treatments-xml/data/23/D2/13/23D2138E2CA25FC495EE6B9D27E6DCB1.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.65620" ID-GBIF-Dataset="657dad35-aa3b-43ba-9ae9-847514dc7914" ID-PMC="PMC8280097" ID-Pensoft-Pub="1313-2970-1048-49" ID-Pensoft-UUID="3DFE13D63D095472B8823D29B6EA1750" ID-PubMed="34305418" ID-ZooBank="EC587B49FB7F4C3A9E9DD11DA6BF3E91" ModsDocID="1313-2970-1048-49" checkinTime="1625677518425" checkinUser="pensoft" docAuthor="Liu, Shuo, Zhang, Peisong &amp; Rao, Dingqi" docDate="2021" docId="23D2138E2CA25FC495EE6B9D27E6DCB1" docLanguage="en" docName="ZooKeys 1048: 49-67" docOrigin="ZooKeys 1048" docPubDate="2021-07-07" docSource="http://dx.doi.org/10.3897/zookeys.1048.65620" docTitle="Nanorana xuelinensis Liu &amp; Zhang &amp; Rao 2021, sp. nov." docType="treatment" docUuid="3BB0CC31-8B68-4EA7-BC7C-DDF7D78C977F" docUuidSource="ZooBank" docVersion="5" id="3DFE13D63D095472B8823D29B6EA1750" lastPageNumber="49" masterDocId="3DFE13D63D095472B8823D29B6EA1750" masterDocTitle="A new species of Nanorana Guenther, 1896 (Anura, Dicroglossidae) from Yunnan, China" masterLastPageNumber="67" masterPageNumber="49" pageNumber="49" updateTime="1668150586060" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Nanorana Guenther, 1896 (Anura, Dicroglossidae) from Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Shuo</mods:namePart>
<mods:affiliation>Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, the Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming, Yunnan 650223, China</mods:affiliation>
<mods:nameIdentifier type="email">liushuo@mail.kiz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Peisong</mods:namePart>
<mods:affiliation>Research Institute of Xishuangbanna National Nature Reserve, No. 6 North Galan Road, Jinghong, Yunnan 666100, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rao, Dingqi</mods:namePart>
<mods:affiliation>Kunming Institute of Zoology, the Chinese Academy of Sciences, No. 17 Longxin Road, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier type="email">raodq@mail.kiz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-07-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1048</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>49</mods:start>
<mods:end>67</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1048.65620</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.65620</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1048-49</mods:identifier>
<mods:identifier type="ZooBank">EC587B49FB7F4C3A9E9DD11DA6BF3E91</mods:identifier>
<mods:identifier type="Pensoft-UUID">3DFE13D63D095472B8823D29B6EA1750</mods:identifier>
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<treatment ID-GBIF-Taxon="183401815" LSID="urn:lsid:zoobank.org:act:3BB0CC31-8B68-4EA7-BC7C-DDF7D78C977F" httpUri="http://treatment.plazi.org/id/23D2138E2CA25FC495EE6B9D27E6DCB1" lastPageNumber="49" pageId="0" pageNumber="49">
<subSubSection pageId="0" pageNumber="49" type="nomenclature">
<paragraph pageId="0" pageNumber="49">
<taxonomicName LSID="http://zoobank.org/3BB0CC31-8B68-4EA7-BC7C-DDF7D78C977F" authority="Liu &amp; Zhang &amp; Rao, 2021" authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis" status="sp. nov.">Nanorana xuelinensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views of the specimens of the type series of Nanorana xuelinensis sp. nov. in preservative." figureDoi="10.3897/zookeys.1048.65620.figure2" httpUri="https://binary.pensoft.net/fig/563876" pageId="0" pageNumber="49">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Various views of the male holotype (KIZL 2019016) of Nanorana xuelinensis sp. nov. in preservative." figureDoi="10.3897/zookeys.1048.65620.figure3" httpUri="https://binary.pensoft.net/fig/563877" pageId="0" pageNumber="49">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Various views of the female paratype (KIZL 2019017) of Nanorana xuelinensis sp. nov. in preservative." figureDoi="10.3897/zookeys.1048.65620.figure4" httpUri="https://binary.pensoft.net/fig/563878" pageId="0" pageNumber="49">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Different views of the male holotype (KIZL 2019016) of Nanorana xuelinensis sp. nov. in life." figureDoi="10.3897/zookeys.1048.65620.figure5" httpUri="https://binary.pensoft.net/fig/563879" pageId="0" pageNumber="49">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Different views of the female paratype (KIZL 2019017) of Nanorana xuelinensis sp. nov. in life." figureDoi="10.3897/zookeys.1048.65620.figure6" httpUri="https://binary.pensoft.net/fig/563880" pageId="0" pageNumber="49">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="holotype">
