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<document id="0F7B35405C3B417FA9CF1C19C6238062" ID-CLB-Dataset="39054" ID-DOI="10.11646/zootaxa.4111.3.5" ID-GBIF-Dataset="7c678046-0584-4c1a-942f-ebcd91fb4947" ID-ISSN="1175-5326" ID-Zenodo-Dep="264878" ID-ZooBank="9344C93C-FF15-4361-8EC0-1B60EE1844EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463468991456" checkinUser="plazi" docAuthor="Rodriguez, Pedro Alexander, Rodriguez, Erick J., Norrbom, Allen L. &amp; Arévalo, Emilio" docDate="2016" docId="03FDB91F9747507233ECFF5D8FF20503" docLanguage="en" docName="zootaxa.4111.3.5.pdf.imf" docOrigin="Zootaxa 4111 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptodacus trinotatus Norrbom &amp; Korytkowski" docType="treatment" docVersion="10" lastPageNumber="289" masterDocId="FFC4C1679740507F337BFFCA8A6E0236" masterDocTitle="A new species and new records of Cryptodacus (Diptera: Tephritidae) from Colombia, Bolivia and Peru" masterLastPageNumber="290" masterPageNumber="276" pageNumber="283" updateTime="1698662529496" updateUser="ExternalLinkService">
<mods:mods id="CA9DD8E6603DDF21BE61039D6832375F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5702753BFDB9524DB374EAC8F8F7574F">
<mods:title id="ACFA78FC16B3B80ED9C0CA569B9EC405">A new species and new records of Cryptodacus (Diptera: Tephritidae) from Colombia, Bolivia and Peru</mods:title>
</mods:titleInfo>
<mods:name id="60C3D22324A1D06FE7B1793066AC864D" type="personal">
<mods:role id="C3C67E8F898E0B9A5207EE7608BC7E32">
<mods:roleTerm id="2CC1E52867927BA48B44F1E7EDB32077">Author</mods:roleTerm>
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<mods:namePart id="D86818CF4143302CCC2AD03158D2A5A0">Rodriguez, Pedro Alexander</mods:namePart>
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<mods:name id="9F8130D98326354E46D616E75936C91E" type="personal">
<mods:role id="B294F4DC5B576B9D8D405214D7A4B767">
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<mods:namePart id="61794240296D6EBA6AED13D85FD4013E">Rodriguez, Erick J.</mods:namePart>
</mods:name>
<mods:name id="DFCD40008D923D137B28855264712208" type="personal">
<mods:role id="61D80349F86DD933F40453675C92C17D">
<mods:roleTerm id="474DB35DC794824CD831785022963C86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6196A472CBFEF1F466244F3DDA58CE1">Norrbom, Allen L.</mods:namePart>
</mods:name>
<mods:name id="B2DB79F14443C970EEBBDF3E396403C8" type="personal">
<mods:role id="C55044AF2095E8FB46F1F341FAC5F31A">
<mods:roleTerm id="1E0CF34FE74A09F1EF2B21351C078E1D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5635B5247153872F30EDF90C034B9B18">Arévalo, Emilio</mods:namePart>
</mods:name>
<mods:typeOfResource id="0D24A0A09D6CB48EA0992233B850A775">text</mods:typeOfResource>
<mods:relatedItem id="BB14BD61A8FED219B152A4953731501D" type="host">
<mods:titleInfo id="8B593ADF70E235037A3E26221F45D837">
<mods:title id="23AB590870CB2F9C03865652066DB893">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8992127EB54E75B8308DAF74AA800A55">
<mods:date id="CA002A54B9BB372EC4302BBF1397AFAB">2016</mods:date>
<mods:detail id="A294D9655848EE730B108A6B7B65E5AE" type="volume">
<mods:number id="2F2CF9B1884AED6F6F46475976EB7AB4">4111</mods:number>
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<mods:detail id="89A9385FA371071197EA760371293CAB" type="issue">
<mods:number id="5D19893A503B5043BE5446BCFE9D5347">3</mods:number>
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<mods:start id="522F7D120D6D5C715BEEB7DE697BFF23">276</mods:start>
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<mods:classification id="FBFF59AB86C8F5C0A1599E77D8ED6EE1">journal article</mods:classification>
<mods:identifier id="2FD6597F27BFFDB04AD9887889095257" type="CLB-Dataset">39054</mods:identifier>
<mods:identifier id="65D5D8ED68445AA353705A86F73F9C5D" type="DOI">10.11646/zootaxa.4111.3.5</mods:identifier>
