141 lines
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141 lines
12 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1023.58733" ID-GBIF-Dataset="d65d87c4-f7ba-4396-8ab8-2d4faca8d85b" ID-PMC="PMC7969582" ID-Pensoft-Pub="1313-2970-1023-29" ID-Pensoft-UUID="345215FD8A6154B2AE1A93C3C10F2425" ID-PubMed="33776513" ID-ZooBank="E9C8BCFD024249879E364533674D5387" ModsDocID="1313-2970-1023-29" checkinTime="1615410361329" checkinUser="pensoft" docAuthor="Bouteille, Emilien, Cesne, Maxime Le & Soulier-Perkins, Adeline" docDate="2021" docId="A49DAF05C29D5EA3AB2FA4681CDC0CEF" docLanguage="en" docName="ZooKeys 1023: 29-47" docOrigin="ZooKeys 1023" docPubDate="2021-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1023.58733" docTitle="Locris Stal 1866" docType="treatment" docVersion="5" id="345215FD8A6154B2AE1A93C3C10F2425" lastPageNumber="29" masterDocId="345215FD8A6154B2AE1A93C3C10F2425" masterDocTitle="Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species" masterLastPageNumber="47" masterPageNumber="29" pageNumber="29" updateTime="1668149888789" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bouteille, Emilien</mods:namePart>
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<mods:affiliation>Mecanismes adaptatifs et evolution (MECADEV), Museum national d'histoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cesne, Maxime Le</mods:namePart>
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<mods:affiliation>Mecanismes adaptatifs et evolution (MECADEV), Museum national d'histoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soulier-Perkins, Adeline</mods:namePart>
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<mods:affiliation>Mecanismes adaptatifs et evolution (MECADEV), Museum national d'histoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France</mods:affiliation>
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<mods:nameIdentifier type="email">adeline.soulier@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-03-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1023</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>47</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1023.58733</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1023.58733</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1023-29</mods:identifier>
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<mods:identifier type="ZooBank">E9C8BCFD024249879E364533674D5387</mods:identifier>
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<mods:identifier type="Pensoft-UUID">345215FD8A6154B2AE1A93C3C10F2425</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="180302201" LSID="urn:lsid:plazi:treatment:A49DAF05C29D5EA3AB2FA4681CDC0CEF" httpUri="http://treatment.plazi.org/id/A49DAF05C29D5EA3AB2FA4681CDC0CEF" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection pageId="0" pageNumber="29" type="nomenclature">
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName LSID="A49DAF05-C29D-5EA3-AB2F-A4681CDC0CEF" authority="Stal, 1866" authorityName="Stal" authorityYear="1866" class="Insecta" family="Cercopidae" genus="Locris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Locris" order="Hemiptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
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Locris
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<normalizedToken originalValue="Stål">Stal</normalizedToken>
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, 1866
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="type species">
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<paragraph pageId="0" pageNumber="29">
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<typeStatus>Type</typeStatus>
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species.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1794" class="Insecta" family="Cercopidae" genus="Locris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Locris rubra" order="Hemiptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="rubra">
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<emphasis italics="true" pageId="0" pageNumber="29">Locris rubra</emphasis>
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</taxonomicName>
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(Fabricius, 1794)
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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The genus
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<taxonomicName authorityName="Stal" authorityYear="1866" class="Insecta" family="Cercopidae" genus="Locris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Locris" order="Hemiptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="29">Locris</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Stål">Stal</normalizedToken>
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, 1866 was largely studied by
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<bibRefCitation author="Lallemand, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="29" refId="B16" refString="Lallemand, V, 1949. Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1-193." title="Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1 - 193." year="1949">Lallemand (1949)</bibRefCitation>
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on the basis of morphological characters that are not all completely consistent in all species according to our observations. These characters are as follows: postclypeus rounded with a medio-longitudinal carina and transverse ridges, when observed laterally; it can be, either rounded (
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<taxonomicName lsidName="L. rubra" pageId="0" pageNumber="29" rank="species" species="rubra">
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<emphasis italics="true" pageId="0" pageNumber="29">L. rubra</emphasis>
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</taxonomicName>
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(Fabricius, 1794)), angular (
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<taxonomicName lsidName="L. vestigans" pageId="0" pageNumber="29" rank="species" species="vestigans">
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<emphasis italics="true" pageId="0" pageNumber="29">L. vestigans</emphasis>
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</taxonomicName>
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Jacobi, 1904) or protruding (