<paragraph pageId="0" pageNumber="49">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="49">
KIZL2019016, adult male, collected on 13 July 2019 by Shuo Liu from Xuelin Township, Lancang County, Puer City, Yunnan Province, China (
<geoCoordinate degrees="23" direction="north" minutes="2" orientation="latitude" precision="15" seconds="38" value="23.043888">23°2'38&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="32" orientation="longitude" precision="15" seconds="35" value="99.54305">99°32'35&quot;E</geoCoordinate>
; at an elevation of 1840 m asl).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="paratypes">
<paragraph pageId="0" pageNumber="49">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="49">KIZL2019012 and KIZL2019015, two subadult males; KIZL2019013 and KIZL2019014, two subadult females; and KIZL2019017, adult female. All with same collection information as for the holotype.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure2" httpUri="https://binary.pensoft.net/fig/563876" pageId="0" pageNumber="49" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 2.</emphasis>
Dorsal and ventral views of the specimens of the type series of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. in preservative.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="diagnosis">
<paragraph pageId="0" pageNumber="49">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="49">Large body size, SVL 101.7-107.3 mm in adults; adult males with keratinized spines on chest, belly, lateral body, posterior dorsum, buttocks, outer side of the fore limbs, the inner metacarpal tubercle, fingers I and II, and upper eyelids; no spines on the inner side of the lower and upper arm; forelimbs strongly hypertrophied in adult males; tympanum big but indistinct, ca 2/3 of eye diameter; anterior dorsum skin smooth; dorsolateral folds absent; finger I longer than finger II; webbing deeply incurved between tips of toes; no tarsal fold; present outer metacarpal tubercle and absent outer metatarsal tubercle; vomerine teeth distinct.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure3" httpUri="https://binary.pensoft.net/fig/563877" pageId="0" pageNumber="49" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 3.</emphasis>
Various views of the male holotype (KIZL2019016) of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. in preservative.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="49">The living specimens were yellowish brown with distinct or indistinct black spots on the dorsum and sides of the body and the dorsal side of limbs; no band on arms and legs. Ventral surface white with no spots, throat yellow in adult males.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure4" httpUri="https://binary.pensoft.net/fig/563878" pageId="0" pageNumber="49" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 4.</emphasis>
Various views of the female paratype (KIZL2019017) of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. in preservative.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="49">
Adult male, habitus very stout, SVL 107.3 mm, large size in genus 
<taxonomicName authorityName="Gunther" authorityYear="1896" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana</emphasis>
</taxonomicName>
; head flat and broader than long (HL/HW 0.85, HH/HL 0.53); snout blunt and rounded in both dorsal and lateral views; canthus rostralis obtuse; tympanum large and very indistinct (TDH/EHD 0.76); supratympanic fold extending from eye over tympanum to shoulder; transversal fold behind eyes; eye relatively large (EHD/HL 0.26), pupil slightly rhombic; vomerine teeth distinct; tongue large and cordiform, deeply notched posteriorly.
