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<document id="289B95529CD712036F1BCAFDE9BB4362" ID-CLB-Dataset="9005" ID-DOI="10.11646/zootaxa.3798.1.1" ID-GBIF-Dataset="f99f3b15-7592-490f-9903-ce6b24131864" ID-ISSN="1175-5326" ID-Zenodo-Dep="4431865" ID-ZooBank="CB8534BA-89CB-44A6-81E3-3A8927055C7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610352857687" checkinUser="plazi" docAuthor="THOMA, MARCO, KRANZ-BALTENSPERGER, YVONNE, KROPF, CHRISTIAN, GRABER, WERNER, NENTWIG, WOLFGANG &amp; FRICK, HOLGER" docDate="2014" docId="094F426DFFA2FFE86FC9F43D814FFB44" docLanguage="en" docName="zootaxa.3798.1.1.pdf" docOrigin="Zootaxa 3798 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3798.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aposphragisma monoceros Thoma 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="F5763A15FF96FFDD6F5EF2608257FFAC" masterDocTitle="The new Southeast Asian goblin spider genus Aposphragisma (Araneae, Oonopidae): diversity and phylogeny" masterLastPageNumber="86" masterPageNumber="1" pageNumber="53" updateTime="1698924184113" updateUser="ExternalLinkService">
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<mods:titleInfo id="0A9D1D5247253B9EAD6DC7603D354B8E">
<mods:title id="FD5264A3294D01C6CCC48125B0B16AAC">The new Southeast Asian goblin spider genus Aposphragisma (Araneae, Oonopidae): diversity and phylogeny</mods:title>
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<mods:namePart id="8973ED3ECCF669B530F24D2039A440B2">THOMA, MARCO</mods:namePart>
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<mods:namePart id="F0300B727BC71B03A3C4706BF2FCFBA4">KRANZ-BALTENSPERGER, YVONNE</mods:namePart>
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<mods:namePart id="AAB5F70130224444D2ED3A8DA5965A58">KROPF, CHRISTIAN</mods:namePart>
</mods:name>
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<mods:roleTerm id="24C7B76293DE4CE5D3B4EBB7ACAEC2E3">Author</mods:roleTerm>
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<mods:namePart id="EAD76953DFAB36CB9E9F27234DB99DBE">GRABER, WERNER</mods:namePart>
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<mods:namePart id="CE65EFF968A351F8F7769684440B9752">NENTWIG, WOLFGANG</mods:namePart>
</mods:name>
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<mods:namePart id="FF6BA82A3444FF92F9637F646B93D88D">FRICK, HOLGER</mods:namePart>
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<mods:title id="4183E34C0E19410FC3929973E4AD8359">Zootaxa</mods:title>
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<mods:date id="501D1CC64058970122798F726ACF135B">2014</mods:date>
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<mods:number id="6C1A3471B0574DEE8D598102C5053B84">2014-05-22</mods:number>
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<mods:number id="AD911D01E981379C3D73FAB7C97A653D">3798</mods:number>
