treatments-xml/data/B4/A0/1B/B4A01B60087A5DF48D79FBA0DBC37007.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.945.51860" ID-GBIF-Dataset="1c694c7e-3d35-4520-b398-6189a61d1430" ID-PMC="PMC7351832" ID-Pensoft-Pub="1313-2970-945-99" ID-Pensoft-UUID="1E1F40C8FAD751509E70897F4DE56527" ID-PubMed="32714007" ID-ZooBank="CCD1B83E262441CDBD0AE1C57EDCA4DC" ModsDocID="1313-2970-945-99" checkinTime="1593797374489" checkinUser="pensoft" docAuthor="Potapov, Mikhail B., Janion-Scheepers, Charlene &amp; Deharveng, Louis" docDate="2020" docId="B4A01B60087A5DF48D79FBA0DBC37007" docLanguage="en" docName="ZooKeys 945: 99-127" docOrigin="ZooKeys 945" docSource="http://dx.doi.org/10.3897/zookeys.945.51860" docTitle="Cryptopygus abulbus Potapov &amp; Janion-Scheepers &amp; Deharveng 2020, sp. nov." docType="treatment" docUuid="EE9161B6-9521-407C-ADA5-1742B864573E" docUuidSource="ZooBank" docVersion="4" id="1E1F40C8FAD751509E70897F4DE56527" lastPageNumber="99" masterDocId="1E1F40C8FAD751509E70897F4DE56527" masterDocTitle="Taxonomy of the Cryptopygus complex. III. The revision of South African species of Cryptopygus and Isotominella (Collembola, Isotomidae)" masterLastPageNumber="127" masterPageNumber="99" pageNumber="99" updateTime="1668169057397" updateUser="ExternalLinkService">
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<mods:title>Taxonomy of the Cryptopygus complex. III. The revision of South African species of Cryptopygus and Isotominella (Collembola, Isotomidae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Potapov, Mikhail B.</mods:namePart>
<mods:affiliation>Senckenberg Museum of Natural History Goerlitz, Am Museum 1, 02826, Goerlitz, Germany &amp; Moscow State Pedagogical University, Kibalchich str., 6, korp. 3, Moscow 129278, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6111-3354</mods:nameIdentifier>
<mods:nameIdentifier type="email">mpnk-abroad@yandex.ru</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Janion-Scheepers, Charlene</mods:namePart>
<mods:affiliation>University of Cape Town, Department of Biological Sciences, Rondebosch, 7701, South Africa &amp; Iziko Museums of South Africa, 25 Queen Victoria Street, Cape Town, 8001, South Africa</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5942-7912</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Deharveng, Louis</mods:namePart>
<mods:affiliation>Institut de Systematique, Evolution, Biodiversite ISYEB-UMR 7205 - CNRS, MNHN, UPMC, EPHE, Museum national d'Histoire naturelle, Sorbonne Universites, 45 rue Buffon, F- 75005, Paris, France</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>945</mods:number>
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<mods:start>99</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.945.51860</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.945.51860</mods:identifier>
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<treatment ID-GBIF-Taxon="164910869" LSID="urn:lsid:zoobank.org:act:EE9161B6-9521-407C-ADA5-1742B864573E" httpUri="http://treatment.plazi.org/id/B4A01B60087A5DF48D79FBA0DBC37007" lastPageNumber="99" pageId="0" pageNumber="99">
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<paragraph pageId="0" pageNumber="99">
<taxonomicName LSID="B4A01B60-087A-5DF4-8D79-FBA0DBC37007" authority="Potapov &amp; Janion-Scheepers &amp; Deharveng, 2020" authorityName="Potapov &amp; Janion-Scheepers &amp; Deharveng" authorityYear="2020" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus abulbus" order="Entomobryomorpha" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="abulbus" status="sp. nov.">Cryptopygus abulbus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="99">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Appearance and macrochaetotaxy of C. bulbus sp. nov., ventral (1) and lateral (2) views, and C. abulbus sp. nov. (3)." figureDoi="10.3897/zookeys.945.51860.figures1-3" httpUri="https://binary.pensoft.net/fig/428159" pageId="0" pageNumber="99">Figures 3</figureCitation>
