treatments-xml/data/12/08/5D/12085D53E4A3319A04A95ADE06883B6C.xml

<document id="F962BF8F44F3AFFC33DD8651E90969DC" ID-CLB-Dataset="35343" ID-DOI="http://dx.doi.org/10.3897/zookeys.329.4961" ID-GBIF-Dataset="1e4ab867-cf67-4105-9918-ebb52be7f061" ID-PMC="PMC3800826" ID-Pensoft-Pub="1313-2970-329-33" ID-PubMed="24146552" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-329-33" ModsDocOrigin="ZooKeys 329" ModsDocTitle="A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species" checkinTime="1451246973851" checkinUser="pensoft" docAuthor="Maier, Crystal A." docDate="2013" docId="12085D53E4A3319A04A95ADE06883B6C" docLanguage="en" docName="ZooKeys 329: 33-91" docOrigin="ZooKeys 329" docSource="http://dx.doi.org/10.3897/zookeys.329.4961" docTitle="Hexanchorus mcdiarmidi Spangler &amp; Staines 2003" docType="treatment" docVersion="5" lastPageNumber="57" masterDocId="307FFFAA5B6E686D5B2FFF86DF568114" masterDocTitle="A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species" masterLastPageNumber="91" masterPageNumber="33" pageNumber="57" updateTime="1732672090439" updateUser="ExternalLinkService">
<mods:mods id="D3D5EDEEF2BB550CDF816B7769753375" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C470D7894091872BC6228E7189F43B06">
<mods:title id="AC83967CAA8AFB708936725E31B81F64">A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species</mods:title>
</mods:titleInfo>
<mods:name id="AA51D78124BBC2B138B0E3F7C1FFBF72" type="personal">
<mods:role id="F42041FEFF30232679F9079E88EF46C0">
<mods:roleTerm id="5051F47E856FF634F234DE672DB0D5BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="853CD2BFD3CD17DC00126071AABF6923">Maier, Crystal A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F8C316D79C1CE38842D5031D5E8EB69E">text</mods:typeOfResource>
<mods:relatedItem id="FCE293117A85B043C56601F2D5F7CE7F" type="host">
<mods:titleInfo id="F3AA56840A91BC99E588094A370C753E">
<mods:title id="8A68F28523AB00A03890F2159E515127">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="C8F2737783AAEAF2C036163753653B37">
<mods:date id="97513B9FB527238EBB56657075A83C3B">2013</mods:date>
<mods:detail id="A94152F52EAC0DF03FBFCAC2A46B019D" type="volume">
<mods:number id="744B6B1B7F09E9D8936EA2BE31A6DF79">329</mods:number>
</mods:detail>
<mods:extent id="F25ECAF9DA53208CECC4FB32A152C7E2" unit="page">
<mods:start id="783D91A682578283DA9B152A92E0F974">33</mods:start>
<mods:end id="D42444935498F7D816512A38B563899B">91</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3E67692B3F096BF0EB1DC661126B592C">
<mods:url id="630EFE567C2A8B19AF552048CEAD742E">http://dx.doi.org/10.3897/zookeys.329.4961</mods:url>
</mods:location>
<mods:classification id="F096069A3E3A3F2F056FB95D7AE55809">journal article</mods:classification>
<mods:identifier id="A55DF7BDE098B67EBBEA382832A61D5D" type="DOI">http://dx.doi.org/10.3897/zookeys.329.4961</mods:identifier>
<mods:identifier id="CA73D131CD9126682C62DF0311EE0D42" type="Pensoft-Pub">1313-2970-329-33</mods:identifier>
</mods:mods>
<treatment id="12085D53E4A3319A04A95ADE06883B6C" ID-GBIF-Taxon="152047497" LSID="urn:lsid:plazi:treatment:12085D53E4A3319A04A95ADE06883B6C" httpUri="http://treatment.plazi.org/id/12085D53E4A3319A04A95ADE06883B6C" lastPageId="25" lastPageNumber="57" pageId="24" pageNumber="57" scope_family="Elmidae" scope_order="Coleoptera" scope_subFamily="Larainae">
<subSubSection id="9EC04B5A67603C17B4A3B5186CA145D0" pageId="24" pageNumber="57" type="nomenclature">
<paragraph id="2FCD17D4FF75B9311BFB9C8769229726" pageId="24" pageNumber="57">
<taxonomicName id="132E070518E39C1AB320DFBD69F02774" ID-CoL="3LGJX" LSID="http://species-id.net/wiki/Hexanchorus_mcdiarmidi" authority="Spangler &amp; Staines, 2003" authorityName="Spangler &amp; Staines" authorityYear="2003" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus mcdiarmidi" order="Coleoptera" pageId="24" pageNumber="57" phylum="Arthropoda" rank="species" species="mcdiarmidi">Hexanchorus mcdiarmidi Spangler &amp; Staines, 2003</taxonomicName>