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<treatment id="094F426DFFA2FFE86FC9F43D814FFB44" ID-DOI="http://doi.org/10.5281/zenodo.4431921" ID-GBIF-Taxon="176566791" ID-Zenodo-Dep="4431921" LSID="urn:lsid:plazi:treatment:094F426DFFA2FFE86FC9F43D814FFB44" httpUri="http://treatment.plazi.org/id/094F426DFFA2FFE86FC9F43D814FFB44" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection id="C9FCA0F0FFA2FFE96FC9F43D80FBF9DB" box="[151,684,1629,1655]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8159F37BFFA2FFE96FC9F43D80FBF9DB" blockId="52.[151,684,1629,1689]" box="[151,684,1629,1655]" pageId="52" pageNumber="53">
<heading id="DA114417FFA2FFE96FC9F43D80FBF9DB" bold="true" box="[151,684,1629,1655]" fontSize="11" level="2" pageId="52" pageNumber="53" reason="4">
<emphasis id="B3922F69FFA2FFE96FC9F43D80FBF9DB" bold="true" box="[151,684,1629,1655]" pageId="52" pageNumber="53">
<taxonomicName id="46E688F8FFA2FFE96FC9F43D8016F9DB" ID-CoL="682PJ" authority="Thoma" authorityName="Thoma" authorityYear="2014" box="[151,577,1629,1655]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="monoceros" status="sp. nov.">
<emphasis id="B3922F69FFA2FFE96FC9F43D8388F9DB" bold="true" box="[151,479,1629,1655]" italics="true" pageId="52" pageNumber="53">Aposphragisma monoceros</emphasis>
Thoma
</taxonomicName>
,
<taxonomicNameLabel id="A8A19212FFA2FFE96D10F43D80FBF9DB" box="[590,684,1629,1655]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C9FCA0F0FFA2FFE96FC9F4E08379F935" box="[151,302,1664,1689]" pageId="52" pageNumber="53" type="description">
<paragraph id="8159F37BFFA2FFE96FC9F4E08379F935" blockId="52.[151,684,1629,1689]" box="[151,302,1664,1689]" pageId="52" pageNumber="53">
(
<figureCitation id="19DDEFFEFFA2FFE96FC1F4E08371F935" box="[159,294,1664,1689]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStartId-0="51.[151,250,1875,1897]" captionStartId-1="52.[151,250,1445,1467]" captionTargetBox-0="[184,1406,228,1819]" captionTargetBox-1="[186,1404,221,1407]" captionTargetId-0="figure-75@51.[151,1436,193,1854]" captionTargetId-1="figure-226@52.[151,1436,193,1424]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionText-0="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." captionText-1="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4431969" figureDoi-1="http://doi.org/10.5281/zenodo.4431971" httpUri-0="https://zenodo.org/record/4431969/files/figure.png" httpUri-1="https://zenodo.org/record/4431971/files/figure.png" pageId="52" pageNumber="53">Figs. 32–33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9FCA0F0FFA2FFE96FC9F4A880F8F8DD" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8159F37BFFA2FFE96FC9F4A880F8F8DD" blockId="52.[151,1436,1736,2013]" pageId="52" pageNumber="53">
<emphasis id="B3922F69FFA2FFE96FC9F4A88312F94D" bold="true" box="[151,325,1736,1761]" pageId="52" pageNumber="53">Type material.</emphasis>
<materialsCitation id="318EF926FFA2FFE96E13F4A883BCF8E1" ID-GBIF-Occurrence="3014709378" collectingDate="2009-10-01" collectionCode="CAS" collectorName="C. Griswold &amp; N. Chousou Polydouri" country="Brunei" elevation="150" latitude="4.5418835" location="Ashton Trail" longLatPrecision="15" longitude="115.155" municipality="Kuala Belalong Field Studies Centre" pageId="52" pageNumber="53" specimenCode="PBI_OON 00036324, UT003, CASENT 9029917" specimenCount="1" stateProvince="Temburong District" typeStatus="holotype">
<emphasis id="B3922F69FFA2FFE96E13F4A8838EF94D" bold="true" box="[333,473,1736,1761]" pageId="52" pageNumber="53">