<figureCitation captionStart="Figures 4–5" captionStartId="F2" captionText="Figures 4 - 5. Macrochaetotaxy and s-ms-pattern of C. bulbus sp. nov. (4) and C. abulbus sp. nov. (5). Abbreviations: msms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures4-5" httpUri="https://binary.pensoft.net/fig/428160" pageId="0" pageNumber="99">, 5</figureCitation>
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">, 15-22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="type material">
<paragraph pageId="0" pageNumber="99">Type material.</paragraph>
<paragraph pageId="0" pageNumber="99">
Holotype and eighteen paratypes: South Africa • Western Cape, Stellenbosch, Jonkershoek Nature Reserve; 
<geoCoordinate degrees="33.986883" direction="south" orientation="latitude" precision="1" value="-33.98688">33.986883S</geoCoordinate>
, 
<geoCoordinate degrees="18.955350" direction="east" orientation="longitude" precision="1" value="18.95535">18.955350E</geoCoordinate>
; 30 July 2009; C. Janion-Scheepers leg.; litter trap (J2_32), Holotype and eight paratypes deposited on four slides at SAMC, four paratypes on two slides deposited at SMNG, four paratypes on two slides deposited at MSPU, two paratypes on one slide at MNHN.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="diagnosis">
<paragraph pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="99">Without globular retractile bulb on Ant. IV. Organite on Ant. IV chili-like. 6+6 ocelli. Maxillary palp simple. Two sublobal hairs. Anterior side of manubrium without chaetae. Tenaculum with two chaetae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
Body size 0.6-0.7 mm. Body with regular blue pigmentation, slender (Fig. 
<figureCitation captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Appearance and macrochaetotaxy of C. bulbus sp. nov., ventral (1) and lateral (2) views, and C. abulbus sp. nov. (3)." figureDoi="10.3897/zookeys.945.51860.figures1-3" httpUri="https://binary.pensoft.net/fig/428159" pageId="0" pageNumber="99">3</figureCitation>
). Abd. V well separated from Abd. IV and fused with Abd. VI (Fig. 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">15</figureCitation>
). Cuticle with orthogonal granulation. Ocelli 6+6 arranged as three in anterior and three in posterior group (Fig. 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">19</figureCitation>
). PAO more than twice as long as ocellus, 0.6-0.7 as long as width of Ant. I and 0.8-1.1 mm as long as inner unguis length. Maxillary outer lobe with two sublobal hairs, one individual with one sublobal hair on one side was found. Maxillary palp simple. Labral formula as 2/5,5,4. Labium with five usual papillae (
<emphasis italics="true" pageId="0" pageNumber="99">A-E</emphasis>
, Fig. 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">16</figureCitation>
) and labial formula as in 
<taxonomicName lsidName="C. bulbus" pageId="0" pageNumber="99" rank="species" species="bulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. bulbus</emphasis>
</taxonomicName>
sp. nov. Ventral side of head with 4+4 postlabial chaetae. Ant. I with eleven common chaetae, two ventral s-chaetae (s) and three small basal micro s-chaetae (bms), two dorsal and one ventral, Ant. II with three bms and one latero-distal s, Ant. III with one bms and five distal s (including one lateral). S-chaetae on Ant. IV weakly differentiated. Organite long, of chili-like shape, set apart from subapical micro s-chaeta (Figs 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">17</figureCitation>
, 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">18</figureCitation>
). Tip of antennae without retractile bulb.
