363 lines
113 KiB
XML
363 lines
113 KiB
XML
|
<document id="0982F09FA45DE5A2930000C29ABDE970" ID-CLB-Dataset="21361" ID-DOI="10.11646/zootaxa.4751.1.8" ID-GBIF-Dataset="d5e751b0-a468-493a-8e76-ef85b5150ce7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3974722" ID-ZooBank="B79BA86F-4F61-4016-8AEE-0AE9110BA253" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1596727948589" checkinUser="felipe" docAuthor="Zavala-Muñoz, Francisca, Bustos, Claudia A., Landaeta, Mauricio F. & Stantinidis, Peter Kon-" docDate="2020" docId="03B59B5DFFB4FFECFF26FE58FDBDF84E" docLanguage="en" docName="zootaxa.4751.1.8.pdf.imf" docOrigin="Zootaxa 4751 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callanthias platei Steindachner 1898" docType="treatment" docVersion="6" lastPageNumber="150" masterDocId="FF8CE325FFB6FFEBFFB1FFCDFF98FFAF" masterDocTitle="Larval development and osteology of Callanthias platei (Teleostei: Callanthiidae) from Desventuradas Islands, South Pacific" masterLastPageNumber="152" masterPageNumber="143" pageNumber="145" updateTime="1698844115900" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
|||
|
|
<mods:mods id="79E9C09F4DF3846A5AE4F0B757199E07" xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo id="B6B6BC1838459F993D0DD3B39B00D1E7">
|
|||
|
|
<mods:title id="EB0915796A6ACCF8EB0D57A394919AD4">Larval development and osteology of Callanthias platei (Teleostei: Callanthiidae) from Desventuradas Islands, South Pacific</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name id="7AA74FDE85E759927598025B11EB68A6" type="personal">
|
|||
|
|
<mods:role id="01E85C8B255F78E06B7AECA6D6EF6A6B">
|
|||
|
|
<mods:roleTerm id="ACEF4075CB6C8044CB79B52D589899E6">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="DA392607DDC361E6B3BA51A7C19D7F84">Zavala-Muñoz, Francisca</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="B3BC45C2D1EE2A4EA4A30EEC214F1552" type="personal">
|
|||
|
|
<mods:role id="C9321E5942489C3E4062C2C90B06CB95">
|
|||
|
|
<mods:roleTerm id="BD530B681B0926A2AB77ECBFB97295FE">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="C70367F6707D58B715FD450CFEAF14AC">Bustos, Claudia A.</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="1EC81048D8C6A8A2808E039CE6FF6068" type="personal">
|
|||
|
|
<mods:role id="700A44CE54BDD92082F0FACDF966BDD4">
|
|||
|
|
<mods:roleTerm id="0CE8A622ECCF8B10B5FB7C1E6EAE7F7E">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="F2B5406327F38FF66555DF328C81AAFC">Landaeta, Mauricio F.</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="A45762646C0462BE58FE6DC38F9789A2" type="personal">
|
|||
|
|
<mods:role id="7F1E4B822835C456D9C869713537295C">
|
|||
|
|
<mods:roleTerm id="75E0910E635AD9FA8F2CDA820177F6BB">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="0D537AA1E933532D7827D901D5090562">Stantinidis, Peter Kon-</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource id="0C017AC8DB130DCC0C060BD2427D503C">text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem id="8F5857E2D97166B683492BAC583E372C" type="host">
|
|||
|
|
<mods:titleInfo id="1E9E8F14554AE5EC640E720B41B9C43F">
|
|||
|
|
<mods:title id="E3DAF9B7D9633D32B0A6FE873777CE73">Zootaxa</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part id="69F5C3A444D1A08282476731B314E49B">
|
|||
|
|
<mods:date id="1E9D4F30015157EC6CF6DACEC155070C">2020</mods:date>
|
|||
|
|
<mods:detail id="5DBFB9D939610AA8C52C762199DF9AFE" type="pubDate">
|
|||
|
|
<mods:number id="FB9152E4F316AB7BC2C1AF9E5930AEE1">2020-03-16</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail id="19847B65BE271EB1881F0722D6F89B0A" type="volume">
|
|||
|
|
<mods:number id="91A4158854B0AA4C47E379A017E74675">4751</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail id="6B28DF07279801331E4492DAC786DB81" type="issue">
|
|||
|
|
<mods:number id="9A4BD12050A54270B708598A7BC384EB">1</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent id="C4A9E67DCDA595D883D5BCA5413BDF64" unit="page">
|
|||
|
|
<mods:start id="E2B575DF393D27A81A199A8B1468288A">143</mods:start>
|
|||
|
|
<mods:end id="5E0EA6876F3B24EC4AEDF555793ED822">152</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:classification id="BBB9619FAB8C02BEAA9C699C57E63CD2">journal article</mods:classification>
|
|||
|
|
<mods:identifier id="F8938A20BC1CCAB788BE8754A299E9C8" type="CLB-Dataset">21361</mods:identifier>
|
|||
|
|
<mods:identifier id="7D793307CF565F1BCD083EB20DCF3F06" type="DOI">10.11646/zootaxa.4751.1.8</mods:identifier>
|
|||
|
|