<typeStatus id="5E5D4DD9FFA2FFE96E13F4A88387F973" box="[333,464,1736,1761]" pageId="52" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
<specimenCount id="97E038F2FFA2FFE96EBFF4A88040F94D" box="[481,535,1736,1761]" pageId="52" pageNumber="53" type="male">male</specimenCount>
(
<specimenCode id="D1405B00FFA2FFE96D79F4A8814BF94D" box="[551,796,1736,1761]" pageId="52" pageNumber="53">PBI_OON 00036324</specimenCode>
),
<collectingRegion id="43223D99FFA2FFE96C71F4A8865CF94D" box="[815,1035,1736,1761]" country="Brunei" name="Temburong" pageId="52" pageNumber="53">Temburong District</collectingRegion>
(
<collectingCountry id="F9F1B3EBFFA2FFE96B42F4A88624F973" box="[1052,1139,1736,1761]" name="Brunei" pageId="52" pageNumber="53">BRUNEI</collectingCountry>
),
<location id="8439A5A0FFA2FFE96BD5F4A8874EF94D" LSID="urn:lsid:plazi:treatment:094F426DFFA2FFE86FC9F43D814FFB44:8439A5A0FFA2FFE96BD5F4A8874EF94D" box="[1163,1305,1736,1761]" country="Brunei" latitude="4.5418835" longLatPrecision="15" longitude="115.155" municipality="Kuala Belalong Field Studies Centre" name="Ashton Trail" pageId="52" pageNumber="53" stateProvince="Temburong District">Ashton Trail</location>
<location id="8439A5A0FFA2FFE96A7FF4A983A2F8A9" LSID="urn:lsid:plazi:treatment:094F426DFFA2FFE86FC9F43D814FFB44:8439A5A0FFA2FFE96A7FF4A983A2F8A9" country="Brunei" latitude="4.5418835" longLatPrecision="15" longitude="115.155" municipality="Kuala Belalong Field Studies Centre" name="near Kuala Belalong Field Studies Centre" pageId="52" pageNumber="53" stateProvince="Temburong District">near Kuala Belalong Field Studies Centre</location>
,
<locationDeviation id="90EBEA59FFA2FFE96D5BF48C8142F8A8" box="[517,789,1772,1797]" location="Bangar" pageId="52" pageNumber="53">
<quantity id="461E5E9EFFA2FFE96D5BF48C8007F8A9" box="[517,592,1772,1797]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="52" pageNumber="53" unit="km" value="21.0">21 km</quantity>
SSW of Bangar
</locationDeviation>
,
<quantity id="461E5E9EFFA2FFE96C7AF48C8138F8AF" box="[804,879,1772,1797]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="52" pageNumber="53" unit="m" value="150.0">
<elevation id="0ACB1448FFA2FFE96C7AF48C8138F8AF" box="[804,879,1772,1797]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="52" pageNumber="53" unit="m" value="150.0">150 m</elevation>
</quantity>
[
<geoCoordinate id="E4D295BCFFA2FFE96CDEF48C867CF8AF" box="[896,1067,1772,1797]" degrees="04" direction="north" minutes="32" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="30.78" value="4.5418835">04°32′30.78′′N</geoCoordinate>
<geoCoordinate id="E4D295BCFFA2FFE96B68F48C869EF8AF" box="[1078,1225,1772,1797]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="52" pageNumber="53" precision="15" seconds="18" value="115.155">115°09′18′′E</geoCoordinate>
; prov.],
<date id="F558D5BBFFA2FFE96A73F48C87CFF8A9" box="[1325,1432,1772,1797]" pageId="52" pageNumber="53" value="2009-10-01">
<collectingDate id="E51C2C53FFA2FFE96A73F48C87CFF8A9" box="[1325,1432,1772,1797]" pageId="52" pageNumber="53" value="2009-10-01">1.X.2009</collectingDate>
</date>
, primary mixed dipterocarp forest, sifting leaf litter, leg.
