<documentid="6AD55D1C2EED50D5897E712DEAE789EB"ID-CLB-Dataset="299719"ID-DOI="10.11646/zootaxa.5474.5.8"ID-GBIF-Dataset="09a20124-10af-4376-a2f8-cedb504e1b52"ID-ISSN="1175-5326"ID-Zenodo-Dep="12731004"ID-ZooBank="170DE3E1-360B-4B15-A29B-C1813F04E7CC"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="tatiana"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1720374090561"checkinUser="plazi"docAuthor="Shavrin, Alexey V."docDate="2024"docId="03FD87C7EC127A15FF6CFAB8734BFB22"docLanguage="en"docName="zootaxa.5474.5.8.pdf"docOrigin="Zootaxa 5474 (5)"docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.8"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Mannerheimia emodensa Coiffait 1977"docType="treatment"docVersion="4"lastPageNumber="589"masterDocId="FFC4FFBFEC117A10FFFBFFE27115FFE7"masterDocTitle="On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)"masterLastPageNumber="592"masterPageNumber="584"pageNumber="587"updateTime="1720775253469"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="6A97D0544EE19181E66481DECED79D5F">On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
<bibRefCitationid="EFC54B20EC127A13FE28FAB87368FA93"author="Coiffait, H."box="[467,637,1370,1396]"pageId="3"pageNumber="587"pagination="243 - 275"refId="ref4812"refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]"type="journal article"year="1977">Coiffait, 1977</bibRefCitation>
<bibRefCitationid="EFC54B20EC127A13FE65FA217377FA3C"author="Coiffait, H."box="[414,610,1475,1499]"pageId="3"pageNumber="587"pagination="243 - 275"refId="ref4812"refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]"type="journal article"year="1977">Coiffait, 1977: 266</bibRefCitation>
<bibRefCitationid="EFC54B20EC127A13FD97FA21723CFA3C"author="Coiffait, H."box="[620,809,1475,1499]"pageId="3"pageNumber="587"pagination="21 - 179"refId="ref4851"refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179."type="journal article"year="1982">Coiffait 1982: 150</bibRefCitation>
</treatmentCitation>
,
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<bibRefCitationid="EFC54B20EC127A13FCCFFA2172FAFA3C"author="Shavrin, A. V."box="[820,1007,1475,1499]"pageId="3"pageNumber="587"pagination="301 - 333"refId="ref4932"refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1"type="journal article"year="2021">Shavrin 2021: 307</bibRefCitation>
; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Umg.Alm Darghari | b. Maharigaon, 4000m’<printed>, ‘Gebiet von Jumla | Westnepal,lg.H.Franz’<printed>, ‘TYPE’<red, printed>, ‘
<bibRefCitationid="EFC54B20EC127A13FD19F95F726AF930"author="Coiffait, H."box="[738,895,1725,1751]"pageId="3"pageNumber="587"pagination="243 - 275"refId="ref4812"refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112, 243 - 275. [1976]"type="journal article"year="1977">Coiffait, 1977</bibRefCitation>
</taxonomicName>
| Shavrin A.
<collectionCodeid="ED45AE14EC127A13FBF3F95C750BF93F"box="[1032,1054,1726,1752]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="3"pageNumber="587"type="Museum">V</collectionCode>
<typeStatusid="54EF8873EC127A13FF3CF9037320F91C"box="[199,565,1761,1787]"isOtherTypeStatus="true"pageId="3"pageNumber="587">Additional material examined</typeStatus>
<materialsCitationid="3B3C3C8CEC127A13FB90F8E773E9F8A4"ID-GBIF-Occurrence="4905213307"collectionCode="NMW"collectorName="H. Franz"country="Nepal"location="Gebiet des Rarasees Westnepal"pageId="3"pageNumber="587"specimenCount="1"specimenCount-female="1">
<locationid="8E8B600AEC127A13FF60F8CB70EAF8A4"LSID="urn:lsid:plazi:treatment:03FD87C7EC127A15FF6CFAB8734BFB22:8E8B600AEC127A13FF60F8CB70EAF8A4"box="[155,511,1833,1860]"country="Nepal"name="Gebiet des Rarasees Westnepal"pageId="3"pageNumber="587">Gebiet des Rarasees Westnepal</location>
. Body yellow-brown to reddish-brown, with distinctly darker head (abdomen of some specimens slightly darker than forebody; lateral and basal portions of pronotum and apical portion of elytra slightly paler in
); antennomeres 5–11 or 6–11 brown; mouthparts, antennomeres 1–4 or 1–5 and paratergites yellow. Forebody without microsculpture; abdomen with fine and dense transverse microreticulation. Head with moderately fine and sparse punctation, distinctly denser in middle; punctation of pronotum denser, larger and deeper than that in middle portion of head, finer and sparser in middle, and sometimes with narrow impunctate portion in middle or in mediobasal third; scutellum with one to four fine punctures in middle or without them; punctation of elytra denser, slightly larger and deeper than that on pronotum, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, but some specimens with markedly coarse surface without longitudinal rows between punctures.