</paragraph>
<caption doi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99" start="Figures 15–22" startId="F4">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figures 15-22.</emphasis>
<taxonomicName lsidName="C. abulbus" pageId="0" pageNumber="99" rank="species" species="abulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. abulbus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="99">15</emphasis>
Abd. IV and fused Abd. V and V 
<emphasis bold="true" pageId="0" pageNumber="99">16</emphasis>
labial palp 
<emphasis bold="true" pageId="0" pageNumber="99">17, 18</emphasis>
apical part of Ant. IV, different views 
<emphasis bold="true" pageId="0" pageNumber="99">19</emphasis>
ocelli and PAO 
<emphasis bold="true" pageId="0" pageNumber="99">20</emphasis>
ventral tube, ventral view 
<emphasis bold="true" pageId="0" pageNumber="99">21</emphasis>
furca, lateral view 
<emphasis bold="true" pageId="0" pageNumber="99">22</emphasis>
furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="99">
Common chaetae slightly (under very high magnification) serrated at the posterior part of abdomen. S-formula as 4,3/2,2,2,3,5 (s), 1,0/1,0,0 (ms) (Figs 
<figureCitation captionStart="Figures 4–5" captionStartId="F2" captionText="Figures 4 - 5. Macrochaetotaxy and s-ms-pattern of C. bulbus sp. nov. (4) and C. abulbus sp. nov. (5). Abbreviations: msms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures4-5" httpUri="https://binary.pensoft.net/fig/428160" pageId="0" pageNumber="99">5</figureCitation>
, 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">15</figureCitation>
). Tergal s-chaetae much shorter than common chaetae and well distinguishable. Medial s-chaetae on Th. II-Abd. III situated in mid-tergal position. Macrochaetae smooth and short, 1,1/3,3,3 in number, medial ones on Abd. VI 1.6-2.0 times longer than dens and 2.8-4.1 times longer than mucro. Foil chaetae at the tip of abdomen absent. Axial chaetotaxy as 6-8,5/3,3,3,5-6. Th. I and II without chaetae, Th. III with 2+2 ventral chaetae.
</paragraph>
<paragraph pageId="0" pageNumber="99">
Unguis without teeth. Empodial appendage 0.5-0.7 as in 
<taxonomicName lsidName="C. bulbus" pageId="0" pageNumber="99" rank="species" species="bulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. bulbus</emphasis>
</taxonomicName>
sp. nov. Tibiotarsi without additional chaetae on Leg I and II (21 chaetae), and with a few additional ones on Leg III, about 26 chaetae. Tibiotarsal tenent hairs clavate, 1,2,2 on Tibiotarsi 1,2,3. Ventral tube with 3+3 laterodistal and four posterior chaetae, anteriorly without chaetae (Fig. 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">20</figureCitation>
). Tenaculum with 4+4 teeth and two chaetae. Anterior furcal subcoxa with 5-7, posterior one with three chaetae. Anterior side of manubrium without chaetae, posterior side with 4+4 laterobasal and 8-9 chaetae on main part, without lateral chaetae. Dens short, without crenulation, with one rigid and short anterior and three posterior chaetae (Figs 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">21</figureCitation>
, 
<figureCitation captionStart="Figures 15–22" captionStartId="F4" captionText="Figures 15 - 22. C. abulbus sp. nov. 15 Abd. IV and fused Abd. V and V 16 labial palp 17, 18 apical part of Ant. IV, different views 19 ocelli and PAO 20 ventral tube, ventral view 21 furca, lateral view 22 furcal area, ventral view. Abbreviations: org organit, sms subapical ms-chaeta, ss s-chaeta." figureDoi="10.3897/zookeys.945.51860.figures15-22" httpUri="https://binary.pensoft.net/fig/428162" pageId="0" pageNumber="99">22</figureCitation>
). Mucro bidentate. Ratio manubrium: dens: mucro = 3.3-3.8: 1.7-2.3: 1.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="etymology">
<paragraph pageId="0" pageNumber="99">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="99">
The name is derived from the absence of apical bulb on Ant. IV to stress the difference from 
<taxonomicName lsidName="C. bulbus" pageId="0" pageNumber="99" rank="species" species="bulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. bulbus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="distribution">
<paragraph pageId="0" pageNumber="99">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="99">Currently known from indigenous vegetation in the Jonkershoek Nature Reserve, Stellenbosch.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="discussion">