<mods:identifier id="F7FF01C94205A998C8442D2551421A35" type="GBIF-Dataset">d5e751b0-a468-493a-8e76-ef85b5150ce7</mods:identifier>
|
|||
|
|
<mods:identifier id="73B0E95190C16FA6CC1238D6FD71CA48" type="ISSN">1175-5326</mods:identifier>
|
|||
|
|
<mods:identifier id="C732F0093A3517E14768934EEEB7B44F" type="Zenodo-Dep">3974722</mods:identifier>
|
|||
|
|
<mods:identifier id="C7C2C1B1F0E103E338E9EE1DCC13B430" type="ZooBank">B79BA86F-4F61-4016-8AEE-0AE9110BA253</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment id="03B59B5DFFB4FFECFF26FE58FDBDF84E" ID-DOI="http://doi.org/10.5281/zenodo.4323840" ID-GBIF-Taxon="166097344" ID-Zenodo-Dep="4323840" LSID="urn:lsid:plazi:treatment:03B59B5DFFB4FFECFF26FE58FDBDF84E" httpUri="http://treatment.plazi.org/id/03B59B5DFFB4FFECFF26FE58FDBDF84E" lastPageId="7" lastPageNumber="150" pageId="2" pageNumber="145">
|
|||
|
|
<subSubSection id="C30679C0FFB4FFE9FF26FE58FF32FE00" box="[151,170,405,431]" pageId="2" pageNumber="145" type="nomenclature">
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF26FE58FF32FE00" blockId="2.[151,1437,405,540]" box="[151,170,405,431]" pageId="2" pageNumber="145">
|
|||
|
|
<heading id="D0EB9D27FFB4FFE9FF26FE58FF32FE00" box="[151,170,405,431]" level="2" pageId="2" pageNumber="145">
|
|||
|
|
<taxonomicName id="4C1C51C8FFB4FFE9FF26FE58FF32FE00" ID-CoL="5WTZH" authorityName="Steindachner" authorityYear="1898" box="[151,170,405,431]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="145" phylum="Chordata" rank="species" species="platei">A</taxonomicName>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30679C0FFB4FFE9FF00FE58FB17FE7C" pageId="2" pageNumber="145" type="materials_examined">
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF00FE58FB17FE7C" blockId="2.[151,1437,405,540]" pageId="2" pageNumber="145">
|
|||
|
|
total of five larvae of
|
|||
|
|
<taxonomicName id="4C1C51C8FFB4FFE9FE16FE58FDEAFE00" authorityName="Steindachner" authorityYear="1898" box="[423,626,405,431]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="145" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB4FFE9FE16FE58FDEAFE00" box="[423,626,405,431]" italics="true" pageId="2" pageNumber="145">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were obtained, ranging from 3.0 mm NL to
|
|||
|
|
<quantity id="4CE487AEFFB4FFE9FBD9FE58FB26FE00" box="[1128,1214,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="145" unit="mm" value="6.1">6.1 mm</quantity>
|
|||
|
|
SL (
|
|||
|
|
<tableCitation id="C69E1FF0FFB4FFE9FB45FE58FAD4FE00" box="[1268,1356,405,431]" captionStart="TABLE 2" captionStartId="1.[151,239,1369,1393]" captionText="TABLE 2. Morphometric measurements and meristic counts for five larval Callanthias platei, collected around Desventuradas Islands, Southeast Pacific." pageId="2" pageNumber="145">Table 2</tableCitation>
|
|||
|
|
). Standardised abundance was low, varying from 0.30 to 3.27 individuals per 1000
|
|||
|
|
<superScript id="7C698703FFB4FFE9FC49FE74FB83FE69" attach="left" box="[1016,1051,440,467]" fontSize="6" pageId="2" pageNumber="145">m-3</superScript>
|
|||
|
|
(
|
|||
|
|
<tableCitation id="C69E1FF0FFB4FFE9FB9BFE74FBE6FE7C" box="[1066,1150,441,467]" captionStart="TABLE 1" captionStartId="1.[151,239,658,682]" captionText="TABLE 1. Summary of the location and time of collection, type of gear and standardised abundance of larval Callanthias platei." pageId="2" pageNumber="145">Table 1</tableCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30679C0FFB4FFECFF76FE10FDBDF84E" lastPageId="7" lastPageNumber="150" pageId="2" pageNumber="145" type="description">
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF76FE10FA1FFDB4" blockId="2.[151,1437,405,540]" pageId="2" pageNumber="145">
|
|||
|
|
We describe three larvae that represent preflexion, flexion, and postflexion stages. The body of the larvae (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FADEFE10FF3BFDB4" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1970]" captionTargetBox="[182,1404,184,1913]" captionTargetId="figure@3.[174,1414,179,1917]" captionTargetPageId="3" captionText="FIGURE 1. Larval development of Callanthias platei, lateral view. A. Preflexion larva, 3 mm SL (MNHNCL ICT 7605). B. Flexion larva, 4.48 mm SL (MNHNCL ICT 7606). C. Postflexion larva, 6.09 mm SL (MNHNCL ICT 7603)." figureDoi="http://doi.org/10.5281/zenodo.3974725" httpUri="https://zenodo.org/record/3974725/files/figure.png" pageId="2" pageNumber="145">Fig. 1</figureCitation>
|
|||
|
|
) is deep at all stages (BD 48-54%), with a large head (HL 40-50%), and a long (PAL 60-69%) and a coiled gut.