</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure5" httpUri="https://binary.pensoft.net/fig/563879" pageId="0" pageNumber="49" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 5.</emphasis>
Different views of the male holotype (KIZL2019016) of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. in life.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="49">Forelimbs short and strongly hypertrophied (LAD 18.8 mm); relative finger length: II &lt;I &lt;IV &lt;III; inner metacarpal tubercle enlarged, dorsal surface of inner metacarpal tubercle, fingers I, and finger II with black keratinized nuptial spines, no spine on inner side of fore limbs, and a few spines on outer side of fore limbs; finger tips rounded but not dilated, fingers free, without webbing, no circum-marginal groove or lateroventral groove; subarticular tubercles distinct, outer metacarpal tubercle indistinct.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure6" httpUri="https://binary.pensoft.net/fig/563880" pageId="0" pageNumber="49" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 6.</emphasis>
Different views of the female paratype (KIZL2019017) of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. in life.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="49">Hindlimbs rather long and stout; relative toe length: I &lt;II &lt;V &lt;III &lt;IV; tips of toes rounded but not dilated; subarticular tubercles oval and distinct, formula is 1, 1, 2, 3, 2; inner metatarsal tubercles elongated and pronounced; outer metatarsal tubercle absent; webbing deeply incurved between tips of toes, formula I 0-0- II 0-0- III 0--0- IV 0--0 V; lateral fringe on the outer side of toe V developed; no circum-marginal groove or lateroventral groove; tarsal fold absent.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure7" httpUri="https://binary.pensoft.net/fig/563881" pageId="0" pageNumber="49" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 7.</emphasis>
Habitat of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. at the type locality.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="49">Anterior dorsum skin smooth; keratinized spines present on chest, belly, lateral body, posterior dorsum, buttocks, and upper eyelids; spines most dense on axilla and each side of chest.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="coloration of holotype in life">
<paragraph pageId="0" pageNumber="49">Coloration of holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="49">The coloration of dorsum is yellowish brown with very indistinct black spots in dorsum, and no band on arms and legs. Ventral surface white with no spots. The throat is yellow. The pupil is black, and the iris is light yellow with many black radial strips around the pupil.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="sexual dimorphism">
<paragraph pageId="0" pageNumber="49">Sexual dimorphism.</paragraph>
<paragraph pageId="0" pageNumber="49">The forelimbs of adult males are strongly hypertrophied; in addition, adult males have keratinized spines on chest, belly, lateral body, posterior dorsum, buttocks, outer side of the fore limbs, the inner metacarpal tubercle, fingers I and II, and upper eyelids. The forelimbs of adult females are not hypertrophied, and adult females have distinct black spots on the dorsum, lateral body, and the dorsal side of limbs, no keratinized spines on chest, belly, lateral body, posterior dorsum, buttocks, and upper eyelids, and only some keratinized spines on finger I and a few small spines on finger II.</paragraph>
<caption doi="10.3897/zookeys.1048.65620.figure8" httpUri="https://binary.pensoft.net/fig/563882" pageId="0" pageNumber="49" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 8.</emphasis>
Map showing the type locality of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. (red dot).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="etymology">
<paragraph pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="49">
The name refers to Xuelin Township, the locality where the new species was found. We propose &quot;Xuelin 
<taxonomicName authorityName="Dubois" authorityYear="1975" class="Amphibia" family="Dicroglossidae" genus="Paa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paa" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="genus">Paa</taxonomicName>
Frog&quot; or &quot;Xuelin Spiny Frog&quot; for the common English name and 
<normalizedToken originalValue="“雪林棘蛙”">&quot;雪林棘蛙&quot;</normalizedToken>
(
<normalizedToken originalValue="Xuě">Xue</normalizedToken>
<normalizedToken originalValue="Lín">Lin</normalizedToken>
<normalizedToken originalValue="Jí">Ji</normalizedToken>
<normalizedToken originalValue="Wā">Wa</normalizedToken>
) for the common Chinese name of the new species.