Figs 4, 48, 49
</paragraph>
</subSubSection>
<subSubSection id="2ED2A82F3B341051E60B7203533C2F8A" pageId="24" pageNumber="57" type="">
<paragraph id="C2D1977BB198FAA3A97D4027370A2A3F" pageId="24" pageNumber="57">
For a complete description and illustrations of genitalia, see 
<bibRefCitation id="9761C43A68BB62674E297FAB991837EA" author="Spangler, PJ" journalOrPublisher="Insecta Mundi" pageId="56" pageNumber="89" pagination="45 - 48" title="Three new species of Hexanchorus Sharp, 1882 (Coleoptera: Elmidae: Larainae) from South America." volume="17" year="2003">Spangler and Staines 2003</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A791537D6FFAD3AC5248CC1F959D7E73" pageId="24" pageNumber="57" type="material examined">
<paragraph id="47FC24E7870074CE83E98A10059EE02F" pageId="24" pageNumber="57">Material examined.</paragraph>
<paragraph id="60690BD427A6B75C8951AE77C92495B5" pageId="24" pageNumber="57">
VENEZUELA: Barinas State: &quot;VENEZUELA: Barinas State; 
<geoCoordinate id="B6AA6681A3E8D0C2E487DF8E0EF209AD" direction="north" orientation="latitude" precision="1" value="8.055683">8°03.341'N</geoCoordinate>
, 
<geoCoordinate id="EAF6906F454C28286B65B77A0E5D2174" direction="west" orientation="longitude" precision="1" value="-70.94328">70°56.597'W</geoCoordinate>
, 203m; nr. Quiri, Rio Quiu; 15.vii.2009; leg. W. Shepard; ex. bulk sample; VZ09-0715-03Z&quot; (99 specimens SEMC). Distrito Capital:&quot;VENEZUELA, Dto. Federal; Los Caracas; 19 January 1985; P. Spangler, R. Faitoute; W. Steiner, &amp; A. Conover&quot; (6 paratypes USNM) 
<normalizedToken id="778ACA558FAB0C5386768482818DD5D7" originalValue="Mérida">Merida</normalizedToken>
State: &quot;VENEZUELA: 
<normalizedToken id="EC7F68EE85A022B24912032B4039B34C" originalValue="Mérida">Merida</normalizedToken>
State; 
<geoCoordinate id="13EE6843EBDB40A39EF422CA7582115E" direction="north" orientation="latitude" precision="1" value="8.953417">8°57.205'N</geoCoordinate>
, 
<geoCoordinate id="B54B537B362AB113FF4DEA5391160058" direction="west" orientation="longitude" precision="1" value="-71.29366">71°17.620'W</geoCoordinate>
, 88m; W. of Tucani; 21.vii2009; Gross sample from river; leg. Shepard; VZ09-0721-04Z&quot; (17 specimens SEMC). Zulia State: &quot;VENEZUELA: Zulia State; 
<geoCoordinate id="3055E8C769B2B3EDFF745F46A32131A5" direction="north" orientation="latitude" precision="1" value="9.841884">9°50.513'N</geoCoordinate>
, 
<geoCoordinate id="CF67D46F104B7B1F3F17795D18522527" direction="west" orientation="longitude" precision="1" value="-72.805565">72°48.334'W</geoCoordinate>
. 252m; Perija N.P. Tukuko: Rio Tukuko; 5.vii.2009; leg. Short &amp; Gustafson; riffle/rocks in river; VZ09-0705-01B&quot; (96 specimens SEMC). &quot;VENEZUELA: Zulia State; P.N. Perija: Tukuko; 18 vii.2008; Rio Marpito&quot; &quot;Andrew E. Z.; Short, leg.&quot; (5 specimens SEMC). &quot;VENEZUELA: Zulia El; Tucuco (51 km S.O. de; Machiques); 24-VI-1992&quot; &quot;Colector:; J. Camacho&quot; (11 specimens MALUZ).
</paragraph>
</subSubSection>
<subSubSection id="C13F4B0003BD280990485C0B974B0DCA" pageId="24" pageNumber="57" type="diagnosis">
<paragraph id="F296AA00D8B9A3F314B14D46C31BF721" pageId="24" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="74F40A4A4C7CEE0D5A26085D8961C929" pageId="24" pageNumber="57">
<taxonomicName id="9DE9115B9E56B00406428F6EE5703D74" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus mcdiarmidi" order="Coleoptera" pageId="24" pageNumber="57" phylum="Arthropoda" rank="species" species="mcdiarmidi">Hexanchorus mcdiarmidi</taxonomicName>
can be disinguished from all other 
<taxonomicName id="C8C50F85C0FC60BAD936886DCFABB724" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="24" pageNumber="57" phylum="Arthropoda" rank="genus">Hexanchorus</taxonomicName>
species in Venezuela by the presence of a greenish iridescent sheen on the dorsal setae (Fig. 48), the narrow parameres, and pronotum with only a shallow postero-median impression (Fig. 48).
