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<document id="BBBC695FBE1EEB2D1E891D8728286622" ID-CLB-Dataset="46667" ID-DOI="10.11646/zootaxa.3754.3.2" ID-GBIF-Dataset="fe375541-8083-43f8-9cf8-b8d75af79599" ID-ISSN="1175-5326" ID-Zenodo-Dep="251551" ID-ZooBank="9E645014-5464-4E7C-8D4A-0B3B52A5AA53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460368973748" checkinUser="plazi" docAuthor="Kantor, Yuri, Lozouet, Pierre, Puillandre, Nicolas &amp; Bouchet, Philippe" docDate="2014" docId="0F4E87BB7B3BFFB6FF22FB71A790FA81" docLanguage="en" docName="zt03754p276.pdf" docOrigin="Zootaxa 3754 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hortia Lozouet 1999" docType="treatment" docVersion="10" lastPageNumber="250" masterDocId="F377FFC37B31FFBDFFB5FFADA776FFE4" masterDocTitle="Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific" masterLastPageNumber="276" masterPageNumber="239" pageNumber="249" updateTime="1698295552789" updateUser="ExternalLinkService">
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<mods:title id="CFDDD7005796FE6ED4D769E88109D7D0">Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific</mods:title>
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<mods:namePart id="75AD2DFDA898E021EA6502AB58ABED23">Kantor, Yuri</mods:namePart>
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<mods:namePart id="630F0295BF1B0F40A9B476777091A25F">Lozouet, Pierre</mods:namePart>
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<mods:namePart id="06EE636D5E2FD4C81641EDDE7C27D58D">Puillandre, Nicolas</mods:namePart>
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<mods:namePart id="32D76C25922515A255B315128D8A147E">Bouchet, Philippe</mods:namePart>
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<mods:title id="999288ECA1B8BD7F3E1CC697AF5293C9">Zootaxa</mods:title>
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<mods:date id="49398388F1160DB2157931859F2D29B4">2014</mods:date>
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<mods:identifier id="0BB9308B1A573A7056C97572F9CC26A2" type="DOI">10.11646/zootaxa.3754.3.2</mods:identifier>
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<treatment id="0F4E87BB7B3BFFB6FF22FB71A790FA81" ID-DOI="http://doi.org/10.5281/zenodo.5676762" ID-GBIF-Taxon="119411397" ID-Zenodo-Dep="5676762" LSID="urn:lsid:plazi:treatment:0F4E87BB7B3BFFB6FF22FB71A790FA81" httpUri="http://treatment.plazi.org/id/0F4E87BB7B3BFFB6FF22FB71A790FA81" lastPageId="11" lastPageNumber="250" pageId="10" pageNumber="249">
<subSubSection id="CFFD65267B3BFFB7FF22FB71A626FADC" pageId="10" pageNumber="249" type="nomenclature">
<paragraph id="875836AD7B3BFFB7FF22FB71A685FB12" blockId="10.[151,499,1244,1270]" box="[151,499,1244,1270]" pageId="10" pageNumber="249">
<heading id="DC1081C17B3BFFB7FF22FB71A685FB12" bold="true" box="[151,499,1244,1270]" fontSize="11" level="1" pageId="10" pageNumber="249" reason="1">