<paragraphid="8BEB36D1EC147A15FF3CFF757312FEA6"blockId="5.[151,1437,150,1221]"pageId="5"pageNumber="589">Head 1.5–1.6 times as broad as long, slightly elevated in middle, with very short and sometimes indistinct anteocellar foveae as long as diameter of ocellus. Antennomere 3 narrower and slightly shorter than 2, 4 distinctly shorter than 3 and slightly longer than broad, 5–6 indistinctly broader than 4, 7–8 slightly longer and distinctly broader than 6, 9–10 slightly longer and broader than 8, apical antennomere about as long as two preceding antennomeres or slightly shorter.</paragraph>
<paragraphid="8BEB36D1EC147A15FF3CFEA97271FE4A"blockId="5.[151,1437,150,1221]"pageId="5"pageNumber="589">Pronotum 1.8 times as broad as long, 1.2 times as broad as head, widest slightly above middle, evenly narrowed toward obtuse posterior angles, sometimes with indistinctly sinuate laterobasal margins; lateral portions slightly impressed, distinctly broader and deeper in laterobasal portions.</paragraph>
<paragraphid="8BEB36D1EC147A15FF3CFE55750DFE36"blockId="5.[151,1437,150,1221]"box="[199,1048,439,465]"pageId="5"pageNumber="589">Elytra 1.2–1.3 times broader than long, slightly or strongly widened apicad.</paragraph>
<paragraphid="8BEB36D1EC147A15FF3CFE397389FE12"blockId="5.[151,1437,150,1221]"box="[199,668,475,501]"pageId="5"pageNumber="589">Metatarsus 1.7 times as long as metatibia.</paragraph>
<paragraphid="8BEB36D1EC147A15FF3CFE1D708DFDDA"blockId="5.[151,1437,150,1221]"pageId="5"pageNumber="589">Abdomen about as broad as elytra, with two small oval or transverse tomentose wing-folding patches in middle of abdominal tergite V.</paragraph>
Male. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed apically, with relatively narrow median lobe and small rounded apex; parameres with wide apical portions, slightly exceeding apex of median lobe; internal sac narrow and moderately long (
<paragraphid="8BEB36D1EC147A15FF3CFD35709EFCF2"blockId="5.[151,1437,150,1221]"pageId="5"pageNumber="589">Female. Apical margin of abdominal tergite VIII straight or somewhat rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
, from which can be distinguished by the slightly broader elytra, larger and denser punctation on the pronotum and elytra, slightly shorter preapical antennomeres, and longer parameres with significantly broader apical portions.
<quantityid="4CAC9B34EC147A15FCB4FC31751CFC0A"box="[847,1033,979,1005]"metricMagnitude="3"metricUnit="m"metricValue="3.5"metricValueMax="4.0"metricValueMin="3.0"pageId="5"pageNumber="589"unit="m"value="3500.0"valueMax="4000.0"valueMin="3000.0">3000 to 4000 m</quantity>
<bibRefCitationid="EFC54B20EC147A15FB70FB657421FB46"author="Shavrin, A. V."box="[1163,1332,1159,1185]"pageId="5"pageNumber="589"pagination="301 - 333"refId="ref4932"refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1"type="journal article"year="2021">Shavrin (2021)</bibRefCitation>
