authorityPageNumber="91" authorityYear="1801" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="fuliginosa">
<emphasis id="169675434876FB060D0A4B8211D6D558" box="[1326,1451,1532,1553]" italics="true" pageId="2" pageNumber="285">C. fuliginosa</emphasis>
<bibRefCitation id="4073D4A04876FB060899485E1508D57C" author="Schkuhr, C." box="[189,373,1568,1589]" pageId="2" pageNumber="285" refId="ref2715" refString="Schkuhr, C. (1801) Beschreibung und Abbildung der Theils bekannten, 1. Der Verfasserm, Wittenberg." type="book" year="1801">Schkuhr (1801:91)</bibRefCitation>
</taxonomicName>
, which is absent from the Himalayas. From
<taxonomicName id="E3E2D2D24876FB060B01485E17DCD57C" authorityName="Koyama" authorityYear="1954" box="[805,929,1568,1589]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="nakaoana">
<emphasis id="169675434876FB060B01485E17DCD57C" box="[805,929,1568,1589]" italics="true" pageId="2" pageNumber="285">C. nakaoana</emphasis>
</taxonomicName>
it differs by having 2 stigmas instead of 3, by having the terminal spike gynecandrous rather than all male as in
<taxonomicName id="E3E2D2D24876FB060ACB483A1717D510" authorityName="Koyama" authorityYear="1954" box="[751,874,1604,1625]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="nakaoana">
<emphasis id="169675434876FB060ACB483A1717D510" box="[751,874,1604,1625]" italics="true" pageId="2" pageNumber="285">C. nakaoana</emphasis>
</taxonomicName>
, and because the utricles are progressively attenuated into a linear beak instead a deltoid one as in
<taxonomicName id="E3E2D2D24876FB060A094816172DD534" authority="(Dai et al. 2010)" baseAuthorityName="Dai" baseAuthorityYear="2010" box="[557,848,1640,1661]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="nakaoana">
<emphasis id="169675434876FB060A09481616D5D534" box="[557,680,1640,1661]" italics="true" pageId="2" pageNumber="285">C. nakaoana</emphasis>
(
<bibRefCitation id="4073D4A04876FB060A9148161734D534" author="Dai, L. K. &amp; Liang, S. Y. &amp; Zhang, S. R. &amp; Tang, Y. C. &amp; Koyama, T. &amp; Tucker, G. C." box="[693,841,1640,1661]" pageId="2" pageNumber="285" pagination="285 - 461" refId="ref2128" refString="Dai, L. K., Liang, S. Y., Zhang, S. R., Tang, Y. C., Koyama, T. &amp; Tucker, G. C. (2010) Carex Linnaeus. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, 23. Beijing and Saint Louis, Science Press and Missouri Botanical Garden Press, pp. 285 - 461." type="book chapter" year="2010">
Dai
<emphasis id="169675434876FB060AFB4816176DD534" box="[735,784,1640,1661]" italics="true" pageId="2" pageNumber="285">et al.</emphasis>
2010
</bibRefCitation>
)
</taxonomicName>
. From
<taxonomicName id="E3E2D2D24876FB060BB04816106DD534" authorityName="Schkuhr" authorityYear="1801" box="[916,1040,1640,1661]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="fuliginosa">
<emphasis id="169675434876FB060BB04816106DD534" box="[916,1040,1640,1661]" italics="true" pageId="2" pageNumber="285">C. fuliginosa</emphasis>
</taxonomicName>
it differs by having 2 stigmas instead of 3, and the rhizomes long instead cespitose, (
<bibRefCitation id="4073D4A04876FB060A6848F216BAD5E8" author="Chater, A. O." box="[588,711,1676,1697]" pageId="2" pageNumber="285" pagination="290 - 323" refId="ref1997" refString="Chater, A. O. (1980) Carex L. In: Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Walters, S. M. &amp; Webb, D. A. (Eds.) Flora Europaea 5. Cambridge University Press, Cambridge, pp. 290 - 323." type="book chapter" year="1980">Chater 1980</bibRefCitation>