<paragraph pageId="0" pageNumber="99">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="99">
Unlike 
<taxonomicName lsidName="C. bulbus" pageId="0" pageNumber="99" rank="species" species="bulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. bulbus</emphasis>
</taxonomicName>
sp. nov., the new species has no antennal bulb. Nevertheless, the two species form a rather well-defined group differing from almost all congeners by a simple maxillary palp, two sublobal hairs, chili-shaped organite on Ant. IV, and the absence of chaetae on the anterior side of the manubrium. Concerning the last character, only 
<taxonomicName lsidName="C. nivicolus" pageId="0" pageNumber="99" rank="species" species="nivicolus">
<emphasis italics="true" pageId="0" pageNumber="99">C. nivicolus</emphasis>
</taxonomicName>
(Salmon, 1965) and 
<taxonomicName lsidName="C. sverdrupi" pageId="0" pageNumber="99" rank="species" species="sverdrupi">
<emphasis italics="true" pageId="0" pageNumber="99">C. sverdrupi</emphasis>
</taxonomicName>
Lawrence, 1978 also lack this pair of chaetae, which is common to other species of the genus. Both mentioned species are inhabitants of Antarctic polar deserts and can hardly be conspecific to 
<taxonomicName lsidName="C. abulbus" pageId="0" pageNumber="99" rank="species" species="abulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. abulbus</emphasis>
</taxonomicName>
sp. nov. found in dry sites in a subtropical climate. These two Antarctic species are very dark and have two clavate tenent hairs (vs. one in 
<taxonomicName lsidName="C. abulbus" pageId="0" pageNumber="99" rank="species" species="abulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. abulbus</emphasis>
</taxonomicName>
sp. nov.) on tibiotarsi I. In addition, 
<taxonomicName lsidName="C. nivicolus" pageId="0" pageNumber="99" rank="species" species="nivicolus">
<emphasis italics="true" pageId="0" pageNumber="99">C. nivicolus</emphasis>
</taxonomicName>
has no mucro (vs. present in the new species) while 
<taxonomicName lsidName="C. sverdrupi" pageId="0" pageNumber="99" rank="species" species="sverdrupi">
<emphasis italics="true" pageId="0" pageNumber="99">C. sverdrupi</emphasis>
</taxonomicName>
has very small PAO (more than twice longer than ocellus in 
<taxonomicName lsidName="C. abulbus" pageId="0" pageNumber="99" rank="species" species="abulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. abulbus</emphasis>
</taxonomicName>
sp. nov.). Recently, 
<taxonomicName genus="Gressitacantha" lsidName="Gressitacantha terranova" pageId="0" pageNumber="99" rank="species" species="terranova">
<emphasis italics="true" pageId="0" pageNumber="99">Gressitacantha terranova</emphasis>
</taxonomicName>
Wise, 1967 was moved to 
<taxonomicName authorityName="Willem" authorityYear="1902" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus" order="Entomobryomorpha" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Cryptopygus</emphasis>
</taxonomicName>
(Greenslade, 2015) adding another 
<taxonomicName authorityName="Willem" authorityYear="1902" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus" order="Entomobryomorpha" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Cryptopygus</emphasis>
</taxonomicName>
species without anterior chaetae on the manubrium. The differences between 
<taxonomicName lsidName="C. abulbus" pageId="0" pageNumber="99" rank="species" species="abulbus">
<emphasis italics="true" pageId="0" pageNumber="99">C. abulbus</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="C. terranovus" pageId="0" pageNumber="99" rank="species" species="terranovus">
<emphasis italics="true" pageId="0" pageNumber="99">C. terranovus</emphasis>
</taxonomicName>
are more numerous than those from 
<taxonomicName lsidName="C. nivicolus" pageId="0" pageNumber="99" rank="species" species="nivicolus">
<emphasis italics="true" pageId="0" pageNumber="99">C. nivicolus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="C. sverdrupi" pageId="0" pageNumber="99" rank="species" species="sverdrupi">
<emphasis italics="true" pageId="0" pageNumber="99">C. sverdrupi</emphasis>
</taxonomicName>
(in furca, arms of abdomen, length of macrochaetae, and others).
</paragraph>
</subSubSection>
</treatment>
</document>