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<document id="4F80895CB822760C27E361AB184B906B" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FF8A3676FF76F9AFBAC2FE7A" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nicagus obscurus" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="11" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="62AE3339555506AE74EAE82BE00662A0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A5D0E8B76D46A55F4464187E443EEA20">
<mods:title id="3CC2B74ED3FFB6C238B6471B16EBDB68">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
</mods:titleInfo>
<mods:name id="EB8E1EA45BB556F49BB432C4501F98D9" type="personal">
<mods:role id="437F2C5897E470DAA62AE5F9FDB9F412">
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<mods:namePart id="5A413D5F550865DA335A8233F1D6FD1C">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="B4C959ABE6528236D45D32E865840A2C">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
</mods:name>
<mods:name id="1B9957FF15BE13739C4BBEECDC0ED0F0" type="personal">
<mods:role id="F2EA837BD2F477324742996F5240A4D2">
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</mods:role>
<mods:namePart id="02A0F88DD41F42DA5306AD11AC5B42B9">Veal, Doug A.</mods:namePart>
<mods:affiliation id="305B13FC292C24FA4EAE69134699CD47">168 Thompson Drive SE Cedar Rapids, Iowa 52403</mods:affiliation>
</mods:name>
<mods:name id="A31940D53E0B50A1110A43A12B67DC76" type="personal">
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</mods:role>
<mods:namePart id="664CD42077B064B41EF119BF9E01B3C5">Lago, Paul K.</mods:namePart>
<mods:affiliation id="4B35D9A783799E4B43BA721AB8846655">Biology Department, University of Mississippi University, Mississippi 38677</mods:affiliation>
</mods:name>
<mods:typeOfResource id="889909DCEB50345CCF09D030D7BF0D7F">text</mods:typeOfResource>
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<mods:title id="DCB4399A436188F3C40952A86120AF84">Insecta Mundi</mods:title>
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<mods:part id="FDBF525FBF64A585376976A05F84AC86">
<mods:date id="DD9A5C8EC4A496EB18BDBC426728FE9E">2020</mods:date>
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<mods:number id="FE2CCCFD35D716394C11EF076BFE451C">2020-08-28</mods:number>
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<mods:number id="A1E80FCE3E0EE0AB18F2064E3560C8EB">2020</mods:number>
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<mods:number id="FA8E787A8DAFB49FD427ADB8847FDE5A">787</mods:number>
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<mods:start id="B79C774FD7F9459941F91E9D7A086A8B">1</mods:start>
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<mods:classification id="D56251C88845F08D70B7A3CF3D101FC3">journal article</mods:classification>
<mods:identifier id="58FBBF2DEB83821B6E6CA605574B4FFA" type="DOI">10.5281/zenodo.5353792</mods:identifier>
<mods:identifier id="98ACCDDF87FD44BF750BBC8D8A08B4A3" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="09DA46CC6D7031BABC9B2454E5993A90" type="Zenodo-Dep">5353792</mods:identifier>
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</mods:mods>
<treatment id="28608792FF8A3676FF76F9AFBAC2FE7A" LSID="urn:lsid:plazi:treatment:28608792FF8A3676FF76F9AFBAC2FE7A" httpUri="http://treatment.plazi.org/id/28608792FF8A3676FF76F9AFBAC2FE7A" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