|
|||
|
|
</paragraph>
|
|||
|
|
<subSection id="E29333A0FFB4FFE9FF26FD84FD8AFC94" pageId="2" pageNumber="145" type="multiple">
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF26FD84FEABFDCC" blockId="2.[151,1437,585,827]" box="[151,307,585,611]" pageId="2" pageNumber="145">
|
|||
|
|
<heading id="D0EB9D27FFB4FFE9FF26FD84FEABFDCC" bold="true" box="[151,307,585,611]" fontSize="11" level="1" pageId="2" pageNumber="145" reason="1">
|
|||
|
|
<emphasis id="B968F659FFB4FFE9FF26FD84FEABFDCC" bold="true" box="[151,307,585,611]" italics="true" pageId="2" pageNumber="145">Pigmentation</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF76FDA0FD8AFC94" blockId="2.[151,1437,585,827]" pageId="2" pageNumber="145">The preflexion larva has a few internal melanophores on the neurocranium and along the base of the dorsal fin. A conspicuous melanophore marks the symphysis of the lower jaw, and is present in all specimens. At flexion, there are external stellated melanophores in the dorsal area of the neurocranium, along the base of the dorsal fin, and an internal dendritic melanophore in the posterior margin of the gut, near the anus. In the postflexion stage, the number of melanophores in the dorsal margin of the neurocranium has increased, and a few long, stellated melanophores are present in the base of the anal fin.</paragraph>
|
|||
|
|
</subSection>
|
|||
|
|
<subSection id="E29333A0FFB4FFECFF26FCA5FDBDF84E" lastPageId="7" lastPageNumber="150" pageId="2" pageNumber="145" type="multiple">
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF26FCA5FE26FC2C" blockId="2.[151,1437,872,1979]" box="[151,446,872,899]" pageId="2" pageNumber="145">
|
|||
|
|
<heading id="D0EB9D27FFB4FFE9FF26FCA5FE26FC2C" bold="true" box="[151,446,872,899]" fontSize="11" level="1" pageId="2" pageNumber="145" reason="1">
|
|||
|
|
<emphasis id="B968F659FFB4FFE9FF26FCA5FE26FC2C" bold="true" box="[151,446,872,899]" italics="true" pageId="2" pageNumber="145">Osteological development</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF76FC43FCD8FA30" blockId="2.[151,1437,872,1979]" pageId="2" pageNumber="145">
|
|||
|
|
<emphasis id="B968F659FFB4FFE9FF76FC43FEEBFC08" box="[199,371,910,935]" italics="true" pageId="2" pageNumber="145">Neurocranium:</emphasis>
|
|||
|
|
The occipital region, the posterior portion of the neurocranium, is closed dorsally already in our smallest specimens (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FE1AFC7CFDBEFC64" box="[427,550,945,971]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A, B</figureCitation>
|
|||
|
|
). It forms a small crest that is later covered by the dome-shaped supraoccipital (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FF11FC18FE81FC40" box="[160,281,981,1007]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C, D</figureCitation>
|
|||
|
|
). The basioccipital has a caudad oriented projection (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FCD6FC18FC27FC40" box="[871,959,981,1007]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
) that approaches the first vertebra in larger specimens (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FEADFC34FEEFFBBC" box="[284,375,1017,1043]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2D</figureCitation>
|
|||
|
|
), and, since dissections are not possible, we are not able to interpret or relate this structure to anything else. The taenia marginalis (anterior and posterior) is connected via an epiphysal bridge to its counterpart (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FF11FB8CFF61FBF4" box="[160,249,1089,1115]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
). Anteriorly, it fuses to the lamina orbitonasalis and is covered by the frontal (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FBE7FB8CFB33FBF4" box="[1110,1195,1089,1115]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
|
|||
|
|
) laterally. The frontal appears as a small ossification lateral to the taenia marginalis, above the orbit (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FBB2FBA8FBC3FBD0" box="[1027,1115,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
|
|||
|
|
). It spreads medially to meet its counterpart in the midline in the larger stages (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FD72FB44FCA6FB0C" box="[707,830,1161,1187]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B, D</figureCitation>
|
|||
|
|
). The parietal starts as a small and narrow ossification posterior to the frontal and in the largest stage it covers the dorsal part of the neurocranium between the frontal anteriorly, the supraoccipital and epioccipital posteriorly, and the pterotic and autosphenotic laterally (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FAA5FB1CFA08FB44" box="[1300,1424,1233,1259]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B, D</figureCitation>
|
|||
|
|
). The prootic and the pterotic form the facet for the articulation with the dorsal head of hyomandibula (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FAB5FB38FAC6FAA0" box="[1284,1374,1269,1295]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2D</figureCitation>
|
|||
|
|
). The ethmoid region is already well developed with a large internasal septum in the smallest specimen (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FAB1FAD4FAC4FA9C" box="[1280,1372,1305,1331]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
). The connection between the ethmoid plate and the otic capsule via the trabeculae is already absorbed (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FB6CFAF0FAAFFAF8" box="[1245,1335,1341,1367]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
). A welldefined lamina orbitonasalis bears a small lateral ethmoid. The posterior edge of the ethmoid cartilage articulates with the pars autopalatina of the palatoquadrate (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FD0BFA48FCABFA30" box="[698,819,1413,1439]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B, C</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB4FFE9FF76FA67FC43F814" blockId="2.[151,1437,872,1979]" pageId="2" pageNumber="145">