<collectorName id="2C1396ADFFA2FFE96C49F57081F6F885" box="[791,929,1808,1833]" pageId="52" pageNumber="53">C. Griswold</collectorName>
&amp;
<collectorName id="2C1396ADFFA2FFE96C97F5718692F885" box="[969,1221,1808,1833]" pageId="52" pageNumber="53">N. Chousou Polydouri</collectorName>
, deposited at
<collectionCode id="E7F76BBEFFA2FFE96A3BF57087CBF885" box="[1381,1436,1808,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CAS</collectionCode>
(
<specimenCode id="D1405B00FFA2FFE96FC1F55482A5F8E1" box="[159,242,1844,1869]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">UT003</specimenCode>
,
<specimenCode id="D1405B00FFA2FFE96E5DF55483B4F8E1" box="[259,483,1844,1869]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CASENT 9029917</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="318EF926FFA2FFE96EA5F55480FBF8DD" ID-GBIF-Occurrence="3014709379" collectingDate="2009-10-01" collectionCode="CAS" collectorName="C. Griswold &amp; N. Chousou Polydouri" country="Brunei" elevation="150" latitude="4.5418835" location="Ashton Trail" longLatPrecision="15" longitude="115.155" municipality="Kuala Belalong Field Studies Centre" pageId="52" pageNumber="53" specimenCode="PBI_OON 00032178, UT004, CASENT 9036043" specimenCount="2" specimenCount-male="2" stateProvince="Temburong" typeStatus="paratype">
<emphasis id="B3922F69FFA2FFE96EA5F55480DAF8E1" bold="true" box="[507,653,1844,1869]" pageId="52" pageNumber="53">
<typeStatus id="5E5D4DD9FFA2FFE96EA5F55480D2F8E7" box="[507,645,1844,1869]" pageId="52" pageNumber="53" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
<specimenCount id="97E038F2FFA2FFE96DC4F55480A2F8E1" box="[666,757,1844,1869]" pageId="52" pageNumber="53" type="male">2 males</specimenCount>
(
<specimenCode id="D1405B00FFA2FFE96C54F5548654F8E1" box="[778,1027,1844,1869]" pageId="52" pageNumber="53">PBI_OON 00032178</specimenCode>
), collected together with holotype, deposited at
<collectionCode id="E7F76BBEFFA2FFE96E7AF538830CF8DD" box="[292,347,1880,1905]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CAS</collectionCode>
(
<specimenCode id="D1405B00FFA2FFE96E37F53883EAF8DD" box="[361,445,1880,1905]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">UT004</specimenCode>
,
<specimenCode id="D1405B00FFA2FFE96E96F53880F4F8DD" box="[456,675,1880,1905]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CASENT 9036043</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FCA0F0FFA2FFE96F99F51C816DF814" pageId="52" pageNumber="53" type="etymology">
<paragraph id="8159F37BFFA2FFE96F99F51C816DF814" blockId="52.[151,1436,1736,2013]" pageId="52" pageNumber="53">
<emphasis id="B3922F69FFA2FFE96F99F51C8319F839" bold="true" box="[199,334,1916,1941]" pageId="52" pageNumber="53">Etymology.</emphasis>
The species epithet, a noun in apposition, is a Greek term meaning ‘unicorn’. It refers to the single ‘horn’ the species bears on the posterior part of its carapace.
</paragraph>
</subSubSection>
<subSubSection id="C9FCA0F0FFA2FFE86F99F5A482E0FEB1" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8159F37BFFA2FFE86F99F5A482E0FEB1" blockId="52.[151,1436,1736,2013]" lastBlockId="53.[151,1437,151,1256]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="B3922F69FFA2FFE96F99F5A48316F871" bold="true" box="[199,321,1988,2013]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
<taxonomicName id="46E688F8FFA2FFE96E14F5A483B2F870" authorityName="Thoma" authorityYear="2014" box="[330,485,1988,2012]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="monoceros" status="sp. nov.">
<emphasis id="B3922F69FFA2FFE96E14F5A483B2F870" box="[330,485,1988,2012]" italics="true" pageId="52" pageNumber="53">A. monoceros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8A19212FFA2FFE96EB0F5A48011F877" box="[494,582,1988,2011]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the stripe-clade and can be distinguished from other species of this clade by the combination of the following traits: carapace with one pair of lateral double-toothed spikes (dts) (
<figureCitation id="19DDEFFEFFA3FFE86FC1F2DC82A9FF79" box="[159,254,188,213]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32</figureCitation>
D–E); carapace margin with pointed denticles, sluice restricted to posterior margin (
<figureCitation id="19DDEFFEFFA3FFE86B9DF2DC8779FF79" box="[1219,1326,188,213]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32D</figureCitation>
). It is the only species of the stripe-clade possessing a single ‘horn’ (ho) at the posterior edge of pars cephalica (
<figureCitation id="19DDEFFEFFA3FFE86A7AF2BF82FDFEB1" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32D, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9FCA0F0FFA3FFE86F99F3478169FB69" pageId="53" pageNumber="54" type="description">
<paragraph id="8159F37BFFA3FFE86F99F34780E7FEEC" blockId="53.[151,1437,151,1256]" box="[199,688,295,320]" pageId="53" pageNumber="54">
<emphasis id="B3922F69FFA3FFE86F99F347830EFEEC" bold="true" box="[199,345,295,320]" pageId="53" pageNumber="54">Description.</emphasis>
Description based on
<specimenCount id="97E038F2FFA3FFE86D0CF34780FBFEEC" box="[594,684,295,320]" pageId="53" pageNumber="54" type="male">3 males</specimenCount>
.