<subSubSection id="E8D3650FFF8A3677FF76F9AFBE3EF903" pageId="12" pageNumber="11" type="nomenclature">
<paragraph id="A0763684FF8A3677FF76F9AFB9BAF9DE" blockId="12.[192,653,1602,1626]" box="[192,653,1602,1626]" pageId="12" pageNumber="11">
<heading id="FB3E81E8FF8A3677FF76F9AFB9BAF9DE" bold="true" box="[192,653,1602,1626]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
<taxonomicName id="67C94D07FF8A3677FF76F9AFB9BAF9DE" ID-CoL="73XS3" authority="(LeConte, 1847)" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[192,653,1602,1626]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="92BDEA96FF8A3677FF76F9AFB9BAF9DE" bold="true" box="[192,653,1602,1626]" pageId="12" pageNumber="11">
<emphasis id="92BDEA96FF8A3677FF76F9AFBA86F9DE" bold="true" box="[192,433,1602,1626]" italics="true" pageId="12" pageNumber="11">Nicagus obscurus</emphasis>
(LeConte, 1847)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A0763684FF8A3677FF76F982BE3EF903" blockId="12.[192,1289,1647,1671]" box="[192,1289,1647,1671]" pageId="12" pageNumber="11">
<bibRefCitation id="C4584B75FF8A3677FF76F982BAC5F903" author="Paulsen, M. J. &amp; A. D. Smith" box="[192,498,1647,1671]" pageId="12" pageNumber="11" pagination="45 - 60" refId="ref33376" refString="Paulsen, M. J., and A. D. Smith. 2005. A new species of stag beetles from sand dunes in west Texas, and a synopsis of the genus Nicagus (Coleoptera: Lucanidae: Aesalinae: Nicagini). Zootaxa 1050: 45 - 60." type="journal article" year="2005">Paulsen and Smith 2005</bibRefCitation>
;
<bibRefCitation id="C4584B75FF8A3677FE48F982B9A7F903" author="Evans, A. V." box="[510,656,1647,1671]" pageId="12" pageNumber="11" refId="ref31489" refString="Evans, A. V. 2014. Beetles of eastern North America. Princeton University Press; Princeton, NJ. 560 p." type="book" year="2014">Evans 2014</bibRefCitation>
; ISIC; USNM; WIRC; DAVC; ELFC; MJPC; PKLC.
</paragraph>
</subSubSection>
<subSubSection id="E8D3650FFF8A3676FF76F97BBAC2FE7A" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="description">
<paragraph id="A0763684FF8A3677FF76F97BB911F92A" blockId="12.[192,550,1686,1710]" box="[192,550,1686,1710]" pageId="12" pageNumber="11">Collection dates: April–June.</paragraph>
<paragraph id="A0763684FF8A3677FF76F953B957F8E1" blockId="12.[192,1442,1726,1894]" pageId="12" pageNumber="11">Males and females collected small wooded stream, under and on debris, on sandbar, and sifted from sand. Specimens also collected in flight above sandbar and near stream. A few females have been collected in a small chamber under debris sandbar small wooded stream. Males have been observed “swarming” or flying zig-zag and circling in groups of ten to twenty or more over a small area of sand- bar. Specimens have also been collected under horse dung on a bike path near wooded stream and flying some distance from stream.</paragraph>
<paragraph id="A0763684FF8B3676FF76FF3AB8EBFEC1" blockId="13.[191,1441,215,326]" pageId="13" pageNumber="12">
<materialsCitation id="10A13CD9FF8B3676FF76FF3ABF48FF6B" box="[192,1151,215,239]" collectingDate="1929-05-23" collectionCode="ISIC" collectorName="H. M. Harris" county="Story Co." location="Ames" municipality="Specimen" pageId="13" pageNumber="12" specimenCount="1" stateProvince="Iowa">
<collectingMunicipality id="4012ACFEFF8B3676FF76FF3ABA00FF6B" box="[192,311,215,239]" pageId="13" pageNumber="12">Specimen</collectingMunicipality>
label data:
<location id="A516605FFF8B3676FE70FF3AB939FF6B" LSID="urn:lsid:plazi:treatment:28608792FF8A3676FF76F9AFBAC2FE7A:A516605FFF8B3676FE70FF3AB939FF6B" box="[454,526,215,239]" county="Story Co." municipality="Specimen" name="Ames" pageId="13" pageNumber="12" stateProvince="Iowa">Ames</location>
,
<collectingCounty id="49174E08FF8B3676FDAFFF3AB9BAFF6B" box="[537,653,215,239]" pageId="13" pageNumber="12">Story Co.</collectingCounty>
,
<collectingRegion id="620DF866FF8B3676FD2EFF3AB9E0FF6A" box="[664,727,215,238]" country="United States of America" name="Iowa" pageId="13" pageNumber="12">Iowa</collectingRegion>
,
<date id="D4771044FF8B3676FD54FF3AB8B6FF6B" box="[738,897,215,239]" pageId="13" pageNumber="12" value="1929-05-23">
<collectingDate id="C433E9ACFF8B3676FD54FF3AB8B6FF6B" box="[738,897,215,239]" pageId="13" pageNumber="12" value="1929-05-23">23 May 1929</collectingDate>
</date>
,
<collectorName id="0D3C5352FF8B3676FC3BFF3ABF1DFF6B" box="[909,1066,215,239]" pageId="13" pageNumber="12">H. M. Harris</collectorName>
(
<collectionCode id="C6D8AE41FF8B3676FB8FFF3ABF41FF6B" box="[1081,1142,215,239]" pageId="13" pageNumber="12">ISIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="10A13CD9FF8B3676FB3CFF3AB88BFE88" collectingDate="1931-04-27" collectionCode="ISIC" collectorName="R. H. Nelson &amp; Howden" county="Story Co." location="Ames" pageId="13" pageNumber="12" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFF8B3676FB3CFF3ABFE5FF6B" LSID="urn:lsid:plazi:treatment:28608792FF8A3676FF76F9AFBAC2FE7A:A516605FFF8B3676FB3CFF3ABFE5FF6B" box="[1162,1234,215,239]" county="Story Co." name="Ames" pageId="13" pageNumber="12" stateProvince="Iowa">Ames</location>
,
<collectingCounty id="49174E08FF8B3676FB6BFF3ABE66FF6B" box="[1245,1361,215,239]" pageId="13" pageNumber="12">Story Co.</collectingCounty>
,
<collectingRegion id="620DF866FF8B3676FAEBFF3ABEABFF6A" box="[1373,1436,215,238]" country="United States of America" name="Iowa" pageId="13" pageNumber="12">Iowa</collectingRegion>
,
<date id="D4771044FF8B3676FF09FF19BA5CFE88" box="[191,363,244,268]" pageId="13" pageNumber="12" value="1931-04-27">
<collectingDate id="C433E9ACFF8B3676FF09FF19BA5CFE88" box="[191,363,244,268]" pageId="13" pageNumber="12" value="1931-04-27">27 April 1931</collectingDate>
</date>
,
<collectorName id="0D3C5352FF8B3676FECEFF19B92EFE88" box="[376,537,244,268]" pageId="13" pageNumber="12">R. H. Nelson</collectorName>
, determined
<collectorName id="0D3C5352FF8B3676FD0AFF19B816FE88" box="[700,801,244,268]" pageId="13" pageNumber="12">Howden</collectorName>
1955 (
<collectionCode id="C6D8AE41FF8B3676FCC0FF19B884FE88" box="[886,947,244,268]" pageId="13" pageNumber="12">ISIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="10A13CD9FF8B3676FC7FFF19B8E0FEC1" collectingDate="2006-05-06" collectingDateMax="2011-05-20" collectingDateMin="2006-05-06" collectionCode="ELFC" county="Lee Co." location="Shimek State Forest" pageId="13" pageNumber="12" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFF8B3676FC7FFF19BFFCFE8F" LSID="urn:lsid:plazi:treatment:28608792FF8A3676FF76F9AFBAC2FE7A:A516605FFF8B3676FC7FFF19BFFCFE8F" box="[969,1227,244,268]" county="Lee Co." name="Shimek State Forest" pageId="13" pageNumber="12" stateProvince="Iowa">Shimek State Forest</location>
,
<collectingCounty id="49174E08FF8B3676FB6EFF19BE0FFE88" box="[1240,1336,244,268]" pageId="13" pageNumber="12">Lee Co.</collectingCounty>
,