|
|||
|
|
<emphasis id="B968F659FFB4FFE9FF76FA67FD2EFA6C" box="[199,694,1449,1475]" italics="true" pageId="2" pageNumber="145">Jaws, suspensorium and dorsal hyoid arch:</emphasis>
|
|||
|
|
All elements of the upper jaw are already present in the smallest stage available to us (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FE25FA00FE6AFA48" box="[404,498,1485,1511]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
). The premaxilla has a well-developed ascending process that articulates in older stages with a large rostral cartilage (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FD9AFA3CFD3EF9A4" box="[555,678,1521,1547]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B, D</figureCitation>
|
|||
|
|
). Initially, the maxilla is a simple leaf-shaped element that develops its complex anterior end for the articulation with the premaxilla and the autopalatine in the early postflexion stages (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FF11F9F4FE80F9FC" box="[160,280,1593,1619]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C, D</figureCitation>
|
|||
|
|
). The quadrate is the first element to ossify in the palatoquadrate (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FC5DF9F4FBDDF9FC" box="[1004,1093,1593,1619]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
), that develops the posteroventral process in a later stage (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FE7DF990FDDEF9D8" box="[460,582,1629,1655]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C, D</figureCitation>
|
|||
|
|
). Following the ossification of the quadrate in sequence is the metapterygoid (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FF11F94CFF6FF934" box="[160,247,1665,1691]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
|
|||
|
|
) and then the autopalatine (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FD84F94CFD2BF934" box="[565,691,1665,1691]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C, D</figureCitation>
|
|||
|
|
). The metapterygoid develops a lamina of membrane bone which contacts a membranous ossification of the hyomandibula (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FC89F968FC0CF910" box="[824,916,1701,1727]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C</figureCitation>
|
|||
|
|
). The autopalatine articulates with the lateral ethmoid of the neurocranium early on, and the maxilla in later stages (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FC05F904FBA9F94C" box="[948,1073,1737,1763]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B, D</figureCitation>
|
|||
|
|
). In the flexion stage larvae, the hyomandibula and the symplectic are present as separate ossifications of the hyosymplectic cartilage (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FAB7F920FAC5F8A8" box="[1286,1373,1773,1799]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
|
|||
|
|
). The hyomandibula develops a crest of membrane bone that attaches to the metapterygoid (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FBC3F8DCFB55F884" box="[1138,1229,1809,1835]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2C</figureCitation>
|
|||
|
|
). The dentary and the anguloarticular are already present in the smallest stage (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FCF0F8F8FC04F8E0" box="[833,924,1845,1871]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
). The dentary has the typical forked posterior end and encloses the anguloarticular (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FD8FF894FD19F8DC" box="[574,641,1881,1907]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2</figureCitation>
|
|||
|
|
A-D). Both elements are laterally attached to Meckel’s cartilage. In the smallest specimen the coronid process of the anguloarticular is already well developed (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FBDBF8B0FB5CF838" box="[1130,1220,1917,1943]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
), and the most posterior end of Meckel’s cartilage has a well ossified retroarticular (
|
|||
|
|
<figureCitation id="132736CEFFB4FFE9FCC5F86CFC56F814" box="[884,974,1953,1979]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="2" pageNumber="145">Fig. 2A</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF637AC3FFB5FFE8FF26F854FB7EF879" ID-DOI="http://doi.org/10.5281/zenodo.3974725" ID-Zenodo-Dep="3974725" httpUri="https://zenodo.org/record/3974725/files/figure.png" pageId="3" pageNumber="146" startId="3.[151,250,1945,1970]" targetBox="[182,1404,184,1913]" targetPageId="3">
|
|||
|
|
<paragraph id="8BA32A4BFFB5FFE8FF26F854FB7EF879" blockId="3.[151,1436,1945,2006]" pageId="3" pageNumber="146">
|
|||
|
|
<emphasis id="B968F659FFB5FFE8FF26F854FE8CF81D" bold="true" box="[151,276,1945,1970]" pageId="3" pageNumber="146">FIGURE 1.</emphasis>
|
|||
|
|
Larval development of
|
|||
|
|
<taxonomicName id="4C1C51C8FFB5FFE8FDBCF857FD5DF81D" authorityName="Steindachner" authorityYear="1898" box="[525,709,1946,1970]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="146" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB5FFE8FDBCF857FD5DF81D" box="[525,709,1946,1970]" italics="true" pageId="3" pageNumber="146">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lateral view. A. Preflexion larva, 3 mm SL (MNHNCL ICT 7605). B. Flexion larva, 4.48 mm SL (MNHNCL ICT 7606). C. Postflexion larva, 6.09 mm SL (MNHNCL ICT 7603).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF637AC3FFB2FFEFFF26F9CBFC20F8B5" ID-DOI="http://doi.org/10.5281/zenodo.3974727" ID-Zenodo-Dep="3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="4" pageNumber="147" startId="4.[151,250,1542,1567]" targetBox="[159,1429,189,1510]" targetPageId="4">
|
|||
|
|
<paragraph id="8BA32A4BFFB2FFEFFF26F9CBFC20F8B5" blockId="4.[151,1437,1542,1819]" pageId="4" pageNumber="147">
|
|||
|
|
<emphasis id="B968F659FFB2FFEFFF26F9CBFE96F9B0" bold="true" box="[151,270,1542,1567]" pageId="4" pageNumber="147">FIGURE 2</emphasis>
|
|||
|
|
. Osteological development of the skull in
|
|||
|
|