<emphasis id="B593EABF7B3BFFB7FF22FB71A685FB12" bold="true" box="[151,499,1244,1270]" pageId="10" pageNumber="249">
Genus
<taxonomicName id="40E74D2E7B3BFFB7FF58FB71A685FB12" ID-CoL="4YW2" authority="Lozouet, 1999" authorityName="Lozouet" authorityYear="1999" box="[237,499,1244,1270]" class="Magnoliopsida" family="Rutaceae" genus="Hortia" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="genus">
<emphasis id="B593EABF7B3BFFB7FF58FB71A64BFB12" bold="true" box="[237,317,1244,1270]" italics="true" pageId="10" pageNumber="249">Hortia</emphasis>
<bibRefCitation id="E3764B5C7B3BFFB7FEF0FB71A685FB12" author="Lozouet" box="[325,499,1244,1270]" pageId="10" pageNumber="249" refString="Lozouet, P. (1999) Nouvelles especes de gasteropodes (Mollusca: Gastropoda) de l'Oligocene et du Miocene inferieur d'Aquitaine (Sud-Ouest de la France). Partie 2. Cossmanniana, 6, 1 - 68, 20 pls." type="journal article" year="1999">Lozouet, 1999</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="875836AD7B3BFFB7FF22FA8EA626FADC" blockId="10.[151,693,1314,1367]" box="[151,336,1314,1336]" pageId="10" pageNumber="249">
<bibRefCitation id="E3764B5C7B3BFFB7FF22FA8EA650FADC" author="Lozouet" box="[151,294,1314,1336]" pageId="10" pageNumber="249" refString="Lozouet, P. (1999) Nouvelles especes de gasteropodes (Mollusca: Gastropoda) de l'Oligocene et du Miocene inferieur d'Aquitaine (Sud-Ouest de la France). Partie 2. Cossmanniana, 6, 1 - 68, 20 pls." type="journal article" year="1999">Lozouet 1999</bibRefCitation>
: 38.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B3BFFB7FF22FAECA3F6FA79" pageId="10" pageNumber="249" type="materials_examined">
<paragraph id="875836AD7B3BFFB7FF22FAECA5C3FAB3" blockId="10.[151,693,1314,1367]" box="[151,693,1345,1367]" pageId="10" pageNumber="249">
<emphasis id="B593EABF7B3BFFB7FF22FAECA654FAB3" bold="true" box="[151,290,1345,1367]" pageId="10" pageNumber="249">
<typeStatus id="585C880F7B3BFFB7FF22FAECA7BAFAB3" box="[151,204,1345,1367]" pageId="10" pageNumber="249">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="40E74D2E7B3BFFB7FE9CFAEFA505FAB3" authority="Lozouet, 1999" authorityName="Lozouet" authorityYear="1999" box="[297,627,1345,1367]" class="Magnoliopsida" family="Rutaceae" genus="Hortia" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="species" species="arriuensis">
<emphasis id="B593EABF7B3BFFB7FE9CFAEFA6AFFAB3" box="[297,473,1346,1367]" italics="true" pageId="10" pageNumber="249">Hortia arriuensis</emphasis>
<bibRefCitation id="E3764B5C7B3BFFB7FE55FAEFA505FAB3" author="Lozouet" box="[480,627,1345,1367]" pageId="10" pageNumber="249" refString="Lozouet, P. (1999) Nouvelles especes de gasteropodes (Mollusca: Gastropoda) de l'Oligocene et du Miocene inferieur d'Aquitaine (Sud-Ouest de la France). Partie 2. Cossmanniana, 6, 1 - 68, 20 pls." type="journal article" year="1999">Lozouet, 1999</bibRefCitation>
</taxonomicName>
(OD).
