treatments-xml/data/27/50/ED/2750ED0DFA485C54870787F934361C04.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.963.53470" ID-GBIF-Dataset="0831704f-4ecf-4989-8072-defde35921ec" ID-PMC="PMC7458944" ID-Pensoft-Pub="1313-2970-963-45" ID-Pensoft-UUID="FE8C7A0A85B45DF7848BFAB844F93C28" ID-PubMed="32922131" ID-ZooBank="9636C9F2C6BD4B34BCC6ED214C7B0D19" ModsDocID="1313-2970-963-45" checkinTime="1598316761636" checkinUser="pensoft" docAuthor="Englund, William F., Njoroge, Laban, Bistroem, Olof, Miller, Kelly B., Bilton, David T. &amp; Bergsten, Johannes" docDate="2020" docId="2750ED0DFA485C54870787F934361C04" docLanguage="en" docName="ZooKeys 963: 45-79" docOrigin="ZooKeys 963" docSource="http://dx.doi.org/10.3897/zookeys.963.53470" docTitle="Agabus raffrayi Sharp 1882" docType="treatment" docVersion="4" id="FE8C7A0A85B45DF7848BFAB844F93C28" lastPageNumber="45" masterDocId="FE8C7A0A85B45DF7848BFAB844F93C28" masterDocTitle="Taxonomic revision of the Afrotropical Agabus raffrayi species group with the description of four new species (Coleoptera, Dytiscidae)" masterLastPageNumber="79" masterPageNumber="45" pageNumber="45" updateTime="1668169378299" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Afrotropical Agabus raffrayi species group with the description of four new species (Coleoptera, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Englund, William F.</mods:namePart>
<mods:affiliation>Swedish Museum of Natural History, Department of Zoology, Box 50007, SE- 10405 Stockholm, Sweden</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Njoroge, Laban</mods:namePart>
<mods:affiliation>National Museums of Kenya, Section of Invertebrate Zoology, Museum Hill, P. O. BOX 40658 - 00100, Nairobi, Kenya</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bistroem, Olof</mods:namePart>
<mods:affiliation>Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bilton, David T.</mods:namePart>
<mods:affiliation>Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bergsten, Johannes</mods:namePart>
<mods:affiliation>Swedish Museum of Natural History, Department of Zoology, Box 50007, SE- 10405 Stockholm, Sweden</mods:affiliation>
<mods:nameIdentifier type="email">johannes.bergsten@nrm.se</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>963</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.963.53470</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.963.53470</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-963-45</mods:identifier>
<mods:identifier type="ZooBank">9636C9F2C6BD4B34BCC6ED214C7B0D19</mods:identifier>
<mods:identifier type="Pensoft-UUID">FE8C7A0A85B45DF7848BFAB844F93C28</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="167293858" LSID="urn:lsid:plazi:treatment:2750ED0DFA485C54870787F934361C04" httpUri="http://treatment.plazi.org/id/2750ED0DFA485C54870787F934361C04" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="2750ED0D-FA48-5C54-8707-87F934361C04" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi Sharp, 1882</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">, 2B</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Agabus spp., protarsomeres in anterior view A A. dytiscoides B A. raffrayi. PT 4 L = measured distance for length of protarsomere 4; CL = measured distance for length of protarsal claw." figureDoi="10.3897/zookeys.963.53470.figure3" httpUri="https://binary.pensoft.net/fig/444400" pageId="0" pageNumber="45">, 3B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">, 5A-C</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">, 8D</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">, 10D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11I</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi</taxonomicName>
Sharp, 1882: 501-502
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
, 1905: 224-225 (Syn. 