<collectingRegion id="620DF866FF8B3676FAF3FF19BEB3FE8F" box="[1349,1412,244,267]" country="United States of America" name="Iowa" pageId="13" pageNumber="12">Iowa</collectingRegion>
,
<date id="D4771044FF8B3676FA26FF19BA08FEAC" pageId="13" pageNumber="12" value="2006-05-06">
<collectingDate id="C433E9ACFF8B3676FA26FF19BA08FEAC" pageId="13" pageNumber="12" value="2006-05-06">6 May 2006</collectingDate>
</date>
, small chamber under piece of debris sandbar [female],
<date id="D4771044FF8B3676FC49FEFCBFA9FEAD" box="[1023,1182,272,297]" pageId="13" pageNumber="12" value="2010-04-02">
<collectingDate id="C433E9ACFF8B3676FC49FEFCBFA9FEAD" box="[1023,1182,272,297]" pageId="13" pageNumber="12" value="2010-04-02">2 April 2010</collectingDate>
</date>
, sifted from sandbar [male],
<date id="D4771044FF8B3676FEA9FEC0BAF7FEC1" box="[287,448,301,326]" pageId="13" pageNumber="12" value="2011-05-20">
<collectingDate id="C433E9ACFF8B3676FEA9FEC0BAF7FEC1" box="[287,448,301,326]" pageId="13" pageNumber="12" value="2011-05-20">20 May 2011</collectingDate>
</date>
, in flight over sandbar [male], ELF (
<collectionCode id="C6D8AE41FF8B3676FC36FEC0B8F8FEC1" box="[896,975,301,325]" pageId="13" pageNumber="12">ELFC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A0763684FF8B3676FF76FEB8B868FEE9" blockId="13.[192,863,341,365]" box="[192,863,341,365]" pageId="13" pageNumber="12">
County records (
<figureCitation id="38F22A01FF8B3676FE3DFEB8B931FEE9" box="[395,518,341,365]" captionStart="Figures 34–41" captionStartId="65.[192,288,1847,1869]" captionTargetBox="[141,1439,220,1805]" captionTargetPageId="65" captionText="Figures 34–41. Distribution of Scarabaeoidea in Iowa. 34) Nicagus obscurus (LeConte, 1847). 35) Ceruchus piceus (Weber, 1801). 36) Platycerus quercus (Weber, 1801). 37) Dorcus parallelus (Say, 1824). 38) Lucanus capreolus (Linnaeus, 1764). 39) Lucanus placidus Say, 1825. 40) Xenochodaeus americanus (Westwood, 1852). 41)" figureDoi="http://doi.org/10.5281/zenodo.5353806" httpUri="https://zenodo.org/record/5353806/files/figure.png" pageId="13" pageNumber="12">Figure 34</figureCitation>
): Johnson, Lee, Linn, Story.
</paragraph>
<paragraph id="A0763684FF8B3676FF76FE41B9F7FE40" blockId="13.[192,719,428,510]" box="[192,704,428,453]" pageId="13" pageNumber="12">
Subfamily
<taxonomicName id="67C94D07FF8B3676FEF0FE41B9F7FE40" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[326,704,428,453]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Syndesinae">SYNDESINAE MacLeay, 1819</taxonomicName>
</paragraph>
<paragraph id="A0763684FF8B3676FF76FE24B9F8FE65" blockId="13.[192,719,428,510]" box="[192,719,457,481]" pageId="13" pageNumber="12">
Tribe
<taxonomicName id="67C94D07FF8B3676FEBCFE24B9F8FE65" authority="Jacquelin du Val, 1857" authorityName="Jacquelin du Val" authorityYear="1857" box="[266,719,457,481]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Ceruchini">CERUCHINI Jacquelin du Val, 1857</taxonomicName>
</paragraph>
<paragraph id="A0763684FF8B3676FF76FE0BBAC2FE7A" blockId="13.[192,719,428,510]" box="[192,501,486,510]" pageId="13" pageNumber="12">
<taxonomicName id="67C94D07FF8B3676FF76FE0BBAC2FE7A" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[192,501,486,510]" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FF8B3676FF76FE0BBA04FE7A" box="[192,307,486,510]" italics="true" pageId="13" pageNumber="12">Ceruchus</emphasis>
MacLeay, 1819
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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