<taxonomicName id="4C1C51C8FFB2FFEFFD77F9CAFCE6F9B0" authorityName="Steindachner" authorityYear="1898" box="[710,894,1543,1567]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="147" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB2FFEFFD77F9CAFCE6F9B0" box="[710,894,1543,1567]" italics="true" pageId="4" pageNumber="147">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA32A4BFFB2FFEEFF76F884FC08FEB0" blockId="4.[151,1437,1865,2035]" lastBlockId="5.[151,1437,153,467]" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147">
|
|||
|
|
<emphasis id="B968F659FFB2FFEFFF76F884FE14F8CC" box="[199,396,1865,1891]" italics="true" pageId="4" pageNumber="147">
|
|||
|
|
Opercular
|
|||
|
|
<taxonomicName id="4C1C51C8FFB2FFEFFEF0F887FE1DF8CC" box="[321,389,1866,1891]" family="Series" pageId="4" pageNumber="151" rank="family">series</taxonomicName>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
We were only able to trace the preopercle and interopercle in the smallest specimen, the opercle might be present but could not be observed (
|
|||
|
|
<figureCitation id="132736CEFFB2FFEFFD1FF8A0FC90F828" box="[686,776,1901,1927]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="4" pageNumber="147">Fig. 2A</figureCitation>
|
|||
|
|
). We suspect that the other elements are present as well but did not stain well. However, all elements are obvious in the flexion larva (
|
|||
|
|
<figureCitation id="132736CEFFB2FFEFFC69F85CFBA8F804" box="[984,1072,1937,1963]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="4" pageNumber="147">Fig. 2B</figureCitation>
|
|||
|
|
), and bear prominent spines and serrations already (
|
|||
|
|
<figureCitation id="132736CEFFB2FFEFFEDDF878FE7FF860" box="[364,487,1973,1999]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="4" pageNumber="147">Fig. 2A, D</figureCitation>
|
|||
|
|
). The opercle has a single spine as a continuation of a ridge on its dorsal margin (
|
|||
|
|
<figureCitation id="132736CEFFB2FFEFFADEF878FF47F85C" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="4" pageNumber="147">Fig. 2A, D</figureCitation>
|
|||
|
|
). There were no signs of a second opercular spine, as reported for adults. The preopercle has three prominent spines in its anterior margin, that are about equal in size and ten spines along the posterior margin of which spines 3, 4 and 5 –counting from the ventral margin- are the largest. The interopercle has six spines and the subopercle four. The dorsal spines on the preopercle appear more as a serration in older stages (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFBBFFF2CFBF1FF54" box="[1038,1129,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="5" pageNumber="148">Fig. 2D</figureCitation>
|
|||
|
|
). The sub- and interopercle are partially covered by the opercle and the preopercle (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFCB9FEC8FC1BFEB0" box="[776,899,261,287]" captionStart="FIGURE 2" captionStartId="4.[151,250,1542,1567]" captionTargetBox="[159,1429,189,1510]" captionTargetId="figure@4.[151,1435,181,1516]" captionTargetPageId="4" captionText="FIGURE 2. Osteological development of the skull in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 MM SL. Cartilage is shown in grey, bone in white. ang-ar: anguloarticular; asph: autosphenotic; boc: basioccipital; d: dentary; ec: ethmoid cartilage; ecp: ectopterygoid; enp: entopterygoid; ep: ethmoid plate; epo: epiotic; epot: epiotic; et: epiphysial tectum; exo: exoccipital; f: frontal; h: hyomandibula; hsc: hyosympletic cartilage; iop: interopercle; is: internasal septum; let: lateral ethmoid; lon: lamina orbitonasalis; Mc: Meckel’s cartilage; mpt: metapterygoid; mx: maxilla; oc: otic capsule; op: opercle; p: parietal; pas: parasphenoid; pl: palatine; pmx: premaxilla; pop: preopercular; pro: prootic; pto: pterotic; q: quadrate; rar: retroarticular; rc: rostral cartilage; s: sympletic; soc: supraoccipital; sop: subopercle; tma: taenia marginalis anterior; tmp: taenia marginalis posterior; ttm: taenia tecti medialis; vo: vomer." figureDoi="http://doi.org/10.5281/zenodo.3974727" httpUri="https://zenodo.org/record/3974727/files/figure.png" pageId="5" pageNumber="148">Fig. 2B, D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB3FFEEFF76FEE4FD49FE7C" blockId="5.[151,1437,153,467]" pageId="5" pageNumber="148">
|
|||
|
|
<emphasis id="B968F659FFB3FFEEFF76FEE4FE0BFEEC" box="[199,403,297,323]" italics="true" pageId="5" pageNumber="148">Vertebral column:</emphasis>
|
|||
|
|
All 22 haemal arches and their associated spines are present in the smallest specimen (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFADEFEE4FF47FEC8" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3A, D</figureCitation>
|
|||
|
|
). They are preformed in cartilage (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFDEEFE80FD2EFEC8" box="[607,694,333,359]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3A</figureCitation>
|
|||
|
|
) and ossify in an anteroposterior direction. In the flexion larva, the six anteriormost vertebrae are ossified (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFDE3FEBCFD32FE24" box="[594,682,369,395]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3B</figureCitation>
|
|||
|
|
). The neural arches and their spines are preformed in cartilage, and the first fully formed neural arch appears on vertebra 11 (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFCA0FE58FCF0FE00" box="[785,872,405,431]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3B</figureCitation>
|
|||
|
|
). The centra and their associated arches and spines are fully ossified in the largest specimen (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFDDAFE74FD5DFE7C" box="[619,709,441,467]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF637AC3FFB3FFEEFF26F912FE94F803" ID-DOI="http://doi.org/10.5281/zenodo.3974729" ID-Zenodo-Dep="3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148" startId="5.[151,250,1759,1784]" targetBox="[173,1413,497,1726]" targetPageId="5">
|
|||
|
|
<paragraph id="8BA32A4BFFB3FFEEFF26F912FE94F803" blockId="5.[151,1437,1759,1964]" pageId="5" pageNumber="148">