</paragraph>
<paragraph id="8159F37BFFA3FFE86F99F32C83C3FE7C" blockId="53.[151,1437,151,1256]" pageId="53" pageNumber="54">
<emphasis id="B3922F69FFA3FFE86F99F32C8341FEC9" bold="true" box="[199,278,332,357]" pageId="53" pageNumber="54">MALE.</emphasis>
With the characters of the genus except as noted. Body length
<quantity id="461E5E9EFFA3FFE86CB1F32C86C3FECF" box="[1007,1172,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.685" metricValueMax="1.71" metricValueMin="1.66" pageId="53" pageNumber="54" unit="mm" value="1.685" valueMax="1.71" valueMin="1.66">1.66–1.71 mm</quantity>
(n=3), carapace length
<quantity id="461E5E9EFFA3FFE86FC9F30F82ACFE2B" box="[151,251,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="53" pageNumber="54" unit="mm" value="0.79">0.79 mm</quantity>
(n=3). Sclerotized parts uniformly coloured orange brown, close to colour 36 ‘amber’; legs slightly paler than body. Setae especially on abdominal scutae slightly longer and darker if compared to most congeners. Habitus:
<figureCitation id="19DDEFFEFFA3FFE86FA3F3D78300FE7C" box="[253,343,439,464]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32</figureCitation>
A–C.
</paragraph>
<paragraph id="8159F37BFFA3FFE86F99F3BC8716FC94" blockId="53.[151,1437,151,1256]" pageId="53" pageNumber="54">
CEPHALOTHORAX. Carapace: surface of elevated portion of pars cephalica smooth, sides strongly reticulate, partly interrupted by small smooth areas (
<figureCitation id="19DDEFFEFFA3FFE86D39F39F809DFDB4" box="[615,714,511,536]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32F</figureCitation>
); posterior edge of pars cephalica with a single ‘horn’ (ho) (
<figureCitation id="19DDEFFEFFA3FFE86A3AF39F82BFFD91" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32D, F</figureCitation>
); carapace margin with pointed denticles, sluice restricted to posterior margin (
<figureCitation id="19DDEFFEFFA3FFE86B00F04486EEFD91" box="[1118,1209,548,573]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32</figureCitation>
D–E), posterolateral corners with a pair of double-toothed spikes (dts) (
<figureCitation id="19DDEFFEFFA3FFE86DB8F0278127FDCC" box="[742,880,583,608]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32D, E</figureCitation>
). Eyes: posterior eye-row recurved from above, procurved from front; ALE largest, separated by their radius to diameter; ALE-PLE touching; PME touching throughout most of their length; PLE-PME touching (
<figureCitation id="19DDEFFEFFA3FFE86DA8F0EF81DAFD04" box="[758,909,655,680]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Figs. 32D, G</figureCitation>
). Sternum coarsely ornamented except smooth median stripe and broadly smooth edges (ornamentation arranged in an ovoid band;
<figureCitation id="19DDEFFEFFA3FFE86B3DF0D48690FD61" box="[1123,1223,692,717]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32E</figureCitation>
); posterior margin with a rather narrow single extension, covered with few blunt denticles (
<figureCitation id="19DDEFFEFFA3FFE86C83F0B78611FD5C" box="[989,1094,727,752]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32E</figureCitation>
). Pleura: surface smooth with pairs of large pits dorsally of each coxa I, II and III (as in
<taxonomicName id="46E688F8FFA3FFE86C19F09C8651FCB9" authorityName="Thoma" authorityYear="2014" box="[839,1030,764,789]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="brunomanseri" status="sp. nov.">
<emphasis id="B3922F69FFA3FFE86C19F09C8651FCB9" box="[839,1030,764,789]" italics="true" pageId="53" pageNumber="54">A. brunomanseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8A19212FFA3FFE86B4EF09C863BFCBF" box="[1040,1132,764,787]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
, cf fig. 10B). Mouthparts: chelicerae promargin distally extending into a short inwards pointing tooth-like projection (tlp) (