</paragraph>
<caption ID-Table-UUID="C482830524CC70107C26314E773CD014" httpUri="http://table.plazi.org/id/C482830524CC70107C26314E773CD014" pageId="0" pageNumber="49" start="Table 4" startId="T4">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Table 4.</emphasis>
Morphological measurements (mm) of the type series of 
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="49">
<table pageId="0" pageNumber="49">
<tr pageId="0" pageNumber="49">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="49" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019016 Holotype Adult male</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019017 Paratype Adult female</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019012 Paratype Subadult male</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019013 Paratype Subadult female</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019014 Paratype Subadult female</th>
<th colspan="1" pageId="0" pageNumber="49" rowspan="1">KIZL2019015 Paratype Subadult male</th>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">SVL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">107.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">101.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">60.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">79.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">75.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">66.9</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">AG</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">36.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">40.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">15.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">29.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">29.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">20.1</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">HL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">35.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">36.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">23.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">27.0</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">27.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">25.8</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">HW</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">42.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">38.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">23.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">28.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">28.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">27.0</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">HH</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">19.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">18.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">14.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">15.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">13.1</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">SL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">16.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">14.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">9.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">11.2</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">ID</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.5</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">IOD</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">3.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">3.3</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">UEW</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.1</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">EHD</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">9.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">6.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.4</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">TDH</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.6</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">TDV</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">6.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">3.6</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">SND</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">6.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.9</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">END</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">4.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">5.4</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">LAl</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">22.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">19.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">15.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">13.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">14.1</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">LAD</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">18.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">10.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">7.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">8.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">9.4</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">HAL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">22.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">19.0</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">14.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">17.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">15.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">15.6</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">FML</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">45.4</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">41.