;
<bibRefCitation id="4073D4A04876FB060AF548F21726D5E8" author="Egorova, T. V." box="[721,859,1676,1697]" pageId="2" pageNumber="285" refId="ref2262" refString="Egorova, T. V. (1999) The Sedges (Carex L.) of Russia and Adjacent States. Missouri Botanical Garden Press, Saint-Louis." type="book" year="1999">Egorova 1999</bibRefCitation>
;
<bibRefCitation id="4073D4A04876FB060B4148F21780D5E8" author="Ball, P. W. &amp; Reznicek, A. A." box="[869,1021,1676,1697]" pageId="2" pageNumber="285" pagination="254 - 273" refId="ref1925" refString="Ball, P. W. &amp; Reznicek, A. A. (2002) Carex Linnaeus. In: Flora of North America Editorial Committee (Eds.) Flora of North America, North of Mexico, 23. Magnoliophyta: Commelinidae (in part): Cyperaceae Oxford University Press, New York, NY, and Oxford, UK., pp. 254 - 273." type="book chapter" year="2002">
Ball
<emphasis id="169675434876FB060BB748F217C0D5E8" box="[915,957,1676,1697]" italics="true" pageId="2" pageNumber="285">et al</emphasis>
. 2002
</bibRefCitation>
).
</paragraph>
<paragraph id="245DA9514876FB0608AC48CE1770D5A0" blockId="2.[136,1455,1491,1769]" pageId="2" pageNumber="285">
<emphasis id="169675434876FB0608AC48CE14C8D58C" box="[136,181,1712,1733]" italics="true" pageId="2" pageNumber="285">Type</emphasis>
:—
<materialsCitation id="948AA30C4876FB0608F548CE1774D5A0" collectingDate="1994-08-03" collectionCode="F, A" country="Nepal" county="Rasuwa Distr." elevation="4300" latitude="28.216667" location="Krharka" longLatPrecision="1233" longitude="85.2" municipality="Paldol Base Camp" pageId="2" pageNumber="285" specimenCount="2" stateProvince="Bagmati" typeStatus="holotype">
<collectingCountry id="5CF5E9C14876FB0608F548CE155FD58C" box="[209,290,1712,1733]" name="Nepal" pageId="2" pageNumber="285">NEPAL</collectingCountry>
.
<collectingRegion id="E62667B34876FB06090848CE15C9D58C" box="[300,436,1712,1733]" country="Nepal" name="Bagmati" pageId="2" pageNumber="285">Bagmati zone</collectingRegion>
,
<collectingCounty id="CD3CD1DD4876FB06099A48CE1634D58C" box="[446,585,1712,1733]" pageId="2" pageNumber="285">Rasuwa Distr.</collectingCounty>
,
<collectingMunicipality id="C439332B4876FB060A7748CE1778D58C" box="[595,773,1712,1733]" pageId="2" pageNumber="285">Paldol Base Camp</collectingMunicipality>
(
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<elevation id="AFCF4E624876FB060B3748CE1722D58C" box="[787,863,1712,1733]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="285" unit="m" value="4300.0">4300 m</elevation>
</quantity>
)—a
<location id="213DFF8A4876FB060BB448CE17A2D58C" LSID="urn:lsid:plazi:treatment:AC4B18474876FB0008AC4BAD16FDD5AE:213DFF8A4876FB060BB448CE17A2D58C" box="[912,991,1712,1733]" country="Nepal" county="Rasuwa Distr." latitude="28.216667" longLatPrecision="1233" longitude="85.2" municipality="Paldol Base Camp" name="Krharka" pageId="2" pageNumber="285" stateProvince="Bagmati">Krharka</location>
(
<quantity id="E31A04B44876FB060BC848CE1047D58C" box="[1004,1082,1712,1733]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="2" pageNumber="285" unit="m" value="4050.0">
<elevation id="AFCF4E624876FB060BC848CE1047D58C" box="[1004,1082,1712,1733]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="2" pageNumber="285" unit="m" value="4050.0">4050 m</elevation>
</quantity>
).