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">raffrayi</emphasis>
</taxonomicName>
<normalizedToken originalValue="“Abyssinia”">&quot;Abyssinia&quot;</normalizedToken>
[Ethiopia]; 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">
<emphasis italics="true" pageId="0" pageNumber="45">limbicollis</emphasis>
</taxonomicName>
&quot;Abyssinie: Auato, au bord du Nil Bleu, dans le Gindeberat&quot; [Ethiopia: Auato, on the banks of the Blue Nile, in the Gindeberat].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Lectotype</emphasis>
♂ of 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">raffrayi</emphasis>
</taxonomicName>
(BMNH) labelled: &quot;♂ Abyssinia, Raffray 782&quot;, 
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, &quot;Sharp Coll 1905-313.&quot;, &quot;Type 782 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Agabus</taxonomicName>
Raffrayi n.sp. Abyssinia&quot;, &quot;LECTOTYPUS ♂ 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi</taxonomicName>
Sharp, 1882 Des. A.Nilsson, 1989&quot;. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Lectotype</emphasis>
♂ of 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">
<emphasis italics="true" pageId="0" pageNumber="45">limbicollis</emphasis>
</taxonomicName>
(MNB) labelled: &quot;N.O. - Africa, Schoa, Falle O. Neumann S.&quot;, &quot;610 Falle&quot;, &quot; 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken originalValue="Rég">Reg</normalizedToken>
. Type.&quot;, &quot;LECTOTYPUS ♂ 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
, 1905. Des. Nilsson -90&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="diagnosis">
<paragraph pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">
With a prolonged preapical section of male aedeagus and a pronotal hypomeron which is not visible in lateral view, this species is most similar to 
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
. From the former it is separated by its broader metasternal wing (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">2</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
) and from the latter by the lack of interocular spots (compare Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4D, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
Habitus as in Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11D, I</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
: Head black, most specimens with a small rufous anterior area, interocular spots not present. Pronotum black with rufous margins. Elytra rufotestaceous to brown. Ventral surface black, hypomeron and epipleuron testaceous. Legs rufous to rufopiceous. Antennae and palpi testaceous to rufotestaceous.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
</taxonomicName>
: Medium impressed on head, pronotum and elytra, similar in both sexes. Composed of a mixture of small and somewhat larger, uneven meshes.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
</taxonomicName>
: Body length: 6.96-8.24 mm (see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
). Hypomeron not visible in strict lateral view (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10D</figureCitation>
), lateral bead of pronotum well defined (see Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10D</figureCitation>
). Metasternal wing broad, WC/WS less than 2.9 (see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">2B</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
). Pronotum broad, more than twice as broad as interocular distance (see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, as in Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
: Protarsal claws long,&gt; 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
as long as protarsomere 4 in all males and most females (see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
, Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
). Metatarsomeres short and broad; metatarsomere 2 &lt;1.6 
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad (see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
), metatarsomere 5 &lt;3.0 times as long as broad (see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
: Subapically broadened, and prolonged between the subapical broadening and the apical and subapical teeth. Subapical tooth with quite variable shape (see Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8D</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">5A-C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Female</emphasis>
: Externally similar to males but colour of the elytra tends to be slightly lighter.
</paragraph>
<caption doi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 8.</emphasis>
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., aedeagus in lateral view 
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">E</emphasis>
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">F</emphasis>
<taxonomicName lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="45">G</emphasis>
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="45">H</emphasis>
<taxonomicName lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="45">I</emphasis>
<taxonomicName lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
Ethiopia (see Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">1</figureCitation>
). 
<bibRefCitation DOI="https://doi.org/10.1080/03749444.1975.10736808" author="Rocchi, S" journalOrPublisher="): Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906. Uppsala: Almquist &amp; Wiksells Boktryckeri-A. B." pageId="0" pageNumber="45" refId="B23" refString="Rocchi, S, 1975. Coleotteri ditiscidi dell'Africa orientale. Monitore Zoologico Italiano (N.S. Suppl. 6) 3: 45-56. https://doi.org/10.1080/03749444.1975.10736808" title="Coleotteri ditiscidi dell'Africa orientale. Monitore Zoologico Italiano (N. S. Suppl. 6) 3: 45 - 56." url="https://doi.org/10.1080/03749444.1975.10736808" year="1975">Rocchi (1975)</bibRefCitation>
listed the distribution of 
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
to also include the Democratic Republic of the Congo, Uganda, Rwanda, Tanzania, Zimbabwe, and South Africa but these specimens are likely to belong to other species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">
Found in small, often temporary, streams and pools in streambeds at elevations between 2100 to 3200 m (
<bibRefCitation DOI="https://doi.org/10.1080/01650429009361398" author="Nilsson, AN" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="45" pagination="135 - 144" refId="B17" refString="Nilsson, AN, Persson, S, 1990. Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned. Aquatic Insects 12 (3): 135 - 144, DOI: https://doi.org/10.1080/01650429009361398" title="Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned." url="https://doi.org/10.1080/01650429009361398" volume="12" year="1990">Nilsson and Persson 1990</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.33338/ef.83753" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="45" pagination="57 - 94" refId="B18" refString="Nilsson, AN, Persson, S, 1993. Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia. Entomologica Fennica 4: 57 - 94, DOI: https://doi.org/10.33338/ef.83753" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." url="https://doi.org/10.33338/ef.83753" volume="4" year="1993">1993</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
The name refers to the collector of the type specimens, Achille Raffray. The name of the synonym 
<taxonomicName lsidName="A. limbicollis" pageId="0" pageNumber="45" rank="species" species="limbicollis">
<emphasis italics="true" pageId="0" pageNumber="45">A. limbicollis</emphasis>
</taxonomicName>
refers to the well-defined lateral bead of the pronotum (Latin: 
<emphasis italics="true" pageId="0" pageNumber="45">limbus</emphasis>
= border, 
<emphasis italics="true" pageId="0" pageNumber="45">collum</emphasis>
= neck).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="comments">
<paragraph pageId="0" pageNumber="45">Comments.</paragraph>
<paragraph pageId="0" pageNumber="45">
The fact that 
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
are distinguishable only on the width of the metasternal wing led some previous authors to suggest the occurrence of a single species which was dimorphic with regard to this character (
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" author="Jackson, DJ" journalOrPublisher="Proceedings of the Royal entomological Society of London (A)" pageId="0" pageNumber="45" pagination="1 - 11" refId="B9" refString="Jackson, DJ, 1956. Dimorphism of the metasternal wings in Agabus raffrayi Sharp and A. labiatus Brahm (Col., Dytiscidae) and its relation to capacity for flight. Proceedings of the Royal entomological Society of London (A) 31: 1 - 11, DOI: https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" title="Dimorphism of the metasternal wings in Agabus raffrayi Sharp and A. labiatus Brahm (Col., Dytiscidae) and its relation to capacity for flight." url="https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" volume="31" year="1956">Jackson 1956</bibRefCitation>
). 