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">26.8</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">34.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">33.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">29.5</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">TIL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">42.5</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">39.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">25.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">33.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">31.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">28.9</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">TFL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">65.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">63.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">41.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">54.2</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">49.9</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">45.8</td>
</tr>
<tr pageId="0" pageNumber="49">
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">FL</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">44.3</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">42.7</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">29.1</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">36.0</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">33.6</td>
<td colspan="1" pageId="0" pageNumber="49" rowspan="1">33.1</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="distribution">
<paragraph pageId="0" pageNumber="49">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. is recorded in Lancang County (
<normalizedToken originalValue="Pu’er">Pu'er</normalizedToken>
City), Shuangjiang County (Lincang City), and Jinghong City (Xishuangbanna Prefecture), Yunnan Province, China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="habitat">
<paragraph pageId="0" pageNumber="49">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="49">
The type series was found in a still-water pond. At the type locality we found three other species of amphibians: 
<taxonomicName authorityName="Boulenger" authorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Chirixalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chirixalus doriae" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="doriae">Chirixalus cf. doriae</taxonomicName>
Boulenger, 1893; 
<taxonomicName authorityName="Jiang, Ren, Guo &amp; L.i.Wang" authorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Raorchestes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raorchestes hillisi" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="hillisi">
<emphasis italics="true" pageId="0" pageNumber="49">Raorchestes hillisi</emphasis>
</taxonomicName>
Jiang Ren, Guo, Wang &amp; Li, 2020; 
<taxonomicName class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tylototriton verrucosus" order="Caudata" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="verrucosus">
<emphasis italics="true" pageId="0" pageNumber="49">Tylototriton verrucosus</emphasis>
</taxonomicName>
Anderson, 1871a; and three species of reptiles: 
<taxonomicName class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Calotes emma" order="Squamata" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="emma">
<emphasis italics="true" pageId="0" pageNumber="49">Calotes emma</emphasis>
</taxonomicName>
Gray, 1845; 
<taxonomicName authorityName="Liu &amp; Rao" authorityYear="2021" class="Amphibia" family="Pareatidae" genus="Pareas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareas xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Pareas xuelinensis</emphasis>
</taxonomicName>
Liu &amp; Rao, 2021; and 
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1888" class="Reptilia" family="Agamidae" genus="Pseudocalotes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pseudocalotes microlepis" order="Squamata" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="microlepis">
<emphasis italics="true" pageId="0" pageNumber="49">Pseudocalotes microlepis</emphasis>
</taxonomicName>
(Boulenger, 1887).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="comparisons">
<paragraph pageId="0" pageNumber="49">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. aenea" pageId="0" pageNumber="49" rank="species" species="aenea">
<emphasis italics="true" pageId="0" pageNumber="49">N. aenea</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="N. annandalii" pageId="0" pageNumber="49" rank="species" species="annandalii">
<emphasis italics="true" pageId="0" pageNumber="49">N. annandalii</emphasis>
</taxonomicName>
(Boulenger, 1920), 
<taxonomicName lsidName="N. gammii" pageId="0" pageNumber="49" rank="species" species="gammii">
<emphasis italics="true" pageId="0" pageNumber="49">N. gammii</emphasis>
</taxonomicName>
(Anderson, 1871b), 
<taxonomicName lsidName="N. liebigii" pageId="0" pageNumber="49" rank="species" species="liebigii">
<emphasis italics="true" pageId="0" pageNumber="49">N. liebigii</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
, 1860), 
<taxonomicName lsidName="N. polunini" pageId="0" pageNumber="49" rank="species" species="polunini">
<emphasis italics="true" pageId="0" pageNumber="49">N. polunini</emphasis>
</taxonomicName>
(Smith, 1951), 
<taxonomicName lsidName="N. rarica" pageId="0" pageNumber="49" rank="species" species="rarica">
<emphasis italics="true" pageId="0" pageNumber="49">N. rarica</emphasis>
</taxonomicName>
(Dubois, Matsui &amp; Ohler, 2001), 
<taxonomicName lsidName="N. rostandi" pageId="0" pageNumber="49" rank="species" species="rostandi">
<emphasis italics="true" pageId="0" pageNumber="49">N. rostandi</emphasis>
</taxonomicName>
(Dubois, 1974), and 
<taxonomicName lsidName="N. unculuanus" pageId="0" pageNumber="49" rank="species" species="unculuanus">