<geoCoordinate id="41D6CF964876FB060C6848CE10D7D58C" box="[1100,1194,1712,1733]" degrees="28" direction="north" minutes="13" orientation="latitude" pageId="2" pageNumber="285" precision="925" value="28.216667">28° 13’ N</geoCoordinate>
<geoCoordinate id="41D6CF964876FB060C9648CE116DD58C" box="[1202,1296,1712,1733]" degrees="85" direction="east" minutes="12" orientation="longitude" pageId="2" pageNumber="285" precision="925" value="85.2">85° 12’ E</geoCoordinate>
.
<date id="505C8F914876FB060D3F48CE11EDD58C" box="[1307,1424,1712,1733]" pageId="2" pageNumber="285" value="1994-08-03">
<collectingDate id="401876794876FB060D3F48CE11EDD58C" box="[1307,1424,1712,1733]" pageId="2" pageNumber="285" value="1994-08-03">3 Aug 1994</collectingDate>
</date>
.
<emphasis id="169675434876FB060DBE48CE15EAD5A0" italics="true" pageId="2" pageNumber="285">
<collectionCode id="42F331944876FB060DBE48CE11D1D58C" box="[1434,1452,1712,1733]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="2" pageNumber="285" type="Herbarium">F</collectionCode>
. Miyamoto et al. 10160
</emphasis>
(
<typeStatus id="FB5917F34876FB06098748AA167CD5A0" box="[419,513,1748,1769]" pageId="2" pageNumber="285" type="holotype">Holotype</typeStatus>
:
<collectionCode id="42F331944876FB060A2D48AA165CD5A0" box="[521,545,1748,1769]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="285" type="Herbarium">A</collectionCode>
!;
<typeStatus id="FB5917F34876FB060A0B48AA1605D5A0" box="[559,632,1748,1769]" pageId="2" pageNumber="285" type="isotype">isotype</typeStatus>
:
<collectionCode id="42F331944876FB060AA448AA16EAD5A0" box="[640,663,1748,1769]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="285" type="Herbarium">A</collectionCode>
!).
<figureCitation id="BCD9B5D44876FB060A8948AA1774D5A0" box="[685,777,1748,1769]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[136,229,1953,1974]" captionStartId-1="4.[136,229,1169,1190]" captionTargetBox-0="[193,1395,190,1929]" captionTargetBox-1="[242,1336,207,1115]" captionTargetId-0="figure-14@3.[193,1395,190,1929]" captionTargetId-1="figure-18@4.[238,1349,190,1144]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 3. Holotype of Carex herbacoeli Jim.-Mejías &amp; Roalson." captionText-1="FIGURE 4. Detailed picture of the holotype of Carex herbacoeli Jim.-Mejías &amp; Roalson. A. Inflorescence. B-C. Glumes. D–E. Utricles. Scale bars represent 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13694282" figureDoi-1="http://doi.org/10.5281/zenodo.13694290" httpUri-0="https://zenodo.org/record/13694282/files/figure.png" httpUri-1="https://zenodo.org/record/13694290/files/figure.png" pageId="2" pageNumber="285">Figs. 3–4</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="245DA9514876FB0008AC496715B0D57A" blockId="2.[136,1452,1817,2058]" lastBlockId="4.[136,1452,1275,1767]" lastPageId="4" lastPageNumber="287" pageId="2" pageNumber="285">
Perennial, with long rhizomes. Stems
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long,
<quantity id="E31A04B44876FB060B2449671724D47B" box="[768,857,1817,1842]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="285" unit="mm" value="0.6">0.6 mm</quantity>
wide below the inflorescence, obscurely trigonous, almost cylindrical, sulcate, smooth. Leaves basal and cauline, shorter than stems, the uppermost ones up to
<quantity id="E31A04B44876FB060D7B494314D7D433" metricMagnitude="-1" metricUnit="m" metricValue="1.125" metricValueMax="1.5" metricValueMin="0.75" pageId="2" pageNumber="285" unit="cm" value="11.25" valueMax="15.0" valueMin="7.5">7.5–15 cm</quantity>
long, the widest ones
<quantity id="E31A04B44876FB06098B491F1645D433" box="[431,568,1889,1914]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="4.3" metricValueMin="2.6" pageId="2" pageNumber="285" unit="mm" value="3.45" valueMax="4.3" valueMin="2.6">2.6–4.3 mm</quantity>