</paragraph>
<paragraph id="875836AD7B3BFFB7FF22FA29A3F6FA79" blockId="10.[151,1436,1412,1976]" box="[151,1152,1412,1437]" pageId="10" pageNumber="249">
<emphasis id="B593EABF7B3BFFB7FF22FA29A640FA79" bold="true" box="[151,310,1412,1437]" pageId="10" pageNumber="249">Locality/Age.</emphasis>
<collectingCountry id="FFF0763D7B3BFFB7FE88FA28A6F8FA78" box="[317,398,1413,1436]" name="France" pageId="10" pageNumber="249">France</collectingCountry>
: Landes, St-Etienne-d’Orthe (Aquitanian Basin); Upper Oligocene.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B3BFFB7FF72FA0AA652FA01" pageId="10" pageNumber="249" type="distribution">
<paragraph id="875836AD7B3BFFB7FF72FA0AA652FA01" blockId="10.[151,1436,1412,1976]" pageId="10" pageNumber="249">
<emphasis id="B593EABF7B3BFFB7FF72FA0AA616FA24" bold="true" box="[199,352,1447,1472]" pageId="10" pageNumber="249">Distribution.</emphasis>
Upper Eocene to Upper Oligocene—France; Recent—New Zealand,
<collectingCountry id="FFF0763D7B3BFFB7FB6FFA05A2EEFA24" box="[1242,1432,1447,1472]" name="New Caledonia" pageId="10" pageNumber="249">New Caledonia</collectingCountry>
,
<collectingCountry id="FFF0763D7B3BFFB7FF22FA61A657FA01" box="[151,289,1484,1509]" name="Madagascar" pageId="10" pageNumber="249">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B3BFFB7FF72FA42A4BEF8CC" pageId="10" pageNumber="249" type="diagnosis">
<paragraph id="875836AD7B3BFFB7FF72FA42A517F959" blockId="10.[151,1436,1412,1976]" pageId="10" pageNumber="249">
<emphasis id="B593EABF7B3BFFB7FF72FA42A637F9EC" bold="true" box="[199,321,1519,1544]" pageId="10" pageNumber="249">Diagnosis.</emphasis>
Shell small, slightly exceeding
<quantity id="401F9B487B3BFFB7FD0EFA5DA478F9E3" box="[699,782,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="249" unit="mm" value="10.0">10 mm</quantity>
in length, narrowly fusiform, with high spire and long, narrow, slightly abaxially recurved siphonal canal. Protoconch multispiral (fossil species) or paucispiral, bulbous (Recent species). Multispiral protoconchs with more than 4 whorls, last whorls with sharp, opisthocyrt, axial ribs. Spiral sculpture of strong, broadly spaced, narrow spiral cords, intersected by prominent axial ribs, often forming beads on intersections. Cords more closely spaced on canal. Aperture narrow, columella smooth. Outer lip with shallow sinus above periphery in profile.
</paragraph>
<paragraph id="875836AD7B3BFFB7FF72F96AA4BEF8CC" blockId="10.[151,1436,1412,1976]" pageId="10" pageNumber="249">Operculum absent. Rachidian teeth of the radula with broad, deeply notched, anterior base and large cusp emanating from posterior margin, sometimes with additional lateral serration; lateral teeth high but narrow, pointed, triangular, and scoop-shaped, with small denticles on inner side.</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B3BFFB6FF72F899A790FA81" lastPageId="11" lastPageNumber="250" pageId="10" pageNumber="249" type="discussion">
<paragraph id="875836AD7B3BFFB7FF72F899A62CF85C" blockId="10.[151,1436,1412,1976]" pageId="10" pageNumber="249">
<emphasis id="B593EABF7B3BFFB7FF72F899A642F8A9" bold="true" box="[199,308,1844,1869]" pageId="10" pageNumber="249">Remarks</emphasis>
.
<taxonomicName id="40E74D2E7B3BFFB7FEF6F898A6F8F8A8" box="[323,398,1845,1868]" class="Magnoliopsida" family="Rutaceae" genus="Hortia" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="249" phylum="Tracheophyta" rank="genus">
<emphasis id="B593EABF7B3BFFB7FEF6F898A6F8F8A8" box="[323,398,1845,1868]" italics="true" pageId="10" pageNumber="249">Hortia</emphasis>
</taxonomicName>
was originally described in the family
<taxonomicName id="40E74D2E7B3BFFB7FCF9F899A4D9F8A9" box="[844,943,1844,1869]" class="Gastropoda" family="Turridae" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="249" phylum="Mollusca" rank="family">Turridae</taxonomicName>
. It is most similar to
<taxonomicName id="40E74D2E7B3BFFB7FB17F899A25DF8A9" box="[1186,1323,1844,1869]" class="Gastropoda" family="Turridae" genus="Vaughanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="249" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B3BFFB7FB17F899A25DF8A9" box="[1186,1323,1844,1869]" italics="true" pageId="10" pageNumber="249">Vaughanites</emphasis>
</taxonomicName>
, differing by a generally shorter, abaxially recurved canal, and by the absence of columellar plaits. It also differs from
<emphasis id="B593EABF7B3BFFB7FF22F8D0A650F870" box="[151,294,1917,1940]" italics="true" pageId="10" pageNumber="249">Pyramimitra</emphasis>
in having narrower axial ribs (rather than varices) and a longer siphonal canal, and the absence of columellar plaits.