<mods:identifier id="47E2B99ECB553A38A194CAD9C466BFB1" type="GBIF-Dataset">7c678046-0584-4c1a-942f-ebcd91fb4947</mods:identifier>
<mods:identifier id="004F174120FB0C5D93AF81028F4A21E9" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FDB91F9747507233ECFF5D8FF20503" ID-DOI="http://doi.org/10.5281/zenodo.6074626" ID-GBIF-Taxon="120678437" ID-Zenodo-Dep="6074626" LSID="urn:lsid:plazi:treatment:03FDB91F9747507233ECFF5D8FF20503" httpUri="http://treatment.plazi.org/id/03FDB91F9747507233ECFF5D8FF20503" lastPageId="13" lastPageNumber="289" pageId="7" pageNumber="283">
<subSubSection id="C34E5B829747507833ECFF5D8B9F03B1" pageId="7" pageNumber="283" type="nomenclature">
<paragraph id="8BEB08099747507833ECFF5D889F0284" blockId="7.[151,753,151,211]" box="[151,753,151,178]" pageId="7" pageNumber="283">
<heading id="D0A3BF659747507833ECFF5D889F0284" bold="true" box="[151,753,151,178]" fontSize="11" level="1" pageId="7" pageNumber="283" reason="1">
<taxonomicName id="4C54738A9747507833ECFF5D889F0284" ID-CoL="8BM4C" authority="Norrbom &amp; Korytkowski" authorityName="Norrbom &amp; Korytkowski" box="[151,753,151,178]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B9747507833ECFF5D889F0284" bold="true" box="[151,753,151,178]" pageId="7" pageNumber="283">
<emphasis id="B920D41B9747507833ECFF5D8BC40287" bold="true" box="[151,426,151,177]" italics="true" pageId="7" pageNumber="283">Cryptodacus trinotatus</emphasis>
Norrbom &amp; Korytkowski
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BEB08099747507833ECFF70886702E5" blockId="7.[151,753,151,211]" box="[151,521,186,211]" pageId="7" pageNumber="283">
<figureCitation id="136F148C9747507833ECFF708A8B02E5" box="[151,229,186,211]" captionStart="FIGURES 3 – 4" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1344,216,1886]" captionTargetId="figure@5.[216,1370,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 4. Female habitus: 3, C. ornatus (Colombia: Tolima); 4, C. trinotatus (Colombia: La Mesa ICAMF 00000046)" httpUri="https://zenodo.org/record/264880/files/figure.png" pageId="7" pageNumber="283">Figs. 4</figureCitation>
,
<figureCitation id="136F148C97475078338AFF718B5602E5" box="[241,312,187,211]" captionStart="FIGURES 11 – 13. 11 – 12, C" captionStartId="7.[151,264,1740,1763]" captionTargetBox="[160,1373,583,1676]" captionTargetId="figure@7.[151,1436,518,1719]" captionTargetPageId="7" captionText="FIGURES 11 – 13. 11 – 12, C. trinotatus (Colombia: La Mesa ICAMF 00000046): 11, head, anteroventral; 12, head lateral. 13, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969), head, dorsal." httpUri="https://zenodo.org/record/264882/files/figure.png" pageId="7" pageNumber="283">11, 12</figureCitation>
,
<figureCitation id="136F148C97475078323FFF718B0C02E5" box="[324,354,187,211]" captionStart="FIGURES 14 – 18. 14 – 15, C" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[231,1350,287,1927]" captionTargetId="figure@8.[225,1362,193,1932]" captionTargetPageId="8" captionText="FIGURES 14 – 18. 14 – 15, C. bernardoi (Colombia: Anolaima ICAMF 00000044): 14, thorax, dorsal; 15, thorax, lateral. 16, C. ornatus (Colombia: Tolima), thorax, dorsal. 17, C. trinotatus, (Colombia: La Mesa ICAMF 00000046), thorax, dorsal. 18, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969), thorax, dorsal." httpUri="https://zenodo.org/record/264883/files/figure.png" pageId="7" pageNumber="283">17</figureCitation>
,
<figureCitation id="136F148C974750783215FF718BE202E5" box="[366,396,187,211]" captionStart="FIGURES 19 – 21" captionStartId="9.[151,264,855,878]" captionTargetBox="[155,1430,199,829]" captionTargetId="figure@9.[151,1435,193,835]" captionTargetPageId="9" captionText="FIGURES 19 – 21. Wings: 19, C. bernardoi (Colombia: Anolaima ICAMF 00000044); 20, C. ornatus (Colombia: Tolima); 21, C. trinotatus (Colombia: La Mesa ICAMF 00000046)." httpUri="https://zenodo.org/record/264884/files/figure.png" pageId="7" pageNumber="283">21</figureCitation>