wide, flat or slightly incurved outwards, apparently hypostomatic, antrorsely scabrid on edges; ligule of cauline leaves
<quantity id="E31A04B44876FB060A7349FB16B9D4D7" box="[599,708,1925,1950]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="2" pageNumber="285" unit="mm" value="7.0" valueMax="12.0" valueMin="2.0">2–12 mm</quantity>
, pale orangey to yellowish or whitish, acute; basal sheaths creamy, sometimes red-brownish tinged, bladeless, splitting in fibers. Inflorescence
<quantity id="E31A04B44876FB060BC349D71030D48B" box="[999,1101,1961,1986]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="2" pageNumber="285" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 cm</quantity>
, with 3–5 (often 4) spikes, the uppermost gynecandrous, the other female, loosely disposed at the top of the stems; lowermost bract shortly leafy to setaceous, shorter than the inflorescence
<quantity id="E31A04B44876FB060A7B498F16A3DB43" box="[607,734,2033,2058]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="5.5" metricValueMin="1.3" pageId="2" pageNumber="285" unit="cm" value="3.4" valueMax="5.5" valueMin="1.3">1.3–5.5 cm</quantity>
×
<quantity id="E31A04B44876FB060B25498F1705DB43" box="[769,888,2033,2058]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="285" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
, flattish, with a sheath (0.5)
<quantity id="E31A04B44876FB060C99498F116CDB43" box="[1213,1297,2033,2058]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="285" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
long; second lowermost bract setaceous, shorter than its spike, sheathless. Uppermost spike 17–25 ×
<quantity id="E31A04B44870FB000C454A8510BDD65D" box="[1121,1216,1275,1300]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="4" pageNumber="287" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
, subclavate, with the male and female parts subequal in length, with a peduncle
<quantity id="E31A04B44870FB000B024B6117F9D671" box="[806,900,1311,1336]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="4" pageNumber="287" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">2–8 mm</quantity>
long. Lateral spikes 12–33 ×
<quantity id="E31A04B44870FB000CFD4B611147D671" box="[1241,1338,1311,1336]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="287" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, narrowly obovate, dense, the lowermost with a peduncle
<quantity id="E31A04B44870FB000AB74B3D1687D615" box="[659,762,1347,1372]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="4" pageNumber="287" unit="cm" value="2.6" valueMax="3.0" valueMin="2.2">2.2–3 cm</quantity>
long, flexuous or suberect, utricles ascending, 15–45. Glumes 3.5–4.5 ×
<quantity id="E31A04B44870FB0008DA4B191525D6C9" box="[254,344,1383,1408]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="287" unit="mm" value="1.2">1.2 mm</quantity>
, obovate to narrowly obovate, dark purplish with a greenish midrib, obtuse, scabrid at the tip on the margins and midrib. Stigmas 2. Utricles 4.5–5 ×
<quantity id="E31A04B44870FB000AFB4BF41714D6EA" box="[735,873,1418,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="4" pageNumber="287" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
, elliptical, flattish, with only the two lateral nerves prominent, sparsely hairy, yellowish, progressively becoming purplish on the abaxial side from the upper half or the upper third up to the beak, but keeping lateral nerves always yellowish, gradually attenuated into a beak
<quantity id="E31A04B44870FB000D014BAC11D1D6A2" box="[1317,1452,1490,1515]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="4" pageNumber="287" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
long, linear, flattish, scabrid, bidentate. No ripe achenes found, the inmature achenes studied conspicuously stipitate, with a stipe ca.