|
|||
|
|
<emphasis id="B968F659FFB3FFEEFF26F912FE88F957" bold="true" box="[151,272,1759,1784]" pageId="5" pageNumber="148">FIGURE 3</emphasis>
|
|||
|
|
. Osteological development of vertebral column, dorsal, anal and pelvic fins of
|
|||
|
|
<taxonomicName id="4C1C51C8FFB3FFEEFBE4F92DFA97F957" authorityName="Steindachner" authorityYear="1898" box="[1109,1295,1760,1784]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB3FFEEFBE4F92DFA97F957" box="[1109,1295,1760,1784]" italics="true" pageId="5" pageNumber="148">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA32A4BFFB3FFEDFF76F816FE2AFF78" blockId="5.[199,1436,2010,2037]" lastBlockId="6.[151,1437,153,899]" lastPageId="6" lastPageNumber="149" pageId="5" pageNumber="148">
|
|||
|
|
Two cartilaginous supraneurals are present, and ossify perichondrally in the larger stages (
|
|||
|
|
<figureCitation id="132736CEFFB3FFEEFB01F817FAB3F85A" box="[1200,1323,2010,2037]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="5" pageNumber="148">Fig. 3A, D</figureCitation>
|
|||
|
|
). Both are obliquely oriented with the distal tip pointing dorsoanterad (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFC86FF54FC2CFF1C" box="[823,948,153,179]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3A, D</figureCitation>
|
|||
|
|
). The proximal ends are associated with the second interneural space.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB0FFEDFF76FF2CFCFFFDB4" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="149">
|
|||
|
|
<emphasis id="B968F659FFB0FFEDFF76FF2CFE2AFF54" box="[199,434,225,251]" italics="true" pageId="6" pageNumber="149">Dorsal and anal fins</emphasis>
|
|||
|
|
: The full complement of dorsal and anal fin pterygiophores, spines and rays are present in the largest specimen (D: XI + 11; A: III + 11;
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFD0BFEC8FC8AFEB0" box="[698,786,261,287]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3C</figureCitation>
|
|||
|
|
). The proximal-middle radials as well as the distal radials appear to develop in a caudad direction (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFDE9FEE4FD4BFEEC" box="[600,723,297,323]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3A, D</figureCitation>
|
|||
|
|
). The first pterygiophore in the dorsal fin is the largest and bears two spines, a serially associated and a smaller supernumerary spine anterior to it (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFBF2FE80FB5AFEC8" box="[1091,1218,333,359]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3B, D</figureCitation>
|
|||
|
|
). The cartilaginous precursor of the last pterygiophore has a small extension (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFC93FEBCFC06FE24" box="[802,926,369,395]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3C, D</figureCitation>
|
|||
|
|
). The first pterygiophore in the anal fin is the largest and bears two supernumerary spines and a serially associated spine (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFC59FE58FBFBFE00" box="[1000,1123,405,431]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3C, D</figureCitation>
|
|||
|
|
). All pterygiophores consist of a distal and a proximal-middle radial. We are uncertain whether middle radials are present in the pterygiophores that are associated with the soft rays (10 to 21). Similar to the last pterygiophore in the dorsal fin, the last pterygiophore in the anal fin has a cartilaginous extension (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFD51FDCCFCC3FDB4" box="[736,859,513,539]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3C, D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB0FFEDFF76FDE8FEC5FC2C" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="149">
|
|||
|
|
<emphasis id="B968F659FFB0FFEDFF76FDE8FE4EFD90" box="[199,470,549,575]" italics="true" pageId="6" pageNumber="149">Pectoral fin and girdle:</emphasis>
|
|||
|
|
We were not able to trace any dermal elements in the smallest larva (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFB4EFDE8FAC4FD90" box="[1279,1372,549,575]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4A</figureCitation>
|
|||
|
|
). The endoskeletal scapulocoracoid and the pectoral plate are present (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFCD6FD84FC53FDCC" box="[871,971,585,611]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Figs. 3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFC66FD84FC65FDCC" box="[983,1021,585,611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">4A</figureCitation>
|
|||
|
|
). The scapulocoracoid has a foramen dorsally. The pectoral plate has an indentation dorsally indicating the separation of the most dorsal radial (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFA8EFDA0FA0FFD28" box="[1343,1431,621,647]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="6" pageNumber="149">Fig. 3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFF26FD5CFF26FD04" box="[151,190,657,683]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">4A</figureCitation>
|
|||
|
|
). The separation of the radials and the development of the distal radials follow a ventral direction (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFAB7FD5CFAC6FD04" box="[1286,1374,657,683]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4B</figureCitation>
|
|||
|
|
). The first distal radial is associated with the scapulocoracoid (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFCA1FD78FC11FD60" box="[784,905,693,719]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4B, C</figureCitation>
|
|||
|
|
). The posttemporal develops as a small pointed element (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFF4EFD14FECDFD5C" box="[255,341,729,755]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4B</figureCitation>
|
|||
|
|
) that just develops its ventral arm in the early postflexion larva (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFB98FD14FB1BFD5C" box="[1065,1155,729,755]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4C</figureCitation>