,
<figureCitation id="136F148C9747507832E3FF718BD802E5" box="[408,438,187,211]" captionStart="FIGURES 24 – 27" captionStartId="10.[151,264,1791,1814]" captionTargetBox="[187,1423,236,1754]" captionTargetId="figure@10.[151,1497,193,1771]" captionTargetPageId="10" captionText="FIGURES 24 – 27. Female abdomen: 24 – 25, C. ornatus (Colombia: Tolima): 24, dorsal; 25, oviscape, lateral. 26, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969), dorsal. 27, C. trinotatus (Colombia: La Mesa ICAMF 00000046), dorsal." httpUri="https://zenodo.org/record/264886/files/figure.png" pageId="7" pageNumber="283">27</figureCitation>
,
<figureCitation id="136F148C9747507832B9FF71886702E5" box="[450,521,187,211]" captionStart="FIGURES 28 – 37" captionStartId="11.[151,264,1786,1809]" captionTargetBox="[191,1412,223,1735]" captionTargetId="figure@11.[151,1436,193,1765]" captionTargetPageId="11" captionText="FIGURES 28 – 37. Female terminalia: 28 – 31, C. bernardoi (Colombia: Guaduas ICAMF 00000045): 28, aculeus; 29, aculeus tip; 30, aculeus tip, line drawing; 31, spermatheca. 32 – 34, C. trinotatus (Colombia: La Mesa ICAMF 00000046): 32, aculeus; 33, aculeus tip; 34, aculeus tip, line drawing. 35, C. ornatus, aculeus tip (Colombia; copied from Norrbom 1994, fig. 5 E). 36 – 37, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969): 36, aculeus; 37, aculeus tip." httpUri="https://zenodo.org/record/264887/files/figure.png" pageId="7" pageNumber="283">32–34</figureCitation>
</paragraph>
<paragraph id="8BEB08099747507833ECFEC88B9F03B1" blockId="7.[151,1436,258,463]" pageId="7" pageNumber="283">
<emphasis id="B920D41B9747507833ECFEC88943032D" bold="true" box="[151,813,258,283]" pageId="7" pageNumber="283">
Specimens examined.
<collectingCountry id="F34348999747507832CEFEC88836032D" box="[437,600,258,283]" name="Colombia" pageId="7" pageNumber="283">COLOMBIA</collectingCountry>
: Cundinamarca:
</emphasis>
La Mesa, entrada al sitio los Pinos [
<geoCoordinate id="EE606ECE97475078365AFEC88FF2032D" box="[1313,1436,258,283]" direction="north" orientation="latitude" pageId="7" pageNumber="283" precision="15" value="5.074167">4º63’87”N</geoCoordinate>
<geoCoordinate id="EE606ECE9747507833ECFEEC8B460309" box="[151,296,294,319]" direction="west" orientation="longitude" pageId="7" pageNumber="283" precision="15" value="-74.78639">74º46’71”W</geoCoordinate>
],
<quantity id="4CACA5EC974750783247FEEC8BFA0308" box="[316,404,294,319]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="7" pageNumber="283" unit="m" value="1125.0">1125 m</quantity>
, McPhail trap 24,
<date id="FFEA2EC997475078311DFEED88810309" box="[614,751,294,319]" pageId="7" pageNumber="283" value="2015-07-14">14 Jul 2015</date>
, L. Guerrero, 3♀ (ICAMF00000046).
<emphasis id="B920D41B9747507837DEFEEC8B460355" bold="true" pageId="7" pageNumber="283">Norte de Santander: Pamplonita,</emphasis>
vereda San Rafael predio El Recuerdo [
<geoCoordinate id="EE606ECE974750783185FE8189020354" box="[766,876,331,355]" direction="north" orientation="latitude" pageId="7" pageNumber="283" precision="5" value="7.4443">7,4443 N</geoCoordinate>
–
<geoCoordinate id="EE606ECE9747507830FFFE808E660354" box="[900,1032,330,355]" direction="west" orientation="longitude" pageId="7" pageNumber="283" precision="5" value="-72.6332">72,6332 W</geoCoordinate>
],
<quantity id="4CACA5EC974750783765FE808E160354" box="[1054,1144,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.658" pageId="7" pageNumber="283" unit="m" value="1658.0">1658 m</quantity>
, McPhail trap
<date id="FFEA2EC997475078365EFE808FF20355" box="[1317,1436,330,355]" pageId="7" pageNumber="283" value="1929-04-16">16, 29 abr</date>
2014. 1 ♀ (ICAMF00000100).