</paragraph>
</subSubSection>
<subSubSection id="5498078599FDB6DCB108AD8A5BCFA0EA" pageId="24" pageNumber="57" type="intraspecific variation">
<paragraph id="59E116F84024359306F764EC40B593A1" pageId="24" pageNumber="57">Intraspecific variation.</paragraph>
<paragraph id="6CF21C9EDDD18042DC18FCC00E889278" pageId="24" pageNumber="57">
This species varies slightly from black to dark brown in color, additionally, there may be slight variation in the greenish iridescence of the elytral setae. 
<taxonomicName id="0206B9731C4A625E4A2E48989863E9B0" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus mcdiarmidi" order="Coleoptera" pageId="24" pageNumber="57" phylum="Arthropoda" rank="species" species="mcdiarmidi">Hexanchorus mcdiarmidi</taxonomicName>
has a purplish and orange iridescent sheen while submerged in alcohol (Short, pers. comm.). There is variation in size (2.9-3.4 mm TL) and slight differences in punctation among specimens.
</paragraph>
</subSubSection>
<subSubSection id="4815099E3CF8B8572DAE9D2A56767CF4" pageId="24" pageNumber="57" type="habitat and distribution">
<paragraph id="8F4AB22F459FA69A9DF66752C5919240" pageId="24" pageNumber="57">Habitat and distribution.</paragraph>
<paragraph id="904020B3E487CB9A28C0DB7709276658" pageId="24" pageNumber="57">
<taxonomicName id="28E319A3F19959FF05C27A3A54C617DF" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus mcdiarmidi" order="Coleoptera" pageId="24" pageNumber="57" phylum="Arthropoda" rank="species" species="mcdiarmidi">Hexanchorus mcdiarmidi</taxonomicName>
is widespread in lower altitude (&lt;300 m elevation) streams throughout western Venezuela (Fig. 4). They can be found in dense aggregations on water-splashed emergent rocks in small to medium sized swift-flowing streams and rivers with rocky substrate.
</paragraph>
</subSubSection>
<subSubSection id="6414F298B17047D9B7856780E203B5F4" lastPageId="25" lastPageNumber="58" pageId="24" pageNumber="57" type="associated species">
<paragraph id="E0F415633F5DFFF3CE6ED0CA77D97F70" pageId="24" pageNumber="57">Associated species.</paragraph>
<paragraph id="EEE8A8EFC3654230C2FEDFE6C87A9FAD" lastPageId="25" lastPageNumber="58" pageId="24" pageNumber="57">
While no other species of 
<taxonomicName id="F3547CF7592820061C44F6F02A9BF5E7" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="genus">
<pageBreakToken id="EE07959ACCC84203578232A9732BB434" pageId="25" pageNumber="58" start="start">Hexanchorus</pageBreakToken>
</taxonomicName>
were collected at the same localities as 
<taxonomicName id="315AF3B592BBE36D46097024F484E303" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus mcdiarmidi" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="species" species="mcdiarmidi">Hexanchorus mcdiarmidi</taxonomicName>
, other laraines collected in the same habitats include 
<taxonomicName id="A305771A9F9330F5D64652C3E8101642" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Disersus dasycolus" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="species" species="dasycolus">Disersus dasycolus</taxonomicName>
, 
<taxonomicName id="1AA7F79B04C1F77A7D050789C75BE106" class="Insecta" family="Elmidae" genus="Phanocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanocerus congener" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="species" species="congener">Phanocerus congener</taxonomicName>
and 
<taxonomicName id="B303F1E47740CFE5B3E7C2C359A6D4D2" class="Insecta" family="Elmidae" genus="Phanocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanocerus clavicornis" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="species" species="clavicornis">Phanocerus clavicornis</taxonomicName>
, and 
<taxonomicName id="22F0652C253F3516420E35007ADF1970" class="Insecta" family="Elmidae" genus="Pharceonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pharceonus volcanus" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="species" species="volcanus">Pharceonus volcanus</taxonomicName>
. Other water beetles collected at the same sites include: 
<taxonomicName id="72FBBEBB7FB630E643F87464E119EA5A" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="genus">Anacaena</taxonomicName>
spp. and 
<taxonomicName id="A4C5F044DCF88D3C27941747657386BB" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="genus">Enochrus</taxonomicName>
spp. (
<taxonomicName id="21090E0F1931B63A11877D2C956F6104" class="Insecta" family="Hydrophilidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="58" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>