<quantity id="E31A04B44870FB000913486415F0D57A" box="[311,397,1562,1587]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="287" unit="mm" value="0.5">0.5 mm</quantity>
long.
</paragraph>
</subSubSection>
<caption id="709DF9D94877FB0708AC49DF176DD4FF" ID-DOI="http://doi.org/10.5281/zenodo.13694282" ID-Zenodo-Dep="13694282" box="[136,784,1953,1974]" httpUri="https://zenodo.org/record/13694282/files/figure.png" pageId="3" pageNumber="286" startId="3.[136,229,1953,1974]" targetBox="[193,1395,190,1929]" targetPageId="3" targetType="figure">
<paragraph id="245DA9514877FB0708AC49DF176DD4FF" blockId="3.[136,784,1953,1974]" box="[136,784,1953,1974]" pageId="3" pageNumber="286">
<emphasis id="169675434877FB0708AC49DF1483D4FF" bold="true" box="[136,254,1953,1974]" pageId="3" pageNumber="286">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="E3E2D2D24877FB07095949DF162CD4FF" authority="Jim." authorityName="Jim. Mejias &amp; Roalson" authorityYear="2017" box="[381,593,1953,1974]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="3" pageNumber="286" phylum="Tracheophyta" rank="species" species="herbacoeli">
<emphasis id="169675434877FB07095949DF165ED4FF" box="[381,547,1953,1974]" italics="true" pageId="3" pageNumber="286">Carex herbacoeli</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; Roalson.
</paragraph>
</caption>
<caption id="709DF9D94870FB0008AC4AEF15F1D783" ID-DOI="http://doi.org/10.5281/zenodo.13694290" ID-Zenodo-Dep="13694290" httpUri="https://zenodo.org/record/13694290/files/figure.png" pageId="4" pageNumber="287" startId="4.[136,229,1169,1190]" targetBox="[242,1336,207,1115]" targetPageId="4" targetType="figure">
<paragraph id="245DA9514870FB0008AC4AEF15F1D783" blockId="4.[136,1453,1169,1226]" pageId="4" pageNumber="287">
<emphasis id="169675434870FB0008AC4AEF1480D7EF" bold="true" box="[136,253,1169,1190]" pageId="4" pageNumber="287">FIGURE 4.</emphasis>
Detailed picture of the holotype of
<taxonomicName id="E3E2D2D24870FB000A774AEF1755D7EF" authority="Jim." authorityName="Jim. Mejias &amp; Roalson" authorityYear="2017" box="[595,808,1169,1190]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="287" phylum="Tracheophyta" rank="species" species="herbacoeli">
<emphasis id="169675434870FB000A774AEF1687D7EF" box="[595,762,1169,1190]" italics="true" pageId="4" pageNumber="287">Carex herbacoeli</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; Roalson.
<emphasis id="169675434870FB000BCE4AEF107FD7EF" bold="true" box="[1002,1026,1169,1190]" pageId="4" pageNumber="287">A.</emphasis>
Inflorescence.
<emphasis id="169675434870FB000CB14AEF10BED7EF" bold="true" box="[1173,1219,1169,1190]" pageId="4" pageNumber="287">B-C.</emphasis>
Glumes.
<emphasis id="169675434870FB000D044AEF112ED7EF" bold="true" box="[1312,1363,1169,1190]" pageId="4" pageNumber="287">D–E.</emphasis>
Utricles. Scale bars represent 1 mm.