</paragraph>
</subSubSection>
<subSubSection id="C34E5B829747507833BCFE588E4F03F9" pageId="7" pageNumber="283" type="discussion">
<paragraph id="8BEB08099747507833BCFE588E4F03F9" blockId="7.[151,1436,258,463]" pageId="7" pageNumber="283">
<emphasis id="B920D41B9747507833BCFE588B21039D" bold="true" box="[199,335,402,427]" pageId="7" pageNumber="283">Comments.</emphasis>
This species has been reported previously only from
<collectingCountry id="F34348999747507830C6FE598E78039D" box="[957,1046,403,427]" name="Panama" pageId="7" pageNumber="283">Panamá</collectingCountry>
(
<bibRefCitation id="EFC575F897475078375DFE588FFF039D" author="Norrbom" box="[1062,1425,402,427]" pageId="7" pageNumber="283" refString="Norrbom, A. L. &amp; Korytkowski, C. A. (2008) New Cryptodacus (Diptera: Tephritidae) from Panama, with a key to the known species. Zootaxa, 1773, 31 - 43." type="journal article" year="2008">Norrbom &amp; Korytkowski 2008</bibRefCitation>
). The examined specimens are the first record from
<collectingCountry id="F34348999747507831B0FE7C895703F9" box="[715,825,438,463]" name="Colombia" pageId="7" pageNumber="283">Colombia</collectingCountry>
and South
<collectingCountry id="F34348999747507830C3FE7C8E7203F9" box="[952,1052,438,463]" name="United States of America" pageId="7" pageNumber="283">America</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF2B58819747507833ECF90689300534" httpUri="https://zenodo.org/record/264882/files/figure.png" pageId="7" pageNumber="283" targetBox="[160,1373,583,1676]" targetPageId="7">
<paragraph id="8BEB08099747507833ECF90689300534" blockId="7.[151,1436,1740,1794]" pageId="7" pageNumber="283">
<emphasis id="B920D41B9747507833ECF9068B3D04D4" bold="true" box="[151,339,1740,1763]" pageId="7" pageNumber="283">FIGURES 11–13.</emphasis>
11–12,
<taxonomicName id="4C54738A9747507832DCF904884704D5" box="[423,553,1742,1763]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B9747507832DCF904884704D5" box="[423,553,1742,1763]" italics="true" pageId="7" pageNumber="283">C. trinotatus</emphasis>
</taxonomicName>
(Colombia: La Mesa ICAMF00000046): 11, head, anteroventral; 12, head lateral. 13,
<taxonomicName id="4C54738A9747507833ECF9268B650534" box="[151,267,1772,1794]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B920D41B9747507833ECF9268B650534" box="[151,267,1772,1794]" italics="true" pageId="7" pageNumber="283">C. obliquus</emphasis>
</taxonomicName>
(Bolivia: Santa Cruz USNMENT00875969), head, dorsal.
</paragraph>
</caption>
<caption id="DF2B58819748507733ECF86B893C05C2" httpUri="https://zenodo.org/record/264883/files/figure.png" pageId="8" pageNumber="284" targetBox="[231,1350,287,1927]" targetPageId="8">
<paragraph id="8BEB08099748507733ECF86B893C05C2" blockId="8.[151,1436,1953,2036]" pageId="8" pageNumber="284">
<emphasis id="B920D41B9748507733ECF86B8B3A0580" bold="true" box="[151,340,1953,1976]" pageId="8" pageNumber="284">FIGURES 14–18.</emphasis>
14–15,
<taxonomicName id="4C54738A9748507732DDF86888470581" box="[422,553,1953,1975]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="284" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B9748507732DDF86888470581" box="[422,553,1953,1975]" italics="true" pageId="8" pageNumber="284">C. bernardoi</emphasis>
</taxonomicName>
(Colombia: Anolaima ICAMF00000044): 14, thorax, dorsal; 15, thorax, lateral. 16,
<taxonomicName id="4C54738A9748507736FFF8688A8A05E0" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="284" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B920D41B9748507736FFF8688A8A05E0" italics="true" pageId="8" pageNumber="284">C. ornatus</emphasis>
</taxonomicName>
(Colombia: Tolima), thorax, dorsal. 17,
<taxonomicName id="4C54738A9748507731F2F80B896505E0" box="[649,779,1985,2006]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="284" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B9748507731F2F80B896505E0" box="[649,779,1985,2006]" italics="true" pageId="8" pageNumber="284">C. trinotatus</emphasis>
</taxonomicName>
, (Colombia: La Mesa ICAMF00000046), thorax, dorsal. 18,
<taxonomicName id="4C54738A9748507736FFF80B8A8005C2" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="284" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B920D41B9748507736FFF80B8A8005C2" italics="true" pageId="8" pageNumber="284">C. obliquus</emphasis>
</taxonomicName>
(Bolivia: Santa Cruz USNMENT00875969), thorax, dorsal.