<emphasis italics="true" pageId="0" pageNumber="49">N. unculuanus</emphasis>
</taxonomicName>
by the absence of dorsolateral fold (vs presence).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. arnoldi" pageId="0" pageNumber="49" rank="species" species="arnoldi">
<emphasis italics="true" pageId="0" pageNumber="49">N. arnoldi</emphasis>
</taxonomicName>
(Dubois, 1975), 
<taxonomicName lsidName="N. maculosa" pageId="0" pageNumber="49" rank="species" species="maculosa">
<emphasis italics="true" pageId="0" pageNumber="49">N. maculosa</emphasis>
</taxonomicName>
(Liu, Hu &amp; Yang, 1960), 
<taxonomicName lsidName="N. yunnanensis" pageId="0" pageNumber="49" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. yunnanensis</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="N. zhaoermii" pageId="0" pageNumber="49" rank="species" species="zhaoermii">
<emphasis italics="true" pageId="0" pageNumber="49">N. zhaoermii</emphasis>
</taxonomicName>
Qi, Zhou, Lu &amp; Li, 2019 by the spines present only on finger I and finger II in adult males (vs present on finger I-III).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. arunachalensis" pageId="0" pageNumber="49" rank="species" species="arunachalensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. arunachalensis</emphasis>
</taxonomicName>
(Saikia, Sinha &amp; Kharkongor, 2017), 
<taxonomicName lsidName="N. blanfordii" pageId="0" pageNumber="49" rank="species" species="blanfordii">
<emphasis italics="true" pageId="0" pageNumber="49">N. blanfordii</emphasis>
</taxonomicName>
(Boulenger, 1882), 
<taxonomicName lsidName="N. chayuensis" pageId="0" pageNumber="49" rank="species" species="chayuensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. chayuensis</emphasis>
</taxonomicName>
(Ye, 1977), 
<taxonomicName lsidName="N. conaensis" pageId="0" pageNumber="49" rank="species" species="conaensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. conaensis</emphasis>
</taxonomicName>
(Fei &amp; Huang, 1981), 
<taxonomicName lsidName="N. minica" pageId="0" pageNumber="49" rank="species" species="minica">
<emphasis italics="true" pageId="0" pageNumber="49">N. minica</emphasis>
</taxonomicName>
(Dubois, 1975), and 
<taxonomicName lsidName="N. mokokchungensis" pageId="0" pageNumber="49" rank="species" species="mokokchungensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. mokokchungensis</emphasis>
</taxonomicName>
(Das &amp; Chanda, 2000) by its larger body size.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. feae" pageId="0" pageNumber="49" rank="species" species="feae">
<emphasis italics="true" pageId="0" pageNumber="49">N. feae</emphasis>
</taxonomicName>
(Boulenger, 1887) by the absence of spines on the inner side of the fore limbs in adult males (vs. presence).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. kangxianensis" pageId="0" pageNumber="49" rank="species" species="kangxianensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. kangxianensis</emphasis>
</taxonomicName>
(Yang, Wang, Hu &amp; Jiang, 2011), 
<taxonomicName lsidName="N. quadranus" pageId="0" pageNumber="49" rank="species" species="quadranus">
<emphasis italics="true" pageId="0" pageNumber="49">N. quadranus</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="N. taihangensis" pageId="0" pageNumber="49" rank="species" species="taihangensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. taihangensis</emphasis>
</taxonomicName>
by the strongly hypertrophied forelimbs in adult males (vs not hypertrophied), and by the presence of nuptial spines on the chest and fingers in adult males (vs absence).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. medogensis" pageId="0" pageNumber="49" rank="species" species="medogensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. medogensis</emphasis>
</taxonomicName>
(Fei &amp; Ye, 1999), 
<taxonomicName lsidName="N. phrynoides" pageId="0" pageNumber="49" rank="species" species="phrynoides">
<emphasis italics="true" pageId="0" pageNumber="49">N. phrynoides</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="N. sichuanensis" pageId="0" pageNumber="49" rank="species" species="sichuanensis">
<emphasis italics="true" pageId="0" pageNumber="49">N. sichuanensis</emphasis>
</taxonomicName>
by smooth anterior dorsum skin (vs many warts present).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. parkeri" pageId="0" pageNumber="49" rank="species" species="parkeri">
<emphasis italics="true" pageId="0" pageNumber="49">N. parkeri</emphasis>
</taxonomicName>
(Stejneger, 1927), 
<taxonomicName lsidName="N. pleskei" pageId="0" pageNumber="49" rank="species" species="pleskei">
<emphasis italics="true" pageId="0" pageNumber="49">N. pleskei</emphasis>
</taxonomicName>
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
, 1896, and 
<taxonomicName lsidName="N. ventripunctata" pageId="0" pageNumber="49" rank="species" species="ventripunctata">
<emphasis italics="true" pageId="0" pageNumber="49">N. ventripunctata</emphasis>
</taxonomicName>
Fei &amp; Huang, 1985 by the shape of the nuptial spines (large and conical spines vs tiny and compact spines).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Liu &amp; Zhang &amp; Rao" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Nanorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanorana xuelinensis" order="Anura" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="xuelinensis">
<emphasis italics="true" pageId="0" pageNumber="49">Nanorana xuelinensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="N. vicina" pageId="0" pageNumber="49" rank="species" species="vicina">
<emphasis italics="true" pageId="0" pageNumber="49">N. vicina</emphasis>
</taxonomicName>
(Stoliczka, 1872) by its toes ca 2/3 webbed (vs fully webbed) and by the absence of bands on the hind limbs (vs presence).
</paragraph>
</subSubSection>
</treatment>
</document>