</paragraph>
</caption>
<subSubSection id="6CF8FADA4870FB000899484016FDD5AE" pageId="4" pageNumber="287" type="etymology">
<paragraph id="245DA9514870FB000899484016B9D532" blockId="4.[136,1452,1275,1767]" pageId="4" pageNumber="287">
<emphasis id="169675434870FB00089948401543D51E" bold="true" box="[189,318,1598,1623]" pageId="4" pageNumber="287">Etymology</emphasis>
:—From the Latin “
<emphasis id="169675434870FB000A3848401620D51E" box="[540,605,1598,1623]" italics="true" pageId="4" pageNumber="287">herba</emphasis>
”, herb or grass, and “
<emphasis id="169675434870FB000B74484017E5D51E" box="[848,920,1598,1623]" italics="true" pageId="4" pageNumber="287">coelus</emphasis>
” (genitive
<emphasis id="169675434870FB000C3148401036D51E" box="[1045,1099,1598,1623]" italics="true" pageId="4" pageNumber="287">coeli</emphasis>
), sky or heaven, in reference to the extreme elevations this species has been found.
</paragraph>
<paragraph id="245DA9514870FB00089948F916FDD5AE" blockId="4.[136,1452,1275,1767]" pageId="4" pageNumber="287">
<emphasis id="169675434870FB00089948F91521D5E9" bold="true" box="[189,348,1671,1696]" pageId="4" pageNumber="287">Observations</emphasis>
:—This new species is unique among the Asian species grouped under section
<taxonomicName id="E3E2D2D24870FB000C9A48F81579D58A" authority="Dumortier (1827: 147)" authorityName="Dumortier" authorityPageNumber="147" authorityYear="1827" class="Hexactinellida" family="Aulocystidae" genus="Aulocystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lychniscosida" pageId="4" pageNumber="287" phylum="Porifera" rank="genus">
<emphasis id="169675434870FB000C9A48F8114CD5D6" box="[1214,1329,1670,1695]" italics="true" pageId="4" pageNumber="287">Aulocystis</emphasis>
Dumortier (1827:147)
</taxonomicName>
by the combination of gynecandrous spikes, 2 stigmas, and utricles hairy, attenuated into a linear beak (see
<bibRefCitation id="4073D4A04870FB0008AC48B0156DD5AE" author="Noltie, H. J." box="[136,272,1742,1767]" pageId="4" pageNumber="287" refId="ref2642" refString="Noltie, H. J. (1994) Flora of Bhutan. Royal Botanic Garden of Edinburgh, Edinburgh." type="book" year="1994">Noltie 1994</bibRefCitation>
;
<bibRefCitation id="4073D4A04870FB00093848B015C0D5AE" author="Egorova, T. V." box="[284,445,1742,1767]" pageId="4" pageNumber="287" refId="ref2262" refString="Egorova, T. V. (1999) The Sedges (Carex L.) of Russia and Adjacent States. Missouri Botanical Garden Press, Saint-Louis." type="book" year="1999">Egorova 1999</bibRefCitation>
;
<bibRefCitation id="4073D4A04870FB0009ED48B01609D5AE" author="Dai, L. K. &amp; Liang, S. Y. &amp; Zhang, S. R. &amp; Tang, Y. C. &amp; Koyama, T. &amp; Tucker, G. C." box="[457,628,1742,1767]" pageId="4" pageNumber="287" pagination="285 - 461" refId="ref2128" refString="Dai, L. K., Liang, S. Y., Zhang, S. R., Tang, Y. C., Koyama, T. &amp; Tucker, G. C. (2010) Carex Linnaeus. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, 23. Beijing and Saint Louis, Science Press and Missouri Botanical Garden Press, pp. 285 - 461." type="book chapter" year="2010">
Dai
<emphasis id="169675434870FB0009DC48B0164DD5AE" box="[504,560,1742,1767]" italics="true" pageId="4" pageNumber="287">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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