</paragraph>
</caption>
<caption id="DF2B58819749507633ECFC9D88D401BA" httpUri="https://zenodo.org/record/264884/files/figure.png" pageId="9" pageNumber="285" targetBox="[155,1430,199,829]" targetPageId="9">
<paragraph id="8BEB08099749507633ECFC9D88D401BA" blockId="9.[151,1435,855,909]" pageId="9" pageNumber="285">
<emphasis id="B920D41B9749507633ECFC9D8B3A015B" bold="true" box="[151,340,855,878]" pageId="9" pageNumber="285">FIGURES 19–21.</emphasis>
Wings: 19,
<taxonomicName id="4C54738A9749507632AAFC93883A0158" box="[465,596,856,878]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="285" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B9749507632AAFC93883A0158" box="[465,596,856,878]" italics="true" pageId="9" pageNumber="285">C. bernardoi</emphasis>
</taxonomicName>
(Colombia: Anolaima ICAMF00000044); 20,
<taxonomicName id="4C54738A974950763749FC938EF30158" box="[1074,1181,857,878]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="285" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B920D41B974950763749FC938EF30158" box="[1074,1181,857,878]" italics="true" pageId="9" pageNumber="285">C. ornatus</emphasis>
</taxonomicName>
(Colombia: Tolima); 21,
<taxonomicName id="4C54738A9749507633ECFCBD8B7801BB" box="[151,278,887,909]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="285" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B9749507633ECFCBD8B7801BB" box="[151,278,887,909]" italics="true" pageId="9" pageNumber="285">C. trinotatus</emphasis>
</taxonomicName>
(Colombia: La Mesa ICAMF00000046).
</paragraph>
</caption>
<caption id="DF2B58819749507633ECF90F881704CC" httpUri="https://zenodo.org/record/264885/files/figure.png" pageId="9" pageNumber="285" targetBox="[157,1411,979,1699]" targetPageId="9">
<paragraph id="8BEB08099749507633ECF90F881704CC" blockId="9.[151,1436,1733,1786]" pageId="9" pageNumber="285">
<emphasis id="B920D41B9749507633ECF90F8B3804ED" bold="true" box="[151,342,1733,1756]" pageId="9" pageNumber="285">FIGURES 22–23.</emphasis>
<taxonomicName id="4C54738A974950763224F90C8B8D04ED" box="[351,483,1733,1755]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="285" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B974950763224F90C8B8D04ED" box="[351,483,1733,1755]" italics="true" pageId="9" pageNumber="285">C. bernardoi</emphasis>
</taxonomicName>
: 22, female abdomen, dorsal (Colombia: Anolaima ICAMF00000044); 23, male abdomen, dorsal (Colombia: Guaduas ICAMF00000045).
</paragraph>
</caption>
<caption id="DF2B5881974A507533ECF9358F2E0503" httpUri="https://zenodo.org/record/264886/files/figure.png" pageId="10" pageNumber="286" targetBox="[187,1423,236,1754]" targetPageId="10">
<paragraph id="8BEB0809974A507533ECF9358F2E0503" blockId="10.[151,1436,1791,1845]" pageId="10" pageNumber="286">
<emphasis id="B920D41B974A507533ECF9358B3A0523" bold="true" box="[151,340,1791,1814]" pageId="10" pageNumber="286">FIGURES 24–27.</emphasis>
Female abdomen: 24–25,
<taxonomicName id="4C54738A974A5075311EF8CB88BF0520" box="[613,721,1793,1814]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B920D41B974A5075311EF8CB88BF0520" box="[613,721,1793,1814]" italics="true" pageId="10" pageNumber="286">C. ornatus</emphasis>
</taxonomicName>
(Colombia: Tolima): 24, dorsal; 25, oviscape, lateral. 26,
<taxonomicName id="4C54738A974A5075365DF8CB8FF20520" box="[1318,1436,1792,1814]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B920D41B974A5075365DF8CB8FF20520" box="[1318,1436,1792,1814]" italics="true" pageId="10" pageNumber="286">C. obliquus</emphasis>
</taxonomicName>
(Bolivia: Santa Cruz USNMENT00875969), dorsal. 27,
<taxonomicName id="4C54738A974A507531A9F8EA893D0503" box="[722,851,1824,1845]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B974A507531A9F8EA893D0503" box="[722,851,1824,1845]" italics="true" pageId="10" pageNumber="286">C. trinotatus</emphasis>
</taxonomicName>
(Colombia: La Mesa ICAMF00000046), dorsal.