<figureCitation id="19DDEFFEFFA3FFE86B90F17F8763FC94" box="[1230,1332,799,824]" captionStart="FIGURE 32" captionStartId="51.[151,250,1875,1897]" captionTargetBox="[184,1406,228,1819]" captionTargetId="figure-75@51.[151,1436,193,1854]" captionTargetPageId="51" captionText="FIGURE 32. Aposphragisma monoceros sp. nov. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. carapace dorsal view; E. prosoma ventral view; F. carapace lateral view; G. carapace front view. Dts, double-toothed spike; tlp, tooth-like projection of male chelicerae; ho, horn (modified hair-base). Scale bars: A–G 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4431969" httpUri="https://zenodo.org/record/4431969/files/figure.png" pageId="53" pageNumber="54">Fig. 32E</figureCitation>
).
</paragraph>
<paragraph id="8159F37BFFA3FFE86F99F124872CFC64" blockId="53.[151,1437,151,1256]" pageId="53" pageNumber="54">
ABDOMEN. Book lung covers large, elongated elliptical (sides slightly concave), about 3 times longer than wide, darker in colour than surroundings (
<figureCitation id="19DDEFFEFFA3FFE86D35F1078084FC2C" box="[619,723,871,896]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33A</figureCitation>
). Epigastric scutum dorsally with widely oval subterminal ridge (sr) (
<figureCitation id="19DDEFFEFFA3FFE86F90F1EC8361FC09" box="[206,310,908,933]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33A</figureCitation>
, partly covered by dorsal abdominal scutum in this case). Postepigastric scutum long, semicircular, with short posteriorly directed lateral apodemes (lap) (
<figureCitation id="19DDEFFEFFA3FFE86DA6F1CF8108FC64" box="[760,863,943,968]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33B</figureCitation>
). Dorsal abdominal setae very long and darkish.
</paragraph>
<paragraph id="8159F37BFFA3FFE86F99F1B48064FBD1" blockId="53.[151,1437,151,1256]" pageId="53" pageNumber="54">
GENITALIA. Epigastric region: sperm pore (sp) situated at level of posterior spiracles (
<figureCitation id="19DDEFFEFFA3FFE86BD8F1B48681FC41" box="[1158,1238,980,1005]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33</figureCitation>
A–B). Palp: basal segments same colour as body but bulb and cymbium much lighter (pale orange); bulb rather stout, tapering apically (
<figureCitation id="19DDEFFEFFA3FFE86FA1F67C830CFB99" box="[255,347,1052,1077]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Figs. 33</figureCitation>
C–D). Embolus (em) tip spatulate (
<figureCitation id="19DDEFFEFFA3FFE86DB7F67C8127FB99" box="[745,880,1052,1077]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33C, E</figureCitation>
); embolic spine (esp) with fringed tip (
<figureCitation id="19DDEFFEFFA3FFE86A74F67C87DAFB99" box="[1322,1421,1052,1077]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33E</figureCitation>
). Conductor (con) with broad tip (appearing ‘snout’-like in lateral view;
<figureCitation id="19DDEFFEFFA3FFE86C89F65F8617FBF4" box="[983,1088,1087,1112]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Fig. 33E</figureCitation>
). Embolus-conductor-complex about as long as bulb (
<figureCitation id="19DDEFFEFFA3FFE86ECDF60483B9FBD1" box="[403,494,1124,1149]" captionStart="FIGURE 33" captionStartId="52.[151,250,1445,1467]" captionTargetBox="[186,1404,221,1407]" captionTargetId="figure-226@52.[151,1436,193,1424]" captionTargetPageId="52" captionText="FIGURE 33. Aposphragisma monoceros sp. nov. Male: A. abdomen lateral view; B. abdomen ventral view; C–D. palp prolateral view (D: hairs omitted except trichobothria); E. embolus-conductor-complex prolateral view. Cb, conical bulge; con, conductor; dc, dorsal comb; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; lc, lateral comb; sp, sperm pore; spp, spermophor; sr, subterminal widely oval scutal ridge. Scale bars: A–B 500 µm; C–D 100 µm; E 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4431971" httpUri="https://zenodo.org/record/4431971/files/figure.png" pageId="53" pageNumber="54">Figs. 33</figureCitation>
C–D).