</paragraph>
<caption id="D39866257B3AFFB6FF22FB2AA402FB5A" httpUri="https://zenodo.org/record/251555/files/figure.png" pageId="11" pageNumber="250" targetBox="[151,1436,195,1133]" targetPageId="11">
<paragraph id="875836AD7B3AFFB6FF22FB2AA402FB5A" blockId="11.[151,1436,1159,1214]" pageId="11" pageNumber="250">
<emphasis id="B593EABF7B3AFFB6FF22FB2AA65EFB7A" bold="true" box="[151,296,1159,1182]" pageId="11" pageNumber="250">FIGURE 4.</emphasis>
<emphasis id="B593EABF7B3AFFB6FE88FB2AA6C4FB7A" box="[317,434,1159,1182]" italics="true" pageId="11" pageNumber="250">Endiatoma</emphasis>
<taxonomicName id="40E74D2E7B3AFFB6FE7DFB2AA44DFB7A" authority="(Cossmann, 1883)" authorityName="Cossmann" authorityYear="1883" baseAuthorityName="Cossmann" baseAuthorityYear="1883" box="[456,827,1159,1182]" class="Gastropoda" family="Borsoniidae" genus="Asthenotoma" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="250" phylum="Mollusca" rank="species" species="quadricincta">
<emphasis id="B593EABF7B3AFFB6FE7DFB2AA526FB7A" box="[456,592,1159,1182]" italics="true" pageId="11" pageNumber="250">quadricincta</emphasis>
(Cossmann, 1883)
</taxonomicName>
, Lower Eocene, St-Gobain, Aisne, France.
<emphasis id="B593EABF7B3AFFB6FAD6FB2AA2E3FB7A" bold="true" box="[1379,1429,1159,1182]" pageId="11" pageNumber="250">A–B</emphasis>
. MNHN.F.
<accessionNumber id="98B4AB4E7B3AFFB6FEB7FB0AA62DFB5A" box="[258,347,1191,1214]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A47757" pageId="11" pageNumber="250" type="EnaNcbi">A47757</accessionNumber>
;
<emphasis id="B593EABF7B3AFFB6FED3FB0AA6EFFB5A" bold="true" box="[358,409,1191,1214]" pageId="11" pageNumber="250">C–E</emphasis>
. MNHN.F.
<accessionNumber id="98B4AB4E7B3AFFB6FDA4FB0AA51DFB5A" box="[529,619,1191,1214]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A47757" pageId="11" pageNumber="250" type="EnaNcbi">A47757</accessionNumber>
;
<emphasis id="B593EABF7B3AFFB6FDC0FB0AA5D2FB5A" bold="true" box="[629,676,1191,1214]" pageId="11" pageNumber="250">F–G</emphasis>
. MNHN.F.
<accessionNumber id="98B4AB4E7B3AFFB6FCAEFB0AA402FB5A" box="[795,884,1191,1214]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A47759" pageId="11" pageNumber="250" type="EnaNcbi">A47759</accessionNumber>
</paragraph>
</caption>
<paragraph id="875836AD7B3AFFB6FF72FAAAA790FA81" blockId="11.[151,1436,1287,1381]" pageId="11" pageNumber="250">
Shells of Recent species of
<taxonomicName id="40E74D2E7B3AFFB6FE49FAA5A531FAFB" box="[508,583,1288,1311]" class="Magnoliopsida" family="Rutaceae" genus="Hortia" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="250" phylum="Tracheophyta" rank="genus">
<emphasis id="B593EABF7B3AFFB6FE49FAA5A531FAFB" box="[508,583,1288,1311]" italics="true" pageId="11" pageNumber="250">Hortia</emphasis>
</taxonomicName>
are similar to fossil ones, but attain a larger adult size. All Recent species known so far have a non-planktotrophic, bulbous, smooth protoconch rather than the multispiral protoconch of the fossils.
</paragraph>
</subSubSection>
</treatment>
</document>
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