</paragraph>
</caption>
<caption id="DF2B5881974B507433ECF9308E2D055A" httpUri="https://zenodo.org/record/264887/files/figure.png" pageId="11" pageNumber="287" targetBox="[191,1412,223,1735]" targetPageId="11">
<paragraph id="8BEB0809974B507433ECF9308E2D055A" blockId="11.[151,1436,1786,1900]" pageId="11" pageNumber="287">
<emphasis id="B920D41B974B507433ECF9308B3B0539" bold="true" box="[151,341,1786,1809]" pageId="11" pageNumber="287">FIGURES 28–37.</emphasis>
Female terminalia: 28–31,
<taxonomicName id="4C54738A974B50743109F93188980526" box="[626,758,1786,1808]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="287" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B974B50743109F93188980526" box="[626,758,1786,1808]" italics="true" pageId="11" pageNumber="287">C. bernardoi</emphasis>
</taxonomicName>
(Colombia: Guaduas ICAMF00000045): 28, aculeus; 29, aculeus tip; 30, aculeus tip, line drawing; 31, spermatheca. 32–34,
<taxonomicName id="4C54738A974B50743189F8D0891A0519" box="[754,884,1818,1839]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="287" phylum="Arthropoda" rank="species" species="trinotatus">
<emphasis id="B920D41B974B50743189F8D0891A0519" box="[754,884,1818,1839]" italics="true" pageId="11" pageNumber="287">C. trinotatus</emphasis>
</taxonomicName>
(Colombia: La Mesa ICAMF00000046): 32, aculeus; 33, aculeus tip; 34, aculeus tip, line drawing. 35,
<taxonomicName id="4C54738A974B507431D8F8F2897E0578" box="[675,784,1848,1870]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="287" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B920D41B974B507431D8F8F2897E0578" box="[675,784,1848,1870]" italics="true" pageId="11" pageNumber="287">C. ornatus</emphasis>
</taxonomicName>
, aculeus tip (Colombia; copied from Norrbom 1994, fig. 5E). 36–37,
<taxonomicName id="4C54738A974B50743398F89D8B39055A" box="[227,343,1878,1900]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="287" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B920D41B974B50743398F89D8B39055A" box="[227,343,1878,1900]" italics="true" pageId="11" pageNumber="287">C. obliquus</emphasis>
</taxonomicName>
(Bolivia: Santa Cruz USNMENT00875969): 36, aculeus; 37, aculeus tip.
</paragraph>
</caption>
<caption id="DF2B5881974C507333ECF86B8B0705C2" httpUri="https://zenodo.org/record/264888/files/figure.png" pageId="12" pageNumber="288" targetBox="[245,1344,215,1921]" targetPageId="12">
<paragraph id="8BEB0809974C507333ECF86B8B0705C2" blockId="12.[151,1436,1953,2036]" pageId="12" pageNumber="288">
<emphasis id="B920D41B974C507333ECF86B8B390580" bold="true" box="[151,343,1953,1976]" pageId="12" pageNumber="288">FIGURES 38–42.</emphasis>
Male terminalia,
<taxonomicName id="4C54738A974C5073316DF86888F50581" box="[534,667,1953,1975]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="288" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B974C5073316DF86888F50581" box="[534,667,1953,1975]" italics="true" pageId="12" pageNumber="288">C. bernardoi</emphasis>
</taxonomicName>
(Colombia: Guaduas ICAMF00000045): 38, abdomen, lateroventral; 39, distiphallus; 40, epandrium and surstyli, lateral and posterodorsal; 41, distiphallus, line drawing; 42, epandrium and surstyli, lateral, line drawing.
</paragraph>
</caption>
<caption id="DF2B5881974D507233ECFA958FE70740" box="[151,1417,1375,1398]" httpUri="https://zenodo.org/record/264889/files/figure.png" pageId="13" pageNumber="289" targetBox="[156,1432,194,1349]" targetPageId="13">
<paragraph id="8BEB0809974D507233ECFA958FE70740" blockId="13.[151,1417,1375,1398]" box="[151,1417,1375,1398]" pageId="13" pageNumber="289">
<emphasis id="B920D41B974D507233ECFA958B3D0743" bold="true" box="[151,339,1375,1398]" pageId="13" pageNumber="289">FIGURES 43–44.</emphasis>
<taxonomicName id="4C54738A974D50723221FAAA8B830740" box="[346,493,1376,1398]" class="Magnoliopsida" family="Santalaceae" genus="Phoradendron" kingdom="Plantae" order="Santalales" pageId="13" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="B920D41B974D50723221FAAA8B830740" box="[346,493,1376,1398]" italics="true" pageId="13" pageNumber="289">Phoradendron</emphasis>
</taxonomicName>
sp. near
<emphasis id="B920D41B974D50723133FAAA88DC0740" box="[584,690,1376,1398]" italics="true" pageId="13" pageNumber="289">piperoides</emphasis>
(Kunth) Trelease (
<taxonomicName id="4C54738A974D5072300BFAAA89B20740" box="[880,988,1376,1398]" class="Magnoliopsida" family="Viscaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Santalales" pageId="13" pageNumber="289" phylum="Tracheophyta" rank="family">Viscaceae</taxonomicName>
): 43, section of the plant; 44, fruits, detail.