<bibRefCitation DOI="https://doi.org/10.1080/01650429009361398" author="Nilsson, AN" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="45" pagination="135 - 144" refId="B17" refString="Nilsson, AN, Persson, S, 1990. Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned. Aquatic Insects 12 (3): 135 - 144, DOI: https://doi.org/10.1080/01650429009361398" title="Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned." url="https://doi.org/10.1080/01650429009361398" volume="12" year="1990">Nilsson and Persson (1990)</bibRefCitation>
provided a detailed account of this argument, analysed a large series of specimens and concluded that the variation should rather be interpreted as two separate species. We agree with this assessment and concur that male genitalia are not diagnostic for these two species, only the width of the metasternal wing being reliable. In our measurements, the pronotum is marginally broader in 
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
but the small sample size forbids any strong conclusions at present (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation DOI="https://doi.org/10.1080/03946975.1992.10539193" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="45" pagination="207 - 217" refId="B16" refString="Nilsson, AN, 1992b. Larval morphology of six species of Afrotropical Agabus Leach 1817 (coleoptera Dytiscidae). Tropical Zoology 5 (2): 207 - 217, DOI: https://doi.org/10.1080/03946975.1992.10539193" title="Larval morphology of six species of Afrotropical Agabus Leach 1817 (coleoptera Dytiscidae)." url="https://doi.org/10.1080/03946975.1992.10539193" volume="5" year="1992 b">Nilsson (1992b)</bibRefCitation>
described the larval morphology of 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
</taxonomicName>
along with some representatives of two other Afrotropical 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
groups.
</paragraph>
<caption doi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 9.</emphasis>
South African 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., microreticulation of male elytral disc 
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. holotype, Keurboom, Western Cape 
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Groote Swartberg, Western Cape 
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Sentinel Peak, KZN Drakensberg 
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Harkerville Forest, Western Cape 
<emphasis bold="true" pageId="0" pageNumber="45">E</emphasis>
<taxonomicName lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov. Kamiesberg, Northern Cape 
<emphasis bold="true" pageId="0" pageNumber="45">F</emphasis>
<taxonomicName lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov. holotype, Rattelrivier, Western Cape.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 10.</emphasis>
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., hypomeron in lateral view 
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
. White arrows indicate hypomeron.
</paragraph>
</caption>
<caption ID-Table-UUID="EC20EA9B30576BE179C66F15DBC0C3C6" httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Table 1.</emphasis>
Morphological characters in the 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
</taxonomicName>
group. WC/WS = width of metacoxal plate /width of metasternal wing, PW/ID = width of pronotum/interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<table pageId="0" pageNumber="45">
<tr pageId="0" pageNumber="45">
<th colspan="1" pageId="0" pageNumber="45" rowspan="2">Species</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="2">Sex</th>
<th colspan="5" pageId="0" pageNumber="45" rowspan="1">WC/WS</th>
<th colspan="5" pageId="0" pageNumber="45" rowspan="1">PW/ID</th>
<th colspan="5" pageId="0" pageNumber="45" rowspan="1">TL</th>
</tr>
<tr pageId="0" pageNumber="45">
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.60</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.69</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.64</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.07</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.04</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.11</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.29</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.65</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.55</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.07</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.13</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">9.12</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.70</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.31</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.00</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.13</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.00</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.60</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.48</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.17</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.88</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.40</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.14</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.22</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.98</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1.96</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.02</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.76</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.55</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.18</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.19</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.36</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.44</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.40</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.11</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4.15</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.55</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.23</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.12</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.30</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.22</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6.80</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.53</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.40</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.12</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.49</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.28</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.04</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.67</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.36</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.90</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.41</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.19</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.67</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.21</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.73</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.63</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.09</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.86</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.20</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.11</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.84</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.58</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.28</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.25</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6.96</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.70</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.45</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.46</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.71</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.61</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.25</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.29</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.01</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.82</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.31</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.29</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.29</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.22</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.79</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.81</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.39</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.30</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.45</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.34</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.92</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.14</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.18</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.00</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.27</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.03</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.21</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.70</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.35</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.65</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.91</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.21</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.37</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.28</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.21</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.83</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.29</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.67</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.75</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.71</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.18</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.16</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.02</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0</td>
</tr>
<tr pageId="0" pageNumber="45">
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">3.67</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4.54</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4.01</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">2.17</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.60</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">8.00</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">7.86</td>
<td colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>