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<taxonomicName lsidName="L. schmidti" pageId="0" pageNumber="29" rank="species" species="schmidti">
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<emphasis italics="true" pageId="0" pageNumber="29">L. schmidti</emphasis>
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</taxonomicName>
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Jacobi, 1910). Rostrum is short and barely extends to base of median trochanters. Antennae are short and their length equals diameter of eyes. Vertex is broader than long and its length equals half width of pronotum. Ocelli are much closer to each other than to compound eyes and are medium to small in size, except in
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<taxonomicName lsidName="L. atra" pageId="0" pageNumber="29" rank="species" species="atra">
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<emphasis italics="true" pageId="0" pageNumber="29">L. atra</emphasis>
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</taxonomicName>
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Lallemand, 1923 where they are very large (
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<bibRefCitation author="Lallemand, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="29" refId="B16" refString="Lallemand, V, 1949. Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1-193." title="Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1 - 193." year="1949">Lallemand 1949</bibRefCitation>
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). Pronotum is large, with a usually straight posterior margin but weakly indented in some species like
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<taxonomicName lsidName="L. maculata" pageId="0" pageNumber="29" rank="species" species="maculata">
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<emphasis italics="true" pageId="0" pageNumber="29">L. maculata</emphasis>
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</taxonomicName>
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(Fabricius, 1781) according to
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<bibRefCitation author="Lallemand, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="29" refId="B16" refString="Lallemand, V, 1949. Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1-193." title="Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1 - 193." year="1949">Lallemand (1949)</bibRefCitation>
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and may have a more or less distinct carina in middle. Scutellum is as long as wide and has three dimples, two small ones on anterior margin and one large centred. Tegmina are about 3 times as long as wide, cubital and median veins are fused from base to middle of tegmen, apical veins network is relatively dense. A spine is present on posterior tibiae. In genitalia, males have thin subgenital plates that look like a filament (
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<bibRefCitation author="Lallemand, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="29" refId="B16" refString="Lallemand, V, 1949. Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1-193." title="Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1 - 193." year="1949">Lallemand 1949</bibRefCitation>
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) curved up or downward; they are wide and sometimes silky at base.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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In describing the genus
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<taxonomicName authorityName="Stal" authorityYear="1866" class="Insecta" family="Cercopidae" genus="Locris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Locris" order="Hemiptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="29">Locris</emphasis>
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</taxonomicName>
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<bibRefCitation author="Lallemand, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="29" refId="B16" refString="Lallemand, V, 1949. Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1-193." title="Revision des Cercopinae (HemipteraHomoptera) Premiere partie. Memoires de l'Institut Royal des Sciences Naturelles de Belgique (Ser. 2) 32: 1 - 193." year="1949">Lallemand (1949)</bibRefCitation>
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listed a series of exceptions in order to include more species in the genus. As a result, our view is that now the homogeneity of the genus and its taxonomic unity is questionable. However, the aim of our work here is not to revise the entire genus (which is present in
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<collectingCountry name="Madagascar">Madagascar</collectingCountry>
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and the whole of Africa except for the northern countries
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<collectingCountry name="Egypt">Egypt</collectingCountry>
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,
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<collectingCountry name="Tunisia">Tunisia</collectingCountry>
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,
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<collectingCountry name="Libya">Libya</collectingCountry>
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,
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<collectingCountry name="Algeria">Algeria</collectingCountry>
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and
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<collectingCountry name="Morocco">Morocco</collectingCountry>
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) but to provide a clear identification for the few species and subspecies of
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<taxonomicName authorityName="Stal" authorityYear="1866" class="Insecta" family="Cercopidae" genus="Locris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Locris" order="Hemiptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="29">Locris</emphasis>
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</taxonomicName>
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present in
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<collectingCountry name="Madagascar">Madagascar</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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