</paragraph>
</caption>
<subSubSection id="C34E5B82974D507233ECFA0D8FF20503" pageId="13" pageNumber="289" type="key">
<paragraph id="8BEB0809974D507233ECFA0D8B600435" blockId="13.[151,1436,1479,1539]" pageId="13" pageNumber="289">
<emphasis id="B920D41B974D507233ECFA0D8AA307D6" bold="true" box="[151,205,1479,1504]" pageId="13" pageNumber="289">Key.</emphasis>
The following modifications to the key of
<bibRefCitation id="EFC575F8974D507231A2FA0D8E3907E9" author="Norrbom" box="[729,1111,1479,1504]" pageId="13" pageNumber="289" refString="Norrbom, A. L. &amp; Korytkowski, C. A. (2008) New Cryptodacus (Diptera: Tephritidae) from Panama, with a key to the known species. Zootaxa, 1773, 31 - 43." type="journal article" year="2008">Norrbom &amp; Korytkowski (2008)</bibRefCitation>
are provided to include
<taxonomicName id="4C54738A974D507236F9FA028B680435" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="289" phylum="Arthropoda" rank="species" species="bernardoi">
<emphasis id="B920D41B974D507236F9FA028B680435" italics="true" pageId="13" pageNumber="289">C. bernardoi</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8BEB0809974D507233ECF9F88FF20503" blockId="13.[151,1436,1586,1849]" pageId="13" pageNumber="289">
2. Female abdominal tergites 5 and 6 with sublateral white areas bordering medial brown vitta or vittae. Crossvein dm-m entirely covered by discal band or, if covered by subapical band (
<taxonomicName id="4C54738A974D5072307AF99A893B0453" box="[769,853,1616,1637]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="289" phylum="Arthropoda" rank="species" species="lopezi">
<emphasis id="B920D41B974D5072307AF99A893B0453" box="[769,853,1616,1637]" italics="true" pageId="13" pageNumber="289">C. lopezi</emphasis>
</taxonomicName>
), face yellow, with single ventromedial brown spot.
<collectingCountry id="F3434899974D50723636F9858FF60453" box="[1357,1432,1615,1637]" name="Mexico" pageId="13" pageNumber="289">Mexico</collectingCountry>
, Central
<collectingCountry id="F3434899974D50723256F9A68B1104B4" box="[301,383,1644,1666]" name="United States of America" pageId="13" pageNumber="289">America</collectingCountry>
...................................................................................... 3 - Female abdominal tergites 5 and 6 without sublateral white areas. Crossvein dm-m at most with posterior half covered by discal band. Face entirely yellow or with more than one brown marking.
<collectingCountry id="F3434899974D50723004F96D8984048B" box="[895,1002,1703,1725]" name="Costa Rica" pageId="13" pageNumber="289">Costa Rica</collectingCountry>
, South
<collectingCountry id="F3434899974D50723749F96D8EEA048B" box="[1074,1156,1703,1725]" name="United States of America" pageId="13" pageNumber="289">America</collectingCountry>
.....................4A 4A. Face yellow with brown markings (
<figureCitation id="136F148C974D50723143F90E88CB04EC" box="[568,677,1732,1754]" captionStart-0="FIGURES 5 – 10. 5 – 8, C" captionStart-1="FIGURES 11 – 13. 11 – 12, C" captionStartId-0="6.[151,264,1944,1967]" captionStartId-1="7.[151,264,1740,1763]" captionTargetBox-0="[181,1398,205,1872]" captionTargetBox-1="[160,1373,583,1676]" captionTargetId-0="figure@6.[151,1436,193,1923]" captionTargetId-1="figure@7.[151,1436,518,1719]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 5 – 10. 5 – 8, C. bernardoi (Colombia: Anolaima ICAMF 00000044): 5, head, anterodorsal; 6, head, anteroventral; 7, head, dorsal; 8, head, lateral. 9 – 10, C. ornatus (Colombia: Tolima): 9, head, dorsal; 10, head, lateral." captionText-1="FIGURES 11 – 13. 11 – 12, C. trinotatus (Colombia: La Mesa ICAMF 00000046): 11, head, anteroventral; 12, head lateral. 13, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969), head, dorsal." httpUri-0="https://zenodo.org/record/264881/files/figure.png" httpUri-1="https://zenodo.org/record/264882/files/figure.png" pageId="13" pageNumber="289">Fig. 10–12</figureCitation>
). Syntergite 1+2 mostly to entirely dark brown (