|
|||
|
|
). The supracleithrum has a posterior oriented spine at its dorsal part, and the cleithrum is a long needle-shaped ossification that runs in an anteroventral direction (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFE19FCECFDBCFC94" box="[424,548,801,827]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4B, C</figureCitation>
|
|||
|
|
). The postcleithrum is a large element that articulates with the supracleithrum and becomes needle-shaped dorsoventrally (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFD36FC88FD78FCF0" box="[647,736,837,863]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4C</figureCitation>
|
|||
|
|
). In the largest specimens the full complement of 18 fin rays is present (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFF46FCA4FEC8FC2C" box="[247,336,873,899]" captionStart="FIGURE 4" captionStartId="6.[151,250,1655,1680]" captionTargetBox="[214,1370,927,1626]" captionTargetId="figure@6.[210,1377,921,1630]" captionTargetPageId="6" captionText="FIGURE 4. Osteological development of the pectoral fin in Callanthias platei, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum." figureDoi="http://doi.org/10.5281/zenodo.3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149">Fig. 4C</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF637AC3FFB0FFEDFF26F9BAFBD5F977" ID-DOI="http://doi.org/10.5281/zenodo.3974731" ID-Zenodo-Dep="3974731" httpUri="https://zenodo.org/record/3974731/files/figure.png" pageId="6" pageNumber="149" startId="6.[151,250,1655,1680]" targetBox="[214,1370,927,1626]" targetPageId="6">
|
|||
|
|
<paragraph id="8BA32A4BFFB0FFEDFF26F9BAFBD5F977" blockId="6.[151,1437,1655,1752]" pageId="6" pageNumber="149">
|
|||
|
|
<emphasis id="B968F659FFB0FFEDFF26F9BAFE96F93F" bold="true" box="[151,270,1655,1680]" pageId="6" pageNumber="149">FIGURE 4</emphasis>
|
|||
|
|
. Osteological development of the pectoral fin in
|
|||
|
|
<taxonomicName id="4C1C51C8FFB0FFEDFCB7F9B5FC25F93F" authorityName="Steindachner" authorityYear="1898" box="[774,957,1656,1680]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB0FFEDFCB7F9B5FC25F93F" box="[774,957,1656,1680]" italics="true" pageId="6" pageNumber="149">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lateral view. A. 3 mm SL. B. 4.48 mm SL. C. 6.09 mm SL. Grey areas, cartilage, open areas, bone. cl: cleithrum; dr: distal radials; pcl: postcleithrum; pp: pectoral plate; pr: pectoral rays; pt: post temporal; r: radials; scc: scapulocoracoid cartilage; scl: supracleithrum.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA32A4BFFB0FFECFF76F8CBFC8EFEB0" blockId="6.[151,1437,1798,2041]" lastBlockId="7.[151,1437,152,287]" lastPageId="7" lastPageNumber="150" pageId="6" pageNumber="149">
|
|||
|
|
<emphasis id="B968F659FFB0FFEDFF76F8CBFED1F88F" box="[199,329,1798,1824]" italics="true" pageId="6" pageNumber="149">Caudal fin:</emphasis>
|
|||
|
|
The elements of the caudal skeleton consist of a parhypural, five hypurals, and three epurals (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFADEF8CBFF3FF8EB" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig. 5</figureCitation>
|
|||
|
|
). In the smallest larva, only hypurals 1-3 are present as small, independent cartilaginous structures, as well as the parahypural (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFE9DF883FE1EF8C7" box="[300,390,1869,1896]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig. 5A</figureCitation>
|
|||
|
|
). In the flexion larva, hypural 4 is already visible (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFC0BF883FB94F8C7" box="[954,1036,1869,1896]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig 5B</figureCitation>
|
|||
|
|
). However, in older stages hypurals 1 and 2 and hypurals 3 and 4 fuse to a single plate, respectively (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFCDEF8BFFC72F822" box="[879,1002,1905,1933]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig. 5C, D</figureCitation>
|
|||
|
|
). The uroneurals are not present or did not pick up the alizarin stain in all our specimens. The first elements that are present in the smallest specimen are the parhypural and hypurals 1, 2 and 3. In the flexion-stage specimen, hypurals 1+2 and 3+4 are partially fused (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFADEF877FF23F857" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig. 5B</figureCitation>
|
|||
|
|
). A fifth, smaller hypural appears in the postflexion larva (
|
|||
|
|
<figureCitation id="132736CEFFB0FFEDFCFDF813FC5FF856" box="[844,967,2013,2041]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="6" pageNumber="149">Fig. 5C, D</figureCitation>
|
|||
|
|
). The three epurals are present dorsally to the notochord in the flexion larva (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFDA7FF54FDF5FF1C" box="[534,621,152,179]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150">Fig. 5B</figureCitation>
|
|||
|
|
). Preural centrum 2 and ural centrum have started to ossify in the younger postflexion stage (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFED6FF70FE58FF78" box="[359,448,188,215]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150">Fig. 5C</figureCitation>
|
|||
|
|
). The ural centrum has a small cartilaginous neural arch (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFBFAFF70FB16FF78" box="[1099,1166,188,215]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150">Fig. 5</figureCitation>
|
|||
|
|
B-D). The fin rays have started to form during the flexion of the notochord (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFD63FF2CFCB1FF54" box="[722,809,224,251]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150">Fig. 5B</figureCitation>
|
|||
|
|
), and the full complement of 17 principal caudal fin rays are present (9+8;