</paragraph>
<paragraph id="8159F37BFFA3FFE86F99F6E78169FB69" blockId="53.[151,1437,151,1256]" pageId="53" pageNumber="54">
<emphasis id="B3922F69FFA3FFE86F99F6E7838EFB0C" bold="true" box="[199,473,1159,1184]" pageId="53" pageNumber="54">Intraspecific variation.</emphasis>
Body size variable (see above). Number of denticles on carapace margin varies. One
<typeStatus id="5E5D4DD9FFA3FFE86FC9F6CC82A0FB68" box="[151,247,1196,1220]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
male much lighter in colour, yellow to pale-orange.
</paragraph>
</subSubSection>
<subSubSection id="C9FCA0F0FFA3FFE86F99F6AF814FFB44" box="[199,792,1231,1256]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8159F37BFFA3FFE86F99F6AF814FFB44" blockId="53.[151,1437,151,1256]" box="[199,792,1231,1256]" pageId="53" pageNumber="54">
<emphasis id="B3922F69FFA3FFE86F99F6AF8337FB44" bold="true" box="[199,352,1231,1256]" pageId="53" pageNumber="54">Distribution.</emphasis>
<collectingRegion id="43223D99FFA3FFE86E39F6AF8011FB44" box="[359,582,1231,1256]" country="Brunei" name="Temburong" pageId="53" pageNumber="54">Temburong District</collectingRegion>
,
<collectingCountry id="F9F1B3EBFFA3FFE86D0FF6B080F0FB4B" box="[593,679,1232,1255]" name="Brunei" pageId="53" pageNumber="54">BRUNEI</collectingCountry>
(
<figureCitation id="19DDEFFEFFA3FFE86DE6F6AF815FFB44" box="[696,776,1231,1256]" captionStart="FIGURE 49" captionStartId="73.[151,250,1482,1504]" captionTargetBox="[173,1409,203,1453]" captionTargetId="figure-249@73.[155,1427,185,1464]" captionTargetPageId="73" captionText="FIGURE 49. Distribution of Aposphragisma gen. nov. in Southeast Asia. Stripe clade: 1) A. baltenspergerae sp. nov., 2) A. borgulai sp. nov., 3) A. brunomanseri sp. nov., 4) A. dayak sp. nov., 5) A. dentatum sp. nov., 6) A. hausammannae sp. nov., 7) A. helvetiorum sp. nov., 8) A. kolleri sp. nov., 9) A. menzi sp. nov., 10) A. monoceros sp. nov., 11) A. retifer sp. nov., 12) A. salewskii sp. nov., 13) A. sepilok sp. nov. Non-stripe clade species: 14) A. confluens sp. nov., 15) A. draconigenum sp. nov., 16) A. nocturnum sp. nov., 17) A. rimba sp. nov., 18) A. scimitar sp. nov., 19) A. stannum sp. nov. Scale bars: 500 km." figureDoi="http://doi.org/10.5281/zenodo.4432005" httpUri="https://zenodo.org/record/4432005/files/figure.png" pageId="53" pageNumber="54">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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