<figureCitation id="136F148C974D5072370CF90E8F7C04EC" box="[1143,1298,1732,1754]" captionStart="FIGURES 24 – 27" captionStartId="10.[151,264,1791,1814]" captionTargetBox="[187,1423,236,1754]" captionTargetId="figure@10.[151,1497,193,1771]" captionTargetPageId="10" captionText="FIGURES 24 – 27. Female abdomen: 24 – 25, C. ornatus (Colombia: Tolima): 24, dorsal; 25, oviscape, lateral. 26, C. obliquus (Bolivia: Santa Cruz USNMENT 00875969), dorsal. 27, C. trinotatus (Colombia: La Mesa ICAMF 00000046), dorsal." httpUri="https://zenodo.org/record/264886/files/figure.png" pageId="13" pageNumber="289">Figs. 24, 26, 27</figureCitation>
), at most with apical white band. Vein R4+5 not sinuous and crossvein dm-m nearly straight or slightly curved (
<figureCitation id="136F148C974D50723709F9288E8B04CE" box="[1138,1253,1762,1784]" captionStart="FIGURES 19 – 21" captionStartId="9.[151,264,855,878]" captionTargetBox="[155,1430,199,829]" captionTargetId="figure@9.[151,1435,193,835]" captionTargetPageId="9" captionText="FIGURES 19 – 21. Wings: 19, C. bernardoi (Colombia: Anolaima ICAMF 00000044); 20, C. ornatus (Colombia: Tolima); 21, C. trinotatus (Colombia: La Mesa ICAMF 00000046)." httpUri="https://zenodo.org/record/264884/files/figure.png" pageId="13" pageNumber="289">Figs. 20, 21</figureCitation>
).............5 - Face entirely yellow, without brown markings (
<figureCitation id="136F148C974D507231D5F8C8894A052E" box="[686,804,1794,1816]" captionStart="FIGURES 5 – 10. 5 – 8, C" captionStartId="6.[151,264,1944,1967]" captionTargetBox="[181,1398,205,1872]" captionTargetId="figure@6.[151,1436,193,1923]" captionTargetPageId="6" captionText="FIGURES 5 – 10. 5 – 8, C. bernardoi (Colombia: Anolaima ICAMF 00000044): 5, head, anterodorsal; 6, head, anteroventral; 7, head, dorsal; 8, head, lateral. 9 – 10, C. ornatus (Colombia: Tolima): 9, head, dorsal; 10, head, lateral." httpUri="https://zenodo.org/record/264881/files/figure.png" pageId="13" pageNumber="289">Figs. 5, 6, 8</figureCitation>
). Syntergite 1+2 entirely yellow (
<figureCitation id="136F148C974D50723717F8C88E8A052E" box="[1132,1252,1794,1816]" captionStart="FIGURES 22 – 23. C" captionStartId="9.[151,264,1733,1756]" captionTargetBox="[157,1411,979,1699]" captionTargetId="figure@9.[151,1436,958,1712]" captionTargetPageId="9" captionText="FIGURES 22 – 23. C. bernardoi: 22, female abdomen, dorsal (Colombia: Anolaima ICAMF 00000044); 23, male abdomen, dorsal (Colombia: Guaduas ICAMF 00000045)." httpUri="https://zenodo.org/record/264885/files/figure.png" pageId="13" pageNumber="289">Figs. 22, 23</figureCitation>
). Apical section of vein R4+5 and crossvein dm-m both sinuous (
<figureCitation id="136F148C974D507231F1F8D588A70503" box="[650,713,1823,1845]" captionStart="FIGURES 19 – 21" captionStartId="9.[151,264,855,878]" captionTargetBox="[155,1430,199,829]" captionTargetId="figure@9.[151,1435,193,835]" captionTargetPageId="9" captionText="FIGURES 19 – 21. Wings: 19, C. bernardoi (Colombia: Anolaima ICAMF 00000044); 20, C. ornatus (Colombia: Tolima); 21, C. trinotatus (Colombia: La Mesa ICAMF 00000046)." httpUri="https://zenodo.org/record/264884/files/figure.png" pageId="13" pageNumber="289">Fig.19</figureCitation>
).......................
<taxonomicName id="4C54738A974D50723089F8EA8F350503" authority="Rodriguez &amp; Rodriguez" authorityName="Rodriguez &amp; Rodriguez" box="[1010,1371,1823,1845]" class="Insecta" family="Tephritidae" genus="Cryptodacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="289" phylum="Arthropoda" rank="species" species="bernardoi" status="sp. nov.">
<emphasis id="B920D41B974D50723089F8EA8E030503" box="[1010,1133,1824,1845]" italics="true" pageId="13" pageNumber="289">C. bernardoi</emphasis>
Rodriguez &amp; Rodriguez
</taxonomicName>
,
<taxonomicNameLabel id="A2136960974D5072361DF8D58FF20503" box="[1382,1436,1823,1845]" pageId="13" pageNumber="289" rank="species">
<emphasis id="B920D41B974D5072361DF8D58FF20503" bold="true" box="[1382,1436,1823,1845]" pageId="13" pageNumber="289">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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