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFEEDFEC8FE2BFEB0" box="[348,435,260,287]" captionStart="FIGURE 5" captionStartId="7.[151,250,652,677]" captionTargetBox="[165,1423,310,621]" captionTargetId="figure@7.[163,1423,309,627]" captionTargetPageId="7" captionText="FIGURE 5. Osteological development of the caudal fin in Callanthias platei, lateral view.A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle." figureDoi="http://doi.org/10.5281/zenodo.3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150">Fig. 5D</figureCitation>
|
|||
|
|
) in the larger postflexion stage.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF637AC3FFB1FFECFF26FD41FD65FD42" ID-DOI="http://doi.org/10.5281/zenodo.3974733" ID-Zenodo-Dep="3974733" httpUri="https://zenodo.org/record/3974733/files/figure.png" pageId="7" pageNumber="150" startId="7.[151,250,652,677]" targetBox="[165,1423,310,621]" targetPageId="7">
|
|||
|
|
<paragraph id="8BA32A4BFFB1FFECFF26FD41FD65FD42" blockId="7.[151,1436,652,749]" pageId="7" pageNumber="150">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFF26FD41FE94FD0A" bold="true" box="[151,268,652,677]" pageId="7" pageNumber="150">FIGURE 5</emphasis>
|
|||
|
|
. Osteological development of the caudal fin in
|
|||
|
|
<taxonomicName id="4C1C51C8FFB1FFECFD54FD41FC02FD0B" authorityName="Steindachner" authorityYear="1898" box="[741,922,652,676]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFD54FD41FC02FD0B" box="[741,922,652,676]" italics="true" pageId="7" pageNumber="150">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lateral view. A. 3 mm SL. B. 4.48 MM SL. C. 5.31 mm SL. D. 6.09 mm SL. Cr: Caudal rays; ep: epural; Ha: Haemal arch; hy: hypural; Na: Neural arch; Nc: Notochord; P: Preural centrum 1; Uc: Ural centrum; ph: parahypural; Ur: Urostyle.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF637AC3FFB1FFECFF26F949FC53F932" box="[151,971,1668,1693]" pageId="7" pageNumber="150">
|
|||
|
|
<paragraph id="8BA32A4BFFB1FFECFF26F949FC53F932" blockId="7.[151,971,1668,1693]" box="[151,971,1668,1693]" pageId="7" pageNumber="150">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFF26F949FE95F932" bold="true" box="[151,269,1668,1693]" pageId="7" pageNumber="150">FIGURE 6</emphasis>
|
|||
|
|
. Illustration of the scales of a
|
|||
|
|
<taxonomicName id="4C1C51C8FFB1FFECFD8EF948FD6EF932" authorityName="Steindachner" authorityYear="1898" box="[575,758,1669,1693]" class="Actinopterygii" family="Callanthiidae" genus="Callanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="platei">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFD8EF948FD6EF932" box="[575,758,1669,1693]" italics="true" pageId="7" pageNumber="150">Callanthias platei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
larva (6.09 mm SL).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA32A4BFFB1FFECFF76F922FC62F8D9" blockId="7.[151,1436,1775,2018]" pageId="7" pageNumber="150">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFF76F922FEDEF8A5" box="[199,326,1775,1802]" italics="true" pageId="7" pageNumber="150">Pelvic fins:</emphasis>
|
|||
|
|
The pelvic element is formed by the flexion stage (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFC32F922FC41F8A5" box="[899,985,1775,1802]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="7" pageNumber="150">Fig. 3B</figureCitation>
|
|||
|
|
) and shows signs of perichondral ossification. The anterior process is in contact with the ventral part of the cleithrum (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFBB6F8DEFB19F881" box="[1031,1153,1811,1838]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="7" pageNumber="150">Fig. 3C, D</figureCitation>
|
|||
|
|
). The posterior process is short (not shown in figures), and the strong spine is the first element of the fin to form (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFBC3F8FAFB53F8FD" box="[1138,1227,1847,1874]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="7" pageNumber="150">Fig. 3B</figureCitation>
|
|||
|
|
). The full complement of fin elements (I+5) is present in the larger postflexion stage (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFC22F896FC75F8D9" box="[915,1005,1883,1910]" captionStart="FIGURE 3" captionStartId="5.[151,250,1759,1784]" captionTargetBox="[173,1413,497,1726]" captionTargetId="figure@5.[172,1415,489,1734]" captionTargetPageId="5" captionText="FIGURE 3. Osteological development of vertebral column, dorsal, anal and pelvic fins of Callanthias platei, lateral view. Red/pink structures, bone; blue structures, cartilage. A. 3 mm SL. B. 4.48 mm SL. C. 5.31 mm SL. D. 6.09 mm SL. adr: anal distal radial; apr: anal proximal radial; ap: anal pterygiophore; as: anal spine; asr: anal soft ray; ddr: dorsal distal radial; dmr: dorsal middle radial; dp: dorsal pterygiophore; dpr: dorsal proximal radial; ds: dorsal spine; dsr: dorsal soft ray; ha: haemal arch; hs: haemal spine; na: neural arch ns: neural spine; p: pelvis; pr: pelvic ray; ps: pelvic spine; snc: supraneural cartilages; v: vertebra." figureDoi="http://doi.org/10.5281/zenodo.3974729" httpUri="https://zenodo.org/record/3974729/files/figure.png" pageId="7" pageNumber="150">Fig. 3D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA32A4BFFB1FFECFF76F84DFDBDF84E" blockId="7.[151,1436,1775,2018]" pageId="7" pageNumber="150">
|
|||
|
|
<emphasis id="B968F659FFB1FFECFF76F84DFECEF835" box="[199,342,1920,1946]" italics="true" pageId="7" pageNumber="150">Squamation:</emphasis>
|
|||
|
|
In the two largest (5.31 and
|
|||
|
|
<quantity id="4CE487AEFFB1FFECFD25F8B2FD60F835" box="[660,760,1919,1946]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="150" unit="mm" value="6.1">6.10 mm</quantity>
|
|||
|
|
SL) collected individuals, the first signs of squamation were observed. Scales are cycloid, each ornamented with two concentric lines and one central spine (
|
|||
|
|
<figureCitation id="132736CEFFB1FFECFB55F86EFAB2F811" box="[1252,1322,1955,1982]" captionStart="FIGURE 6" captionStartId="7.[151,250,1668,1693]" captionText="FIGURE 6. Illustration of the scales of a Callanthias platei larva (6.09 mm SL)." pageId="7" pageNumber="150">Fig. 6</figureCitation>
|
|||
|
|
), forming rows along the midline